treatments-xml/data/03/E0/87/03E087A4FFC6543BACB5FC22FAC6FC7E.xml

<document id="C62D5D037A3E53B40AD91E6BAEFC922E" ID-CLB-Dataset="22045" ID-DOI="10.5852/ejt.2016.183" ID-GBIF-Dataset="29033971-cc71-44ce-9a46-9ab104d684ea" ID-ISSN="2118-9773" ID-Zenodo-Dep="3833125" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589841522561" checkinUser="valdenar" docAuthor="Turner, Ian M. &amp; Utteridge, Timothy M. A." docDate="2016" docId="03E087A4FFC6543BACB5FC22FAC6FC7E" docLanguage="en" docName="ejt-2016-183.pdf.imf" docOrigin="European Journal of Taxonomy 183" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Polyalthia pakdin I. M. Turner &amp; Utteridge 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFD9FFDCFFC95429AD5AFFFDFFAFFFB5" masterDocTitle="Whither Polyalthia (Annonaceae) in Peninsular Malaysia? Synopses of Huberantha, Maasia, Monoon and Polyalthia s. s." masterLastPageNumber="26" masterPageNumber="1" pageNumber="16" updateTime="1698840243520" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CA5C5418F23FB5611F49EC8EEF801EB6">Whither Polyalthia (Annonaceae) in Peninsular Malaysia? Synopses of Huberantha, Maasia, Monoon and Polyalthia s. s.</mods:title>
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<mods:namePart id="A920D0A44829A150B3C40ED8F39C7B96">Turner, Ian M.</mods:namePart>
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<mods:namePart id="CB77501999133E666692B8DDFC2B3C22">Utteridge, Timothy M. A.</mods:namePart>
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<mods:title id="BEB5E21F4B09AB0438761134184277D6">European Journal of Taxonomy</mods:title>
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<mods:date id="238AB19BC14E525F2CD54E006A22380C">2016</mods:date>
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<treatment id="03E087A4FFC6543BACB5FC22FAC6FC7E" ID-DOI="http://doi.org/10.5281/zenodo.3850937" ID-GBIF-Taxon="164167864" ID-Zenodo-Dep="3850937" LSID="urn:lsid:plazi:treatment:03E087A4FFC6543BACB5FC22FAC6FC7E" httpUri="http://treatment.plazi.org/id/03E087A4FFC6543BACB5FC22FAC6FC7E" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
<subSubSection id="C3536539FFC65426ACB5FC22FBEAFC4F" box="[495,1093,991,1018]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BF636B2FFC65426ACB5FC22FBEAFC4F" blockId="15.[495,1093,991,1091]" box="[495,1093,991,1018]" pageId="15" pageNumber="16">
<heading id="D0BE81DEFFC65426ACB5FC22FBEAFC4F" box="[495,1093,991,1018]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="4C494D31FFC65426ACB5FC22FC4CFC4C" authority="I.M.Turner &amp; Utteridge" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[495,995,991,1018]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFC65426ACB5FC22FD69FC4F" bold="true" box="[495,710,991,1018]" italics="true" pageId="15" pageNumber="16">Polyalthia pakdin</emphasis>
I.M.Turner &amp; Utteridge
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFC65426AEB0FC1DFBEAFC4F" box="[1002,1093,992,1018]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3536539FFC65426AFBDFBF9FC42FBF6" pageId="15" pageNumber="16" type="description">
<paragraph id="8BF636B2FFC65426AFBDFBF9FC42FBF6" blockId="15.[495,1093,991,1091]" pageId="15" pageNumber="16">
<figureCitation id="13722A37FFC65426AFBDFBF9FCE2FBAB" box="[743,845,1028,1054]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1824,1850]" captionStartId-1="18.[189,232,2000,2026]" captionTargetBox-0="[236,1350,276,1765]" captionTargetBox-1="[189,1398,1355,1960]" captionTargetId-0="figure@16.[233,1355,265,1784]" captionTargetId-1="figure@18.[189,1398,1355,1960]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionText-0="Fig. 1. Polyalthia pakdin I.M.Turner &amp; Utteridge sp. nov. A. Leafy twig with flowers. B. Inflorescence and base of leaf (abaxial view). C. Flower. D. Magnified view of abaxial surface of outer petal. E. Calyx from below. F. Stamen. G. Carpel. H. Infructescense with immature monocarp. I. Mature monocarp. J. Seed. Scale bars: single bar = 1 mm, graduated single bar = 5 mm; double bar = 1 cm, graduated double bar = 5 cm. Drawn from Carrick s.n. (A, C–E), KL 3033 (B, F, G), SFN 40941 (H), FRI 11853 (I), FRI 13467 (J)." captionText-1="Fig. 2. Map showing the collecting localities of Polyalthia pakdin I.M.Turner &amp; Utteridge sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.3833127" figureDoi-1="http://doi.org/10.5281/zenodo.3833129" httpUri-0="https://zenodo.org/record/3833127/files/figure.png" httpUri-1="https://zenodo.org/record/3833129/files/figure.png" pageId="15" pageNumber="16">Figs 1–2</figureCitation>
urn:lsid:ipni.org:names:77153637-1
</paragraph>
</subSubSection>
<subSubSection id="C3536539FFC65426ADE7FB96FD86FAFC" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BF636B2FFC65426ADE7FB96FE99FB30" blockId="15.[189,310,1131,1157]" box="[189,310,1131,1157]" pageId="15" pageNumber="16">
<heading id="D0BE81DEFFC65426ADE7FB96FE99FB30" bold="true" box="[189,310,1131,1157]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B93DEAA0FFC65426ADE7FB96FE99FB30" bold="true" box="[189,310,1131,1157]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFC65426ADE7FB66FD86FAFC" blockId="15.[189,1399,1179,1353]" pageId="15" pageNumber="16">
With its large, multi-nerved leaves the new species has generally been confused with 
<taxonomicName id="4C494D31FFC65426A9F0FB66FF54FB6F" authority="King" authorityName="King" class="Magnoliopsida" family="Annonaceae" genus="Polyaltha" kingdom="Plantae" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="bullata">
<emphasis id="B93DEAA0FFC65426A9F0FB66FAD9FB00" box="[1194,1398,1179,1205]" italics="true" pageId="15" pageNumber="16">Polyaltha bullata</emphasis>
King
</taxonomicName>
, but it lacks the villose tomentum on the twigs and the strongly auriculate to cordate leaf bases. In comparison with 
<taxonomicName id="4C494D31FFC65426ACF0FB1BFDB4FB4A" box="[426,539,1253,1279]" class="Magnoliopsida" family="Annonaceae" genus="Polyaltha" kingdom="Plantae" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="bullata">
