treatments-xml/data/03/E0/87/03E087A1E554FFCC88BE9C9C62A87A69.xml

<document id="373E0B4693ED451DDCD7F83BFF9329C4" ID="10.11646/zootaxa.4147.1.4" ID-DOI="10.11646/zootaxa.4147.1.4" ID-GBIF-Dataset="3166e193-d052-4efa-953b-4e5ac1b6f568" ID-ISSN="1175-5326" ID-Zenodo-Dep="266251" ID-ZooBank="39ECCC38-BC63-40D2-9B04-F435841AA0D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470120521773" checkinUser="plazi" docAuthor="Daniel, Gimo M., Davis, Adrian Lv. &amp; Scholtz, Clarke H." docDate="2016" docId="03E087A1E554FFCC88BE9C9C62A87A69" docLanguage="en" docName="zootaxa.4147.1.4.pdf" docOrigin="Zootaxa 4147 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sisyphus oralensis Daniel &amp; Davis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="71" masterDocId="FFD9FFD9E557FFC888299C0B66117C33" masterDocTitle="Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa" masterLastPageNumber="74" masterPageNumber="67" pageNumber="70" updateTime="1698366689639" updateUser="plazi">
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<mods:title id="CF24D8B212CC4731E1DFC262FF94FF78">Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa</mods:title>
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<mods:namePart id="7A67F663F871FBB2254B88ACE5FBCE07">Daniel, Gimo M.</mods:namePart>
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<mods:namePart id="45EF68DA0D13ED4D6EEA373864C0140C">Davis, Adrian Lv.</mods:namePart>
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<mods:namePart id="3664A200FECE81574D822A997B35C0AB">Scholtz, Clarke H.</mods:namePart>
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<mods:title id="69B54FA7F4716BB239F9174AC6F99036">Zootaxa</mods:title>
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<mods:date id="7427312180198527FC093D421B17A377">2016</mods:date>
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<mods:number id="3508BB049D0548915AEBAE4AC2214008">4147</mods:number>
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<mods:identifier id="6D572988EC5223D76F69AC3983AFD8ED" type="DOI">10.11646/zootaxa.4147.1.4</mods:identifier>
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<treatment id="03E087A1E554FFCC88BE9C9C62A87A69" ID-DOI="http://doi.org/10.5281/zenodo.6087201" ID-GBIF-Taxon="123513975" ID-Zenodo-Dep="6087201" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69" httpUri="http://treatment.plazi.org/id/03E087A1E554FFCC88BE9C9C62A87A69" lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70">
<subSubSection id="C353653CE554FFCB88BE9C9C66E67CE0" pageId="3" pageNumber="70" type="nomenclature">
<paragraph id="8BF636B7E554FFCB88BE9C9C64B07C81" blockId="3.[151,673,151,211]" box="[151,673,151,178]" pageId="3" pageNumber="70">
<heading id="D0BE81DBE554FFCB88BE9C9C64B07C81" bold="true" box="[151,673,151,178]" fontSize="11" level="1" pageId="3" pageNumber="70" reason="1">
<emphasis id="B93DEAA5E554FFCB88BE9C9C64B07C81" bold="true" box="[151,673,151,178]" pageId="3" pageNumber="70">
<taxonomicName id="4C494D34E554FFCB88BE9C9C642D7C81" ID-CoL="7ZRH7" authority="Daniel &amp; Davis" authorityName="Daniel &amp; Davis" box="[151,572,151,178]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB88BE9C9C67667C82" bold="true" box="[151,375,151,177]" italics="true" pageId="3" pageNumber="70">Sisyphus oralensis</emphasis>
Daniel &amp; Davis
</taxonomicName>
<taxonomicNameLabel id="A20E57DEE554FFCB8A6D9C9364B07C81" box="[580,673,152,178]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B7E554FFCB88BE9CB166E67CE0" blockId="3.[151,673,151,211]" box="[151,247,186,211]" pageId="3" pageNumber="70">
(
<figureCitation id="13722A32E554FFCB88B69CB166FE7CE0" box="[159,239,186,211]" captionStart="FIGURE 2. A. S" captionStartId="3.[151,250,1161,1183]" captionTargetBox="[394,1198,429,1144]" captionTargetId="figure@3.[340,1248,429,1149]" captionTargetPageId="3" captionText="FIGURE 2. A. S. oralensis sp. nov. habitus. Diagnostic characters: Bi. Gena rounded forming obtuse angle with genal suture. Bii. Patterned arrangement of setae on pronotal disc separated by bare patches. C. Aedeagus." httpUri="https://zenodo.org/record/266253/files/figure.png" pageId="3" pageNumber="70">Fig. 2a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C353653CE554FFCB88BE9D09623D7DB4" pageId="3" pageNumber="70" type="diagnosis">