<emphasis id="B93DEAA0FFC65426ACF0FB1BFDB4FB4A" box="[426,539,1253,1279]" italics="true" pageId="15" pageNumber="16">P. bullata</emphasis>
</taxonomicName>
, the flowers are borne on shorter pedicels (&lt;1 versus 
<quantity id="4CB19B57FFC65426A9C7FB18FB76FB4A" box="[1181,1241,1253,1279]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="cm" value="2.0">2 cm</quantity>
) with shorter sepals (3–4 versus 
<quantity id="4CB19B57FFC65426ACC6FAF7FE4DFA91" box="[412,482,1290,1316]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="7.0">7 mm</quantity>
) and petals (to 
<quantity id="4CB19B57FFC65426AFCCFAF7FD7FFA91" box="[662,720,1290,1316]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="cm" value="1.0">1 cm</quantity>
versus up to 
<quantity id="4CB19B57FFC65426AE37FAF7FC04FA91" box="[877,939,1290,1316]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="cm" value="4.0">4 cm</quantity>
); and with monocarps on shorter stipes (to 
<quantity id="4CB19B57FFC65426ADBFFAD3FE86FAFC" box="[229,297,1326,1353]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="6.0">6 mm</quantity>
versus up to 
<quantity id="4CB19B57FFC65426AC9FFAD3FDB2FAFC" box="[453,541,1326,1353]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="10.0">10 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536539FFC65426ADE7FA8CFAD8F99C" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BF636B2FFC65426ADE7FA8CFEEBFA3E" blockId="15.[189,324,1393,1419]" box="[189,324,1393,1419]" pageId="15" pageNumber="16">
<heading id="D0BE81DEFFC65426ADE7FA8CFEEBFA3E" bold="true" box="[189,324,1393,1419]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B93DEAA0FFC65426ADE7FA8CFEEBFA3E" bold="true" box="[189,324,1393,1419]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFC65426ADE7FA5FFAD8F99C" blockId="15.[189,1399,1441,1577]" pageId="15" pageNumber="16">
We are pleased to commemorate the late Prof. Kamarudin Mat Salleh (
<date id="FFF71072FFC65426AEB6FA5CFBD8FA09" box="[1004,1143,1441,1468]" pageId="15" pageNumber="16" value="1959" valueMax="2009">1959–2009</date>
). A leading Malaysian student of the 
<taxonomicName id="4C494D31FFC65426AC36FA3BFDAFFA55" box="[364,512,1478,1504]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
, he was widely known as ‘Pak Din’ (Uncle Din in Malay, Din being the common abbreviation of names with that ending). The epithet represents a compounding of this soubriquet. Kamarudin was native to the state of 
<collectingRegion id="498DF850FFC65426AFACF9F2FCCCF99C" box="[758,867,1551,1577]" country="Malaysia" name="Kelantan" pageId="15" pageNumber="16">Kelantan</collectingRegion>
, which is part of the range of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3536539FFC65438ADE7F9AFFE29FEFC" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="materials_examined">
<materialsCitation id="3B213CEFFFC65426ADE7F9AFFC6AF974" ID-GBIF-Occurrence="2620554304" collectingDate="1972-03-21" collectionCode="L, E, G, KL, T, P, K" country="Malaysia" latitude="4.8333335" location="Teo" longLatPrecision="1306" longitude="103.36667" municipality="Jambu Bongkok Forest Reserve" pageId="15" pageNumber="16" specimenCount="1" typeStatus="type">
<paragraph id="8BF636B2FFC65426ADE7F9AFFF55F9D9" blockId="15.[189,250,1618,1644]" box="[189,250,1618,1644]" pageId="15" pageNumber="16">
<heading id="D0BE81DEFFC65426ADE7F9AFFF55F9D9" bold="true" box="[189,250,1618,1644]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B93DEAA0FFC65426ADE7F9AFFF55F9D9" bold="true" box="[189,250,1618,1644]" pageId="15" pageNumber="16">
<typeStatus id="54F28810FFC65426ADE7F9AFFF55F9D9" box="[189,250,1618,1644]" pageId="15" pageNumber="16">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFC65426ADE7F97FFC6AF974" blockId="15.[189,1399,1665,1729]" pageId="15" pageNumber="16">
PENINSULAR 
<collectingCountry id="F35E7622FFC65426AC20F97FFDB8F929" box="[378,535,1666,1692]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
: TRENGGANU, 
<collectingMunicipality id="6B92ACC8FFC65426AFE9F97FFB89F929" box="[691,1062,1666,1692]" pageId="15" pageNumber="16">Jambu Bongkok Forest Reserve</collectingMunicipality>
[
<geoCoordinate id="EE7D5075FFC65426A96EF97FFB21F928" box="[1076,1166,1666,1693]" degrees="4" direction="north" minutes="50" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="4.8333335">4°50’N</geoCoordinate>
<geoCoordinate id="EE7D5075FFC65426A9CEF97CFAA6F929" box="[1172,1289,1665,1692]" degrees="103" direction="east" minutes="22" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="103.36667">103°22’E</geoCoordinate>
], 
<date id="FFF71072FFC65426A846F97FFF55F974" pageId="15" pageNumber="16" value="1972-03-21">
<collectingDate id="EFB3E99AFFC65426A846F97FFF55F974" pageId="15" pageNumber="16" value="1972-03-21">21 Mar. 1972</collectingDate>
</date>
, 
<emphasis id="B93DEAA0FFC65426AC5CF95AFD03F974" box="[262,684,1702,1729]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426AC5CF95AFEB6F975" box="[262,281,1703,1728]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="15" pageNumber="16" type="Herbarium">L</collectionCode>
. 
<collectionCode id="ED58AE77FFC65426AC44F95AFE9BF975" box="[286,308,1703,1728]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="15" pageNumber="16" type="Herbarium">E</collectionCode>
. 
<location id="8E966069FFC65426AC65F95AFEC7F975" LSID="urn:lsid:plazi:treatment:03E087A4FFC6543BACB5FC22FAC6FC7E:8E966069FFC65426AC65F95AFEC7F975" box="[319,360,1703,1728]" country="Malaysia" latitude="4.8333335" longLatPrecision="1306" longitude="103.36667" municipality="Jambu Bongkok Forest Reserve" name="Teo" pageId="15" pageNumber="16">Teo</location>
&amp; 
<collectionCode id="ED58AE77FFC65426ACD7F95AFE0AF974" box="[397,421,1703,1729]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="15" pageNumber="16" type="Herbarium">G</collectionCode>
. 