<paragraph id="8BF636B7E554FFCB88BE9D09623D7DB4" blockId="3.[151,1436,258,391]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88BE9D0967007D28" bold="true" box="[151,273,258,283]" pageId="3" pageNumber="70">Diagnosis.</emphasis>
<taxonomicName id="4C494D34E554FFCB89359D0867E07D28" box="[284,497,259,283]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB89359D0867E07D28" box="[284,497,259,283]" italics="true" pageId="3" pageNumber="70">Sisyphus oralensis</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E554FFCB89D59D09644B7D28" bold="true" box="[508,602,258,283]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A20E57DEE554FFCB89D59D09644B7D28" box="[508,602,258,283]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="4C494D34E554FFCB8AD69D0865BC7D28" box="[767,941,259,283]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="umbraphilus" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB8AD69D0865BC7D28" box="[767,941,259,283]" italics="true" pageId="3" pageNumber="70">S. umbraphilus</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E554FFCB8B909D0962067D28" bold="true" box="[953,1047,258,283]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A20E57DEE554FFCB8B909D0962067D28" box="[953,1047,258,283]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished by the following combination of characters: genae rounded forming obtuse angle with genal suture; less dense cover of long setae on pronotal disc, not distributed uniformly but mostly arranged in a linear pattern separated by bare patches; 
<taxonomicName id="4C494D34E554FFCB88D39D64676D7DB5" box="[250,380,366,390]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB88D39D64676D7DB5" box="[250,380,366,390]" italics="true" pageId="3" pageNumber="70">S. oralensis</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E554FFCB89AA9D6567CC7DB4" bold="true" box="[387,477,366,391]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A20E57DEE554FFCB89AA9D6567CC7DB4" box="[387,477,366,391]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is smaller than 
<taxonomicName id="4C494D34E554FFCB8AA79D6465267DB5" box="[654,823,366,390]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="umbraphilus" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB8AA79D6465267DB5" box="[654,823,366,390]" italics="true" pageId="3" pageNumber="70">S. umbraphilus</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E554FFCB8B179D6565897DB4" bold="true" box="[830,920,366,391]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A20E57DEE554FFCB8B179D6565897DB4" box="[830,920,366,391]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B93DEAA5E554FFCB8B819D65620A7DB4" bold="true" box="[936,1051,366,391]" pageId="3" pageNumber="70">
<figureCitation id="13722A32E554FFCB8B819D6565FB7DB4" box="[936,1002,366,391]" captionStart="FIGURE 2. A. S" captionStartId="3.[151,250,1161,1183]" captionTargetBox="[394,1198,429,1144]" captionTargetId="figure@3.[340,1248,429,1149]" captionTargetPageId="3" captionText="FIGURE 2. A. S. oralensis sp. nov. habitus. Diagnostic characters: Bi. Gena rounded forming obtuse angle with genal suture. Bii. Patterned arrangement of setae on pronotal disc separated by bare patches. C. Aedeagus." httpUri="https://zenodo.org/record/266253/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
bi-ii
</emphasis>
).
</paragraph>
</subSubSection>
<caption id="DF36663FE554FFCB88BE98826251788D" httpUri="https://zenodo.org/record/266253/files/figure.png" pageId="3" pageNumber="70" targetBox="[394,1198,429,1144]" targetPageId="3">
<paragraph id="8BF636B7E554FFCB88BE98826251788D" blockId="3.[151,1437,1161,1214]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88BE9882672378AC" bold="true" box="[151,306,1161,1183]" pageId="3" pageNumber="70">FIGURE 2. A.</emphasis>
<taxonomicName id="4C494D34E554FFCB8910988167A178AC" box="[313,432,1161,1183]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E554FFCB8910988167A178AC" box="[313,432,1161,1183]" italics="true" pageId="3" pageNumber="70">S. oralensis</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E554FFCB899F9882641678AC" bold="true" box="[438,519,1161,1183]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A20E57DEE554FFCB899F9882641678AC" box="[438,519,1161,1183]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
habitus. Diagnostic characters: 
<emphasis id="B93DEAA5E554FFCB8B649882657D78AC" bold="true" box="[845,876,1161,1183]" pageId="3" pageNumber="70">Bi.</emphasis>
Gena rounded forming obtuse angle with genal suture. 
<emphasis id="B93DEAA5E554FFCB88BE98A366A7788D" bold="true" box="[151,182,1192,1214]" pageId="3" pageNumber="70">Bii</emphasis>
. Patterned arrangement of setae on pronotal disc separated by bare patches. 
<emphasis id="B93DEAA5E554FFCB8B9398A365DF788D" bold="true" box="[954,974,1192,1214]" pageId="3" pageNumber="70">C.</emphasis>
Aedeagus.
</paragraph>
</caption>
<subSubSection id="C353653CE554FFCB88EE98EE655C7BC1" pageId="3" pageNumber="70" type="description">
<paragraph id="8BF636B7E554FFCB88EE98EE66E57912" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE98EE67F778CD" bold="true" box="[199,486,1253,1278]" pageId="3" pageNumber="70">Description. Size: Male:</emphasis>
Length: 
<quantity id="4CB19B52E554FFCB8A6298ED64C578CE" box="[587,724,1254,1278]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.8" metricValueMin="3.4" pageId="3" pageNumber="70" unit="mm" value="4.1" valueMax="4.8" valueMin="3.4">3.4–4.8 mm</quantity>
, width: 
<quantity id="4CB19B52E554FFCB8B0798EE65A978CE" box="[814,952,1253,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="3" pageNumber="70" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1–2.5 mm</quantity>
. 
<emphasis id="B93DEAA5E554FFCB8BEA98EE620B78CD" bold="true" box="[963,1050,1253,1278]" pageId="3" pageNumber="70">Female</emphasis>
: Length: 
<quantity id="4CB19B52E554FFCB8CAE98ED630178CE" box="[1159,1296,1254,1278]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="3" pageNumber="70" unit="mm" value="4.3" valueMax="4.7" valueMin="3.9">3.9–4.7 mm</quantity>
, width: 
<quantity id="4CB19B52E554FFCB8D4298ED66E17912" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="3" pageNumber="70" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1– 2.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE992664097975" blockId="3.[151,1437,1253,2034]" box="[199,536,1325,1350]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9926670A7975" bold="true" box="[199,283,1325,1350]" pageId="3" pageNumber="70">Colour</emphasis>
: Dark brown to black.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE995A678F79E5" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE995A67017959" bold="true" box="[199,272,1361,1386]" pageId="3" pageNumber="70">Head:</emphasis>
Clypeus with a strong cleft between two medial teeth, deep excavation lateral to each tooth, lateral convex sinuosity between excavation and genal suture, genae rounded forming obtuse angle with genal suture; frons finely punctate and setose, post-occipital suture fully rounded and arciform, eyes reniforme with dark ocular stains, antennae brown.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE99EA63227A71" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE99EA675879C9" bold="true" box="[199,329,1505,1530]" pageId="3" pageNumber="70">Pronotum:</emphasis>
Convex, dark brown, coarsely punctate; anteriorly inclined; central diameter wider than elytra; complete lateral prothoracic ridge separating prothoracic disc and prothoracic episternum; long setae of pronotal disc not uniformly spaced but forming a clear, mostly linear pattern separated by bare patches (
<emphasis id="B93DEAA5E554FFCB8CE99A2263337A71" bold="true" box="[1216,1314,1577,1602]" pageId="3" pageNumber="70">
<figureCitation id="13722A32E554FFCB8CE99A2263137A71" box="[1216,1282,1577,1602]" captionStart="FIGURE 2. A. S" captionStartId="3.[151,250,1161,1183]" captionTargetBox="[394,1198,429,1144]" captionTargetId="figure@3.[340,1248,429,1149]" captionTargetPageId="3" captionText="FIGURE 2. A. S. oralensis sp. nov. habitus. Diagnostic characters: Bi. Gena rounded forming obtuse angle with genal suture. Bii. Patterned arrangement of setae on pronotal disc separated by bare patches. C. Aedeagus." httpUri="https://zenodo.org/record/266253/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
bii
</emphasis>
).