<location id="8E966069FFC65426ACEBF95BFD90F975" LSID="urn:lsid:plazi:treatment:03E087A4FFC6543BACB5FC22FAC6FC7E:8E966069FFC65426ACEBF95BFD90F975" box="[433,575,1702,1728]" country="Malaysia" latitude="4.8333335" longLatPrecision="1306" longitude="103.36667" municipality="Jambu Bongkok Forest Reserve" name="Pachiappan" pageId="15" pageNumber="16">Pachiappan</location>
<collectionCode id="ED58AE77FFC65426AF1CF95AFDC6F975" box="[582,617,1703,1728]" country="Austria" lsid="urn:lsid:biocol.org:col:14990" name="Landesmuseum für Kärnten" pageId="15" pageNumber="16" type="Herbarium">KL</collectionCode>
3033
</emphasis>
[
<emphasis id="B93DEAA0FFC65426AFE7F95AFC98F974" box="[701,823,1703,1729]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426AFE7F95AFD62F975" box="[701,717,1703,1728]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
&amp; 
<collectionCode id="ED58AE77FFC65426AFA8F95AFCACF975" box="[754,771,1703,1728]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="15" pageNumber="16" type="Herbarium">P</collectionCode>
433
</emphasis>
] (holo-: 
<collectionCode id="ED58AE77FFC65426AEC5F95AFC17F974" box="[927,952,1703,1729]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="15" pageNumber="16" type="Herbarium">K</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BF636B2FFC65426ADE7F914FDE6F8B6" blockId="15.[189,585,1769,1795]" box="[189,585,1769,1795]" pageId="15" pageNumber="16">
<heading id="D0BE81DEFFC65426ADE7F914FDE6F8B6" bold="true" box="[189,585,1769,1795]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B93DEAA0FFC65426ADE7F914FDE6F8B6" bold="true" box="[189,585,1769,1795]" pageId="15" pageNumber="16">Additional specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFC65438ADE7F8E7FE29FEFC" blockId="15.[189,1399,1817,2027]" lastBlockId="17.[189,1399,267,329]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
PENINSULAR 
<collectingCountry id="F35E7622FFC65426AC26F8E7FDB6F881" box="[380,537,1818,1844]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
: 
<collectingRegion id="498DF850FFC65426AF7FF8E7FD36F884" box="[549,665,1818,1844]" country="Malaysia" name="Kelantan" pageId="15" pageNumber="16">KELANTAN</collectingRegion>
, Chabang Tongkat Forest Reserve [
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<geoCoordinate id="EE7D5075FFC65426A9F9F8E4FAB7F881" box="[1187,1304,1817,1844]" degrees="102" direction="east" minutes="17" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="102.28333">102°17’E</geoCoordinate>
], compt 77, alt. 250’ 
<date id="FFF71072FFC65426AC0BF8C3FE43F8ED" box="[337,492,1854,1880]" pageId="15" pageNumber="16" value="1967-04-28">28 Apr. 1967</date>
, 
<emphasis id="B93DEAA0FFC65426ACA3F8C2FCE9F8ED" box="[505,838,1854,1880]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426ACA3F8C2FDBFF8ED" box="[505,528,1855,1880]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="15" pageNumber="16" type="Herbarium">K</collectionCode>
. 
<collectionCode id="ED58AE77FFC65426AF49F8C2FD9FF8ED" box="[531,560,1855,1880]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="15" pageNumber="16" type="Herbarium">M</collectionCode>
. Kochummen 
<collectionCode id="ED58AE77FFC65426AF8FF8C2FCACF8ED" box="[725,771,1855,1880]" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="15" pageNumber="16">FRI</collectionCode>
2317
</emphasis>
(
<collectionCode id="ED58AE77FFC65426AE0FF8C3FC3CF8ED" box="[853,915,1854,1880]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="15" pageNumber="16" type="Herbarium">KEP</collectionCode>
). 
<collectingRegion id="498DF850FFC65426AEFCF8C2FB84F8E3" box="[934,1067,1855,1881]" country="Malaysia" name="TRENGGANU" pageId="15" pageNumber="16">TRENGGANU</collectingRegion>
, Dunggun, Jambu Bongkok [
<geoCoordinate id="EE7D5075FFC65426AD9EF89FFEB1F8C8" box="[196,286,1890,1917]" degrees="4" direction="north" minutes="50" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="4.8333335">4°50’N</geoCoordinate>
<geoCoordinate id="EE7D5075FFC65426AC7DF89FFE34F8C8" box="[295,411,1890,1917]" degrees="103" direction="east" minutes="21" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="103.35">103°21’E</geoCoordinate>
], 
<date id="FFF71072FFC65426ACE8F89EFDFAF8C8" box="[434,597,1891,1917]" pageId="15" pageNumber="16" value="1961-03-12">12 Mar. 1961</date>
, 
<emphasis id="B93DEAA0FFC65426AF39F899FCA2F8C8" box="[611,781,1891,1917]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426AF39F899FDDBF8C8" box="[611,628,1892,1917]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="15" pageNumber="16" type="Herbarium">J</collectionCode>
. Carrick s.n.
</emphasis>
(
<collectionCode id="ED58AE77FFC65426AE45F89EFC99F8C8" box="[799,822,1891,1917]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="15" pageNumber="16" type="Herbarium">K</collectionCode>
×2); ibid., Sekayu, Bukit Lanjut Forest Reserve [
<geoCoordinate id="EE7D5075FFC65426AD9EF87AFEB1F817" box="[196,286,1927,1954]" degrees="4" direction="north" minutes="58" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="4.9666667">4°58’N</geoCoordinate>
<geoCoordinate id="EE7D5075FFC65426AC7EF87AFE35F817" box="[292,410,1927,1954]" degrees="102" direction="east" minutes="58" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="102.96667">102°58’E</geoCoordinate>
], 
<date id="FFF71072FFC65426ACF7F87AFDFFF814" box="[429,592,1927,1953]" pageId="15" pageNumber="16" value="1969-09-18">18 Sept. 1969</date>
, 
<emphasis id="B93DEAA0FFC65426AF06F875FCE8F814" box="[604,839,1927,1954]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426AF06F875FDD9F817" box="[604,630,1928,1954]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="15" pageNumber="16" type="Herbarium">H</collectionCode>
. 
<collectionCode id="ED58AE77FFC65426AF20F875FD24F817" box="[634,651,1928,1954]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="15" pageNumber="16" type="Herbarium">S</collectionCode>
. Loh 
<collectionCode id="ED58AE77FFC65426AF90F875FD57F814" box="[714,760,1928,1953]" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="15" pageNumber="16">FRI</collectionCode>
13467
</emphasis>
(
<collectionCode id="ED58AE77FFC65426AE0DF87AFC3BF814" box="[855,916,1927,1953]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="15" pageNumber="16" type="Herbarium">KEP</collectionCode>
); ibid., Dungun, near 36th mile, Jerangau Road [
<geoCoordinate id="EE7D5075FFC65426AC56F851FE20F872" box="[268,399,1964,1991]" degrees="4" direction="north" minutes="56" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="36" value="4.9433336">4°56’36”N</geoCoordinate>
<geoCoordinate id="EE7D5075FFC65426ACC2F856FD97F873" box="[408,568,1963,1990]" degrees="103" direction="east" minutes="10" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="42" value="103.17833">103°10’42”E</geoCoordinate>
], 
<date id="FFF71072FFC65426AF14F851FD58F872" box="[590,759,1964,1991]" pageId="15" pageNumber="16" value="1955-09-22">22 Sept. 1955</date>
, 
<emphasis id="B93DEAA0FFC65426AE5FF850FB5BF873" box="[773,1268,1964,1991]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426AE5FF850FCB9F873" box="[773,790,1965,1990]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="15" pageNumber="16" type="Herbarium">J</collectionCode>
. Sinclair &amp; 
<taxonomicName id="4C494D31FFC65426AEF3F851FBCBF873" authority="bin Salleh" authorityName="bin Salleh" box="[937,1124,1964,1990]" class="Insecta" family="Mogoplistidae" genus="Kiah" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Kiah bin Salleh</taxonomicName>
<collectionCode id="ED58AE77FFC65426A937F851FB0EF873" box="[1133,1185,1964,1990]" pageId="15" pageNumber="16">SFN</collectionCode>
40941
</emphasis>
(
<collectingRegion id="498DF850FFC65426A85CF851FAB1F873" box="[1286,1310,1964,1990]" country="Malaysia" name="K" pageId="15" pageNumber="16">K</collectingRegion>
, 
<collectionCode id="ED58AE77FFC65426A877F850FAC6F872" box="[1325,1385,1965,1991]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="15" pageNumber="16" type="Herbarium">KEP</collectionCode>
); ibid., Sekayu Forest Reserve [
<geoCoordinate id="EE7D5075FFC65426AF4CF82DFDC0F85E" box="[534,623,2000,2027]" degrees="4" direction="north" minutes="58" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="4.9666667">4°58’N</geoCoordinate>
<geoCoordinate id="EE7D5075FFC65426AF2FF82DFD45F85E" box="[629,746,2000,2027]" degrees="102" direction="east" minutes="57" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="102.95">102°57’E</geoCoordinate>
], compt 43, 
<date id="FFF71072FFC65426AE2CF82CFBB8F85E" box="[886,1047,2001,2027]" pageId="15" pageNumber="16" value="1969-09-22">22 Sept. 1969</date>
, 
<emphasis id="B93DEAA0FFC65426A978F82FFAB2F85E" box="[1058,1309,2001,2027]" italics="true" pageId="15" pageNumber="16">
<collectionCode id="ED58AE77FFC65426A978F82FFB9BF85E" box="[1058,1076,2002,2027]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Suppiah 
<collectionCode id="ED58AE77FFC65426A9F9F82FFB7EF85E" box="[1187,1233,2002,2027]" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="15" pageNumber="16">FRI</collectionCode>
11853
</emphasis>
(
<collectionCode id="ED58AE77FFC65426A876F82CFAC6F85E" box="[1324,1385,2001,2027]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="15" pageNumber="16" type="Herbarium">KEP</collectionCode>
); ibid., Bukit Bauk Forest Reserve, compartment 8 
<collectionCode id="ED58AE77FFD85438AE5CFEF6FCB3FE90" box="[774,796,267,293]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="17" pageNumber="18" type="Herbarium">A</collectionCode>
[
<geoCoordinate id="EE7D5075FFD85438AE76FEF6FC29FE93" box="[812,902,267,294]" degrees="4" direction="north" minutes="45" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="4.75">4°45’N</geoCoordinate>
<geoCoordinate id="EE7D5075FFD85438AED5FEF6FBACFE93" box="[911,1027,267,294]" degrees="103" direction="east" minutes="21" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="103.35">103°21’E</geoCoordinate>
], 
<date id="FFF71072FFD85438A940FEF6FB13FE90" box="[1050,1212,267,294]" pageId="17" pageNumber="18" value="1967-06-17">17 June 1967</date>
, 
<emphasis id="B93DEAA0FFD85438A990FEF1FE82FEFC" italics="true" pageId="17" pageNumber="18">
<collectionCode id="ED58AE77FFD85438A990FEF1FB71FE93" box="[1226,1246,268,294]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="18">T</collectionCode>
. 
<collectionCode id="ED58AE77FFD85438A9B8FEF1FB58FE93" box="[1250,1271,268,294]" country="Denmark" name="University of Copenhagen" pageId="17" pageNumber="18" type="Herbarium">C</collectionCode>
. Whitmore 
<collectionCode id="ED58AE77FFD85438ADE7FECDFF44FEFC" box="[189,235,304,329]" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="17" pageNumber="18">FRI</collectionCode>
3903
</emphasis>
(
<collectionCode id="ED58AE77FFD85438AC66FED2FED5FEFC" box="[316,378,303,329]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="17" pageNumber="18" type="Herbarium">KEP</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF36663AFFD95439ADE7F8DDFE9BF85E" ID-DOI="http://doi.org/10.5281/zenodo.3833127" ID-Zenodo-Dep="3833127" httpUri="https://zenodo.org/record/3833127/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1824,1850]" targetBox="[236,1350,276,1765]" targetPageId="16">
<paragraph id="8BF636B2FFD95439ADE7F8DDFE9BF85E" blockId="16.[189,1399,1824,2027]" pageId="16" pageNumber="17">
<emphasis id="B93DEAA0FFD95439ADE7F8DDFEA4F88E" bold="true" box="[189,267,1824,1851]" pageId="16" pageNumber="17">Fig. 1.</emphasis>
<taxonomicName id="4C494D31FFD95439AC48F8DCFCADF88E" authority="I.M.Turner &amp; Utteridge" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[274,770,1825,1851]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFD95439AC48F8DCFE4BF88E" box="[274,484,1825,1851]" italics="true" pageId="16" pageNumber="17">Polyalthia pakdin</emphasis>
I.M.Turner &amp; Utteridge
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFD95439AE53F8DCFCCBF88E" box="[777,868,1825,1851]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93DEAA0FFD95439AE31F8DDFC2FF88F" bold="true" box="[875,896,1824,1850]" pageId="16" pageNumber="17">A</emphasis>
. Leafy twig with flowers. 