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE9A4663447A55" blockId="3.[151,1437,1253,2034]" box="[199,1365,1613,1638]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9A46670D7A55" bold="true" box="[199,284,1613,1638]" pageId="3" pageNumber="70">Elytra:</emphasis>
Dark brown; narrowed posteriorly; finely punctate; well-spaced; fine curled setae on interstriae.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE9A7A63647ABA" blockId="3.[151,1437,1253,2034]" box="[199,1397,1649,1674]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9A7A672B7AB9" bold="true" box="[199,314,1649,1674]" pageId="3" pageNumber="70">Pygidium</emphasis>
: Dark brown; strongly narrowed and &quot;v&quot; shaped towards apical base; finely punctate and setose.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE9A9E62D07AE2" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9A9E67287A9D" bold="true" box="[199,313,1685,1710]" pageId="3" pageNumber="70">Sternites:</emphasis>
Abdominal sternites punctate with fine setae arranged in rows; meso- and metasternum black, metallic shiny, and finely punctate; mesepimerum and metepisternum: punctate with fine setae.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE9AD667657BB5" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9AD667197AC5" bold="true" box="[199,264,1757,1782]" pageId="3" pageNumber="70">Legs:</emphasis>
Profemur with well-developed setae laterally, punctate ventrally; meso- and metafemur brown, punctate and setose; meso- and metatrochanter contiguous with femur, forming a divergent spine at the tip in latter; promeso- and metatibia setose with single terminal spur, protibia with three teeth; pro- meso- and metatarsus five segmented with two claws; the first tarsal segment of the meso- and meta-thoracic legs with a row of strong setae on the ventral edge.
</paragraph>
<paragraph id="8BF636B7E554FFCB88EE9B9A655C7BC1" blockId="3.[151,1437,1253,2034]" pageId="3" pageNumber="70">
<emphasis id="B93DEAA5E554FFCB88EE9B9A67527B99" bold="true" box="[199,323,1937,1962]" pageId="3" pageNumber="70">Aedeagus:</emphasis>
Phallobase curved dorsally, forming a strong angle with parameres, visible medial line ventrally; parameres nearly symmetrical, separated by membranous area visible dorsally on apical edge; basal edges slightly excavated longitudinally both ventrally and dorsally. (
<emphasis id="B93DEAA5E554FFCB8ADD9BD265527BC1" bold="true" box="[756,835,2009,2034]" pageId="3" pageNumber="70">
<figureCitation id="13722A32E554FFCB8ADD9BD265267BC1" box="[756,823,2009,2034]" captionStart="FIGURE 2. A. S" captionStartId="3.[151,250,1161,1183]" captionTargetBox="[394,1198,429,1144]" captionTargetId="figure@3.[340,1248,429,1149]" captionTargetPageId="3" captionText="FIGURE 2. A. S. oralensis sp. nov. habitus. Diagnostic characters: Bi. Gena rounded forming obtuse angle with genal suture. Bii. Patterned arrangement of setae on pronotal disc separated by bare patches. C. Aedeagus." httpUri="https://zenodo.org/record/266253/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
c
</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="C353653CE553FFCC88EE9C9C62007CE6" pageId="4" pageNumber="71" type="etymology">
<paragraph id="8BF636B7E553FFCC88EE9C9C62007CE6" blockId="4.[151,1436,151,1626]" pageId="4" pageNumber="71">
<emphasis id="B93DEAA5E553FFCC88EE9C9C675F7C83" bold="true" box="[199,334,151,176]" pageId="4" pageNumber="71">Etymology.</emphasis>
We named this species 
<taxonomicName id="4C494D34E553FFCC8A4F9C9364FD7C83" box="[614,748,152,176]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="oralensis">
<emphasis id="B93DEAA5E553FFCC8A4F9C9364FD7C83" box="[614,748,152,176]" italics="true" pageId="4" pageNumber="71">S. oralensis</emphasis>
</taxonomicName>
because it is found on the Indian Ocean coastline of 
<collectingCountry id="F35E7627E553FFCC8D729C9C66CE7CE6" name="South Africa" pageId="4" pageNumber="71">South Africa</collectingCountry>
and 
<collectingCountry id="F35E7627E553FFCC893C9CB767B87CE6" box="[277,425,188,213]" name="Mozambique" pageId="4" pageNumber="71">Mozambique</collectingCountry>
where it is associated with dense woodland and forest.