<emphasis id="B93DEAA0FFD95439A9E2F8DCFB63F88E" bold="true" box="[1208,1228,1825,1851]" pageId="16" pageNumber="17">B</emphasis>
. Inflorescence and base of leaf (abaxial view). 
<emphasis id="B93DEAA0FFD95439AF75F8B9FDEBF8EB" bold="true" box="[559,580,1860,1886]" pageId="16" pageNumber="17">C</emphasis>
. Flower. 
<emphasis id="B93DEAA0FFD95439AFEBF8B9FD69F8EB" bold="true" box="[689,710,1860,1886]" pageId="16" pageNumber="17">D</emphasis>
. Magnified view of abaxial surface of outer petal. 
<emphasis id="B93DEAA0FFD95439A855F8B9FA8CF8EB" bold="true" box="[1295,1315,1860,1886]" pageId="16" pageNumber="17">E</emphasis>
. Calyx from below. 
<emphasis id="B93DEAA0FFD95439AC0BF89AFECCF834" bold="true" box="[337,355,1895,1921]" pageId="16" pageNumber="17">F</emphasis>
. Stamen. 
<emphasis id="B93DEAA0FFD95439AC82F89AFE40F834" bold="true" box="[472,495,1895,1921]" pageId="16" pageNumber="17">G</emphasis>
. Carpel. 
<emphasis id="B93DEAA0FFD95439AF00F89AFDDEF834" bold="true" box="[602,625,1895,1921]" pageId="16" pageNumber="17">H</emphasis>
. Infructescense with immature monocarp. 
<emphasis id="B93DEAA0FFD95439A93DF89AFBDDF834" bold="true" box="[1127,1138,1895,1921]" pageId="16" pageNumber="17">I</emphasis>
. Mature monocarp. 
<emphasis id="B93DEAA0FFD95439A83AF89AFAC0F834" bold="true" box="[1376,1391,1895,1921]" pageId="16" pageNumber="17">J</emphasis>
. Seed. Scale bars: single bar = 1 mm, graduated single bar = 5 mm; double bar = 1 cm, graduated double bar = 5 cm. Drawn from 
<emphasis id="B93DEAA0FFD95439ACB9F850FDC8F872" box="[483,615,1965,1991]" italics="true" pageId="16" pageNumber="17">Carrick s.n</emphasis>
. (A, C–E), 
<emphasis id="B93DEAA0FFD95439AFABF853FCF8F87D" box="[753,855,1966,1992]" italics="true" pageId="16" pageNumber="17">KL 3033</emphasis>
(B, F, G), 
<emphasis id="B93DEAA0FFD95439AE8FF853FBF6F87D" box="[981,1113,1966,1992]" italics="true" pageId="16" pageNumber="17">SFN 40941</emphasis>
(H), 
<emphasis id="B93DEAA0FFD95439A9C3F853FABAF87D" box="[1177,1301,1966,1992]" italics="true" pageId="16" pageNumber="17">FRI 11853</emphasis>
(I), 
<emphasis id="B93DEAA0FFD95439A813F853FEA9F85E" italics="true" pageId="16" pageNumber="17">FRI 13467</emphasis>
(J).
</paragraph>
</caption>
<subSubSection id="C3536539FFD85438ADE7FE8CFCF8FA8E" pageId="17" pageNumber="18" type="description">
<paragraph id="8BF636B2FFD85438ADE7FE8CFEE1FE3E" blockId="17.[189,334,369,395]" box="[189,334,369,395]" pageId="17" pageNumber="18">
<heading id="D0BE81DEFFD85438ADE7FE8CFEE1FE3E" bold="true" box="[189,334,369,395]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B93DEAA0FFD85438ADE7FE8CFEE1FE3E" bold="true" box="[189,334,369,395]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFD85438ADE7FE5DFD81FB8E" blockId="17.[189,1399,416,1083]" pageId="17" pageNumber="18">
Shrub 
<quantity id="4CB19B57FFD85438AC4BFE5DFECCFE0F" box="[273,355,416,442]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="17" pageNumber="18" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">1–4 m</quantity>
tall. Young twigs drying red-brown, rather smooth and shiny, with some irregular longitudinal ridges with rounded tops, varying from glabrous to densely pale short hairy, twigs becoming black to brown with age. Leaves chartaceous to subcoriaceous (tend to look more leathery than they are to the touch), often bullate, drying grey or grey-brown, more rarely dark brown, above, brown or greybrown below with brown venation, midrib and laterals sunken above in dry leaves, prominent beneath, more or less glabrous though sometimes with pale hairs along midrib below near base, lamina oblongoblanceolate to narrowly oblong-elliptic, 17–36 × 
<quantity id="4CB19B57FFD85438AE44FD88FC3DFD25" box="[798,914,629,656]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="9.0" metricValueMin="3.5" pageId="17" pageNumber="18" unit="cm" value="6.25" valueMax="9.0" valueMin="3.5">3.5–9 cm</quantity>
, apex acuminate, base obtuse, generally slightly asymmetric, lateral nerves 19–24 pairs, arching and looping within margin to form a scalloped intramarginal nerve, reticulations distinct. Petiole drying dark brown to black, 
<quantity id="4CB19B57FFD85438A900FD40FB65FD62" box="[1114,1226,701,727]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="17" pageNumber="18" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
long, 
<quantity id="4CB19B57FFD85438A84FFD40FAD8FD62" box="[1301,1399,701,727]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
in diameter, sometimes with short erect pale hairs. Inflorescences subopposite leaves, single- or fewflowered. Flowering pedicel 
<quantity id="4CB19B57FFD85438AF57FCF9FDC1FCAB" box="[525,622,772,798]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">2–8 mm</quantity>
long, 
<quantity id="4CB19B57FFD85438AFEDFCF9FD55FCAB" box="[695,762,772,798]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
diameter widening distally, drying brown with minute pale hairs [some 4-merous flowers present on 
<emphasis id="B93DEAA0FFD85438AF81FCD5FCEEFCF7" box="[731,833,808,834]" italics="true" pageId="17" pageNumber="18">KL 3033</emphasis>
], sepals ovate 
<emphasis id="B93DEAA0FFD85438AEAAFCD5FBABFCF4" box="[1008,1028,808,833]" italics="true" pageId="17" pageNumber="18">c.</emphasis>
3–4 × 
<quantity id="4CB19B57FFD85438A902FCD5FB13FCF7" box="[1112,1212,808,834]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, outside brown, minutely bumpy with short pale hairs, inside glabrous, petals thin-textured, ovate to ovate-lanceolate 6–10 × 
<quantity id="4CB19B57FFD85438AC40FC92FE39FC3C" box="[282,406,879,906]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="17" pageNumber="18" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
, drying dark brown to blackish, outside minutely bumpy with adpressed pale hairs, inside glabrous, stamens many 
<emphasis id="B93DEAA0FFD85438AF6CFC6EFDE5FC19" box="[566,586,915,940]" italics="true" pageId="17" pageNumber="18">c.</emphasis>
<quantity id="4CB19B57FFD85438AF0EFC6FFD1FFC18" box="[596,688,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="mm" value="1.5">1.5 mm</quantity>
long, carpels many. Fruiting pedicel to 
<quantity id="4CB19B57FFD85438A9C9FC6FFB75FC18" box="[1171,1242,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="2.0">2 mm</quantity>
thick, sepals often persisting, monocarps to 9 or more, irregularly ellipsoidal, 8–10 × 
<quantity id="4CB19B57FFD85438A970FC4BFB3EFC64" box="[1066,1169,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="17" pageNumber="18" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
, drying red-brown, surface minutely pimply with pale hairs, apiculate, stipe to 
<quantity id="4CB19B57FFD85438AED3FC24FC60FC41" box="[905,975,985,1012]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="mm" value="6.0">6 mm</quantity>
long, 
<quantity id="4CB19B57FFD85438A947FC27FBCBFC41" box="[1053,1124,986,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
diameter, pimply, pale hairy. Seeds generally two, lenticular, 
<emphasis id="B93DEAA0FFD85438AF26FC03FD3FFBA2" box="[636,656,1022,1047]" italics="true" pageId="17" pageNumber="18">c.</emphasis>
7 × 6 × 
<quantity id="4CB19B57FFD85438AFA8FC00FC96FBAD" box="[754,825,1021,1048]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="mm" value="4.0">4 mm</quantity>
, drying golden brown with a longitudinal groove running parallel to the flat face.
</paragraph>
<paragraph id="8BF636B2FFD85438ADE7FB9FFEEAFBC8" blockId="17.[189,325,1122,1149]" box="[189,325,1122,1149]" pageId="17" pageNumber="18">
<heading id="D0BE81DEFFD85438ADE7FB9FFEEAFBC8" bold="true" box="[189,325,1122,1149]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B93DEAA0FFD85438ADE7FB9FFEEAFBC8" bold="true" box="[189,325,1122,1149]" pageId="17" pageNumber="18">Field notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFD85438ADE7FB6FFCF8FA8E" blockId="17.[189,1399,1170,1339]" pageId="17" pageNumber="18">
Leaves bullate (
<emphasis id="B93DEAA0FFD85438AC24FB6FFDAAFB19" box="[382,517,1170,1196]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
), dark glossy green above, paler green beneath (
<emphasis id="B93DEAA0FFD85438A90CFB6FFB72FB19" box="[1110,1245,1170,1196]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
), shiny dark green above, slightly yellowish green below (
<emphasis id="B93DEAA0FFD85438AFBEFB4BFCE2FB7A" box="[740,845,1205,1231]" italics="true" pageId="17" pageNumber="18">KL 3033</emphasis>
); flowers yellowish (
<emphasis id="B93DEAA0FFD85438A914FB4BFB18FB7A" box="[1102,1207,1205,1231]" italics="true" pageId="17" pageNumber="18">KL 3033</emphasis>
), petals reddish orange (
<emphasis id="B93DEAA0FFD85438AC45FB24FE0BFB46" box="[287,420,1241,1267]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
), petals and sepals orange (
<emphasis id="B93DEAA0FFD85438AFB7FB27FCF1FB46" box="[749,862,1241,1267]" italics="true" pageId="17" pageNumber="18">FRI 2317</emphasis>
), stamens paler reddish orange (
<emphasis id="B93DEAA0FFD85438A9BAFB24FACAFB46" box="[1248,1381,1241,1267]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
), no scent with flowers (
<emphasis id="B93DEAA0FFD85438AC92FB00FDE3FAA2" box="[456,588,1277,1303]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
); fruits green (
<emphasis id="B93DEAA0FFD85438AFA7FB00FCC3FAA2" box="[765,876,1277,1303]" italics="true" pageId="17" pageNumber="18">FRI 3903</emphasis>
), fruits green turning red (
<emphasis id="B93DEAA0FFD85438A9FEFB00FA8FFAA2" box="[1188,1312,1277,1303]" italics="true" pageId="17" pageNumber="18">FRI 11853</emphasis>
), fruits red (
<emphasis id="B93DEAA0FFD85438ADAEFADCFEDEFA8F" box="[244,369,1312,1338]" italics="true" pageId="17" pageNumber="18">FRI 13467</emphasis>
, 
<emphasis id="B93DEAA0FFD85438ACDAFADCFDAAFA8F" box="[384,517,1312,1339]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
); seeds brown (
<emphasis id="B93DEAA0FFD85438AF9BFADCFCE9FA8F" box="[705,838,1312,1339]" italics="true" pageId="17" pageNumber="18">SFN 40941</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536539FFD8543BADE7FA9FFAC6FC7E" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BF636B2FFD85438ADE7FA9FFE41FAC9" blockId="17.[189,494,1378,1404]" box="[189,494,1378,1404]" pageId="17" pageNumber="18">
<heading id="D0BE81DEFFD85438ADE7FA9FFE41FAC9" bold="true" box="[189,494,1378,1404]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B93DEAA0FFD85438ADE7FA9FFE41FAC9" bold="true" box="[189,494,1378,1404]" pageId="17" pageNumber="18">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFD85438ADE7FA6CFD5AFA7B" blockId="17.[189,1398,1425,1487]" pageId="17" pageNumber="18">
Recorded from the eastern side of Peninsular 
<collectingCountry id="F35E7622FFD85438AF9DFA6CFC9CFA1E" box="[711,819,1425,1451]" name="Malaysia" pageId="17" pageNumber="18">Malaysia</collectingCountry>
from 
<collectingRegion id="498DF850FFD85438AE2DFA6CFC4EFA1E" box="[887,993,1425,1451]" country="Malaysia" name="Kelantan" pageId="17" pageNumber="18">Kelantan</collectingRegion>
and, with most records, Trengganu (
<figureCitation id="13722A37FFD85438AD9CFA49FEA1FA7A" box="[198,270,1460,1487]" captionStart="Fig" captionStartId="18.[189,232,2000,2026]" captionTargetBox="[189,1398,1355,1960]" captionTargetId="figure@18.[189,1398,1355,1960]" captionTargetPageId="18" captionText="Fig. 2. Map showing the collecting localities of Polyalthia pakdin I.M.Turner &amp; Utteridge sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3833129" httpUri="https://zenodo.org/record/3833129/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
). It occurs in lowland tropical rain forest.