</paragraph>
</subSubSection>
<subSubSection id="C353653CE553FFCC88EE9CE966F179DD" pageId="4" pageNumber="71" type="materials_examined">
<paragraph id="8BF636B7E553FFCC88EE9CE964FC7909" blockId="4.[151,1436,151,1626]" pageId="4" pageNumber="71">
<materialsCitation id="3B213CEAE553FFCC88EE9CE9651C7D11" ID-GBIF-Occurrence="1290984948" collectingDate="2000-01-27" collectionCode="TMSA" country="South Africa" location="Richard's Bay" municipality="Examined" pageId="4" pageNumber="71" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC88EE9CE9643D7CC8" bold="true" box="[199,556,226,251]" pageId="4" pageNumber="71">
<collectingMunicipality id="6B92ACCDE553FFCC88EE9CE967517CC8" box="[199,320,226,251]" pageId="4" pageNumber="71">Examined</collectingMunicipality>
material. 
<typeStatus id="54F28815E553FFCC89969CE9643D7CC8" box="[447,556,226,251]" pageId="4" pageNumber="71" type="holotype">Holotype</typeStatus>
</emphasis>
: (
<specimenCount id="9D4FFD3EE553FFCC8A6F9CE8644A7CC8" box="[582,603,227,251]" pageId="4" pageNumber="71" type="male">♂</specimenCount>
<emphasis id="B93DEAA5E553FFCC8A4D9CE964A47CC8" bold="true" box="[612,693,226,251]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8A4D9CE964A47CC8" box="[612,693,226,251]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
</emphasis>
): 
<emphasis id="B93DEAA5E553FFCC8AE69CE965B97CC8" bold="true" box="[719,936,226,251]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8AE69CE965B97CC8" box="[719,936,226,251]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC8C2B9CE962B27CC8" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8C2B9CE962B27CC8" box="[1026,1187,226,251]" country="South Africa" municipality="Examined" name="Richard's Bay" pageId="4" pageNumber="71">Richard’s Bay</location>
28[◦] 39[’] 29.1[”] S; 32[◦] 15[’] 19.2[”] E, 
<date id="FFF71077E553FFCC89BF9D0264137D11" box="[406,514,265,290]" pageId="4" pageNumber="71" value="2000-01-27">
<collectingDate id="EFB3E99FE553FFCC89BF9D0264137D11" box="[406,514,265,290]" pageId="4" pageNumber="71" value="2000-01-27">27.i.2000</collectingDate>
</date>
, leg: Davis &amp; Delport
</materialsCitation>
. 