</paragraph>
<paragraph id="8BF636B2FFD85438ADE7FA0BFE1AF9A4" blockId="17.[189,437,1526,1553]" box="[189,437,1526,1553]" pageId="17" pageNumber="18">
<heading id="D0BE81DEFFD85438ADE7FA0BFE1AF9A4" bold="true" box="[189,437,1526,1553]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B93DEAA0FFD85438ADE7FA0BFE1AF9A4" bold="true" box="[189,437,1526,1553]" pageId="17" pageNumber="18">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFD8543BADE7F9D8FDDBFEC0" blockId="17.[189,1399,1573,2027]" lastBlockId="18.[189,1399,267,373]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<taxonomicName id="4C494D31FFD85438ADE7F9D8FE23F98A" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[189,396,1573,1599]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFD85438ADE7F9D8FE23F98A" box="[189,396,1573,1599]" italics="true" pageId="17" pageNumber="18">Polyalthia pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFD85438ACCBF9DBFE45F9F5" box="[401,490,1574,1600]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is restricted to the lowland forests of eastern Peninsular 
<collectingCountry id="F35E7622FFD85438A936F9D8FB77F98A" box="[1132,1240,1573,1599]" name="Malaysia" pageId="17" pageNumber="18">Malaysia</collectingCountry>
and currently known from seven collections from five localities (two are from the Sekayu area and two from Jambu Bongkok). The Extent of Occurrence (EOO) is 
<quantity id="4CB19B57FFD85438AFBDF991FCFAF933" box="[743,853,1644,1671]" metricMagnitude="6" metricUnit="m" metricValue="1.8559999999999999" pageId="17" pageNumber="18" unit="km" value="1856.0">1,856 km</quantity>
<superScript id="7C3C9BFAFFD85438AE0FF996FCF1F9CC" attach="left" box="[853,862,1643,1657]" fontSize="6" pageId="17" pageNumber="18">2</superScript>
falling within the Endangered (EN) threshold of less than 
<quantity id="4CB19B57FFD85438AC16F96DFE14F91F" box="[332,443,1680,1707]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="5000.0">5,000 km</quantity>
<superScript id="7C3C9BFAFFD85438ACE1F972FE6BF928" attach="left" box="[443,452,1679,1693]" fontSize="6" pageId="17" pageNumber="18">2</superScript>
, and the Area of Occupancy (AOO; based on a user-defined cell width of 
<quantity id="4CB19B57FFD85438A86BF96DFAC1F91E" box="[1329,1390,1680,1707]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="2.0">2 km</quantity>
) is 
<quantity id="4CB19B57FFD85438AD82F949FE8EF97B" box="[216,289,1716,1742]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="17" pageNumber="18" unit="km" value="28.0">28 km</quantity>
<superScript id="7C3C9BFAFFD85438AC78F94FFE84F975" attach="left" box="[290,299,1714,1728]" fontSize="6" pageId="17" pageNumber="18">2</superScript>
falling within the Endangered (EN) threshold. Whilst none of the collections is contemporary (the most recent is 1972), nearly all are from forests reserves, and, in addition, one collection is from ‘roadside forest’ (
<emphasis id="B93DEAA0FFD85438ACD1F901FCCAF8A0" box="[395,869,1787,1813]" italics="true" pageId="17" pageNumber="18">
J. Sinclair &amp; 
<taxonomicName id="4C494D31FFD85438AF7DF906FD74F8A0" authority="bin Salleh" authorityName="bin Salleh" box="[551,731,1787,1813]" class="Insecta" family="Mogoplistidae" genus="Kiah" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Kiah bin Salleh</taxonomicName>
SFN 40941
</emphasis>
). Whilst the EOO is within the EN threshold, the presence of the species within protected areas would not place the species within this category because recent satellite imagery shows the protected areas still with relatively large areas of intact forest (using the most recent Google Earth imagery). The Sekayu collections, for example, are on the edge of the very poorly known and under-collected Taman Negara National Park and more field collections are needed in such areas to better understand the distribution of this species. Without any other information, we must assume that the species is still present in these protected areas and that the protected areas still harbour the appropriate habitat for 
<taxonomicName id="4C494D31FFDB543BAF0FFEF1FD6EFE90" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[597,705,267,293]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFDB543BAF0FFEF1FD6EFE90" box="[597,705,267,293]" italics="true" pageId="18" pageNumber="19">P. pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFDB543BAF9DFEF1FC8EFE93" box="[711,801,268,294]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
It is difficult to demonstrate trends in the reduction of habitat etc. without modern collections, and for these reasons we assign a preliminary conservation assessment of Near Threatened (NT).
</paragraph>
<paragraph id="8BF636B2FFDB543BADE7FE5CFE80FE0E" blockId="18.[189,303,417,443]" box="[189,303,417,443]" pageId="18" pageNumber="19">
<heading id="D0BE81DEFFDB543BADE7FE5CFE80FE0E" bold="true" box="[189,303,417,443]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B93DEAA0FFDB543BADE7FE5CFE80FE0E" bold="true" box="[189,303,417,443]" pageId="18" pageNumber="19">Affinities</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B2FFDB543BADE7FE29FAC6FC7E" blockId="18.[189,1399,468,971]" pageId="18" pageNumber="19">
<taxonomicName id="4C494D31FFDB543BADE7FE29FE3EFE5B" authorityName="King" authorityYear="1892" box="[189,401,468,494]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="bullata">
<emphasis id="B93DEAA0FFDB543BADE7FE29FE3EFE5B" box="[189,401,468,494]" italics="true" pageId="18" pageNumber="19">Polyalthia bullata</emphasis>
</taxonomicName>
, the species with which 
<taxonomicName id="4C494D31FFDB543BAFF6FE28FCB7FE5B" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[684,792,468,494]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFDB543BAFF6FE28FCB7FE5B" box="[684,792,468,494]" italics="true" pageId="18" pageNumber="19">P. pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFDB543BAE44FE29FCD7FE5B" box="[798,888,468,494]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
was formerly confused, was included in the informal group of 
<taxonomicName id="4C494D31FFDB543BACC8FE01FDA3FDA3" authorityName="Blume" authorityYear="1830" box="[402,524,508,534]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B93DEAA0FFDB543BACC8FE01FDA3FDA3" box="[402,524,508,534]" italics="true" pageId="18" pageNumber="19">Polyalthia</emphasis>
</taxonomicName>
species termed the 
<taxonomicName id="4C494D31FFDB543BAFB5FE01FC65FDA0" box="[751,970,508,534]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="insignis">
<emphasis id="B93DEAA0FFDB543BAFB5FE01FC65FDA0" box="[751,970,508,534]" italics="true" pageId="18" pageNumber="19">Polyalthia insignis</emphasis>
</taxonomicName>
group by 
<bibRefCitation id="EFD84B43FFDB543BA965FE06FADCFDA3" author="Johnson D. M. &amp; Murray N. A." box="[1087,1395,507,534]" pageId="18" pageNumber="19" pagination="95 - 104" refId="ref14398" refString="Johnson D. M. &amp; Murray N. A. 1999. Four new species of Polyalthia (Annonaceae) from Borneo and their relationship to Polyalthia insignis. Contributions from the University of Michigan Herbarium 22: 95 - 104." type="journal article" year="1999">Johnson &amp; Murray (1999)</bibRefCitation>
. They characterised the group as follows: shrubs or small trees; leaves oblique, subcordate or auriculate at base, often drying grey above, brown below; inflorescences internodal to leaf opposed or sometimes cauliflorous; petals orange or red; apex of ovary woody and persistant as a short beak on the monocarp; stigmas capitate and cohering to drop as a cap at the end of anthesis, several ovules per carpel, laterally attached in vertical row and seed not pitted but with encircling groove. While 
<taxonomicName id="4C494D31FFDB543BA91CFD3FFAB8FD69" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[1094,1303,706,732]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFDB543BA91CFD3FFAB8FD69" box="[1094,1303,706,732]" italics="true" pageId="18" pageNumber="19">Polyalthia pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFDB543BA847FD3EFAD8FD68" box="[1309,1399,707,733]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
does exhibit some of these characters, it does not have auriculate or cordate leaf bases and the stigmas do not seem to cohere. In fact, 
<taxonomicName id="4C494D31FFDB543BAF7AFCEEFD23FC99" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[544,652,786,812]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFDB543BAF7AFCEEFD23FC99" box="[544,652,786,812]" italics="true" pageId="18" pageNumber="19">P. pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFDB543BAFC8FCEFFD44FC99" box="[658,747,786,812]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
also comes close to the Bornean 
<taxonomicName id="4C494D31FFDB543BA931FCEFFF54FCE6" authority="Diels" authorityName="Diels" authorityYear="1931" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="polyphlebia">
<emphasis id="B93DEAA0FFDB543BA931FCEFFAD8FC99" box="[1131,1399,786,812]" italics="true" pageId="18" pageNumber="19">Polyalthia polyphlebia</emphasis>
Diels
</taxonomicName>
which was not included in the 
<taxonomicName id="4C494D31FFDB543BAF32FCC7FD4FFCE6" box="[616,736,826,851]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="insignis">
<emphasis id="B93DEAA0FFDB543BAF32FCC7FD4FFCE6" box="[616,736,826,851]" italics="true" pageId="18" pageNumber="19">P. insignis</emphasis>
</taxonomicName>
group by Johnson &amp; Murray. The leaf form of 
<taxonomicName id="4C494D31FFDB543BA850FCC7FAD8FCE6" authority="Turner &amp; Utteridge, 2016" authorityName="I. M. Turner &amp; Utteridge" authorityYear="2016" box="[1290,1399,825,851]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="pakdin" status="sp. nov.">
<emphasis id="B93DEAA0FFDB543BA850FCC7FAD8FCE6" box="[1290,1399,825,851]" italics="true" pageId="18" pageNumber="19">P. pakdin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DBFFDB543BADE7FC9FFEB4FCC9" box="[189,283,866,892]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C494D31FFDB543BAC01FC9FFDA9FCCE" authorityName="Diels" authorityYear="1931" box="[347,518,865,891]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="polyphlebia">
<emphasis id="B93DEAA0FFDB543BAC01FC9FFDA9FCCE" box="[347,518,865,891]" italics="true" pageId="18" pageNumber="19">P. polyphlebia</emphasis>
</taxonomicName>
are similar but 
<taxonomicName id="4C494D31FFDB543BAF91FC9FFCD8FCCE" authorityName="Diels" authorityYear="1931" box="[715,887,865,891]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="polyphlebia">
<emphasis id="B93DEAA0FFDB543BAF91FC9FFCD8FCCE" box="[715,887,865,891]" italics="true" pageId="18" pageNumber="19">P. polyphlebia</emphasis>
</taxonomicName>
generally has greyish corky twigs, longer pedicels (8–15 versus 
<quantity id="4CB19B57FFDB543BACE5FC74FD8EFC16" box="[447,545,905,931]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">2–8 mm</quantity>
) with the flowers larger (outer petals 
<quantity id="4CB19B57FFDB543BAE8AFC74FBE2FC16" box="[976,1101,905,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="18" pageNumber="19" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
versus 
<quantity id="4CB19B57FFDB543BA9FEFC74FABAFC16" box="[1188,1301,905,931]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="18" pageNumber="19" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
) and the petals fleshy rather than thin in texture; with the monocarps on longer stipes (to 
<quantity id="4CB19B57FFDB543BA932FC4DFB15FC7E" box="[1128,1210,944,971]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="25.0">25 mm</quantity>
versus 
<quantity id="4CB19B57FFDB543BA849FC4DFAF3FC7E" box="[1299,1372,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="6.0">6 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>