<emphasis id="B93DEAA5E553FFCC8B339D0265837D11" bold="true" box="[794,914,265,290]" pageId="4" pageNumber="71">Paratypes</emphasis>
: (5♂, 8♀ 
<emphasis id="B93DEAA5E553FFCC8C399D0263407D11" bold="true" box="[1040,1361,265,290]" pageId="4" pageNumber="71">
UPSA): 
<materialsCitation id="3B213CEAE553FFCC8C5E9D0263407D11" ID-GBIF-Occurrence="1290984945" box="[1143,1361,265,290]" collectingDate="2000-01-26" collectingDateMax="2000-01-27" collectingDateMin="2000-01-26" collectionCode="UPSA" collectorName="Davis &amp; Delport." country="South Africa" location="Richard's Bay" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8C5E9D0263407D11" box="[1143,1361,265,290]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</materialsCitation>
</emphasis>
, KZN 
<location id="8E96606CE553FFCC88BE9D3B672C7D7A" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC88BE9D3B672C7D7A" box="[151,317,304,329]" country="South Africa" name="Richard's Bay" pageId="4" pageNumber="71">Richard’s Bay</location>
28[◦] 39[’] S; 32[◦] 15[’] E, 
<date id="FFF71077E553FFCC8A8C9D3B65267D7A" box="[677,823,304,329]" pageId="4" pageNumber="71" value="2000-01-26" valueMax="2000-01-27" valueMin="2000-01-26">
<collectingDate id="EFB3E99FE553FFCC8A8C9D3B65267D7A" box="[677,823,304,329]" pageId="4" pageNumber="71" value="2000-01-26" valueMax="2000-01-27" valueMin="2000-01-26">26-27.i.2000</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8BA89D3B65D37D7A" box="[897,962,304,329]" pageId="4" pageNumber="71">Davis</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8BDD9D3B62407D7A" box="[1012,1105,304,329]" pageId="4" pageNumber="71">Delport.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8C419D3A629C7D7A" box="[1128,1165,305,329]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8CB69D3A62D37D7A" box="[1183,1218,305,329]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8CF99D3B67157D43" bold="true" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8CF99D3B630E7D7A" box="[1232,1311,304,329]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="4" pageNumber="71">UPSA</collectionCode>
): 
<materialsCitation id="3B213CEAE553FFCC8D139D3B67157D43" ID-GBIF-Occurrence="1290984929" collectingDate="1999-01-13" collectionCode="UPSA, SANC" collectorName="Chown &amp; McGeogh &amp; Davis." country="South Africa" location="Itala Game Reserve" pageId="4" pageNumber="71" specimenCount="9" specimenCount-female="6" specimenCount-male="3" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8D139D3B67157D43" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</materialsCitation>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC89739D5C64297D5C" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC89739D5C64297D5C" box="[346,568,343,368]" country="South Africa" name="Itala Game Reserve" pageId="4" pageNumber="71">Itala Game Reserve</location>
27[◦] 27[’] 53.6[”] S; 31[◦] 15[’] 36.1[”] E, 
<date id="FFF71077E553FFCC8C0A9D5C629E7D43" box="[1059,1167,343,368]" pageId="4" pageNumber="71" value="1999-01-13">
<collectingDate id="EFB3E99FE553FFCC8C0A9D5C629E7D43" box="[1059,1167,343,368]" pageId="4" pageNumber="71" value="1999-01-13">13.i.1999</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8CE59D5C63317D43" box="[1228,1312,343,368]" pageId="4" pageNumber="71">Chown</collectorName>
, 
<collectorName id="26BC5361E553FFCC8D059D5C638D7D43" box="[1324,1436,343,368]" pageId="4" pageNumber="71">McGeogh</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC889D9D7666EC7DA5" box="[180,253,381,406]" pageId="4" pageNumber="71">Davis.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC89279D7567207DA5" box="[270,305,382,406]" pageId="4" pageNumber="71" type="female">4♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89109D7667937DA5" bold="true" box="[313,386,381,406]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC89109D7667937DA5" box="[313,386,381,406]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="4" pageNumber="71">UPSA</collectionCode>
</emphasis>
; 
<specimenCount id="9D4FFD3EE553FFCC89BB9D7567A67DA5" box="[402,439,382,406]" pageId="4" pageNumber="71" type="male">3♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC89EA9D7567F77DA5" box="[451,486,382,406]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89C79D76653D7DA5" bold="true" box="[494,812,381,406]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC89C79D7664517DA5" box="[494,576,381,406]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
): 
<materialsCitation id="3B213CEAE553FFCC8A7C9D76653D7DA5" ID-GBIF-Occurrence="1290984947" box="[597,812,381,406]" collectingDate="1996-12-17" collectionCode="UPSA" collectorName="B. J. Van Rensburg." country="South Africa" location="Thembe Elephant Park" pageId="4" pageNumber="71" specimenCount="17" specimenCount-female="8" specimenCount-male="9" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8A7C9D76653D7DA5" box="[597,812,381,406]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</materialsCitation>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC8BAA9D7662947DA5" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8BAA9D7662947DA5" box="[899,1157,381,406]" country="South Africa" name="Thembe Elephant Park" pageId="4" pageNumber="71">Thembe Elephant Park</location>
27[◦] 01[’] S; 32[◦] 24[’] E, 
<date id="FFF71077E553FFCC88919DAF672B7D8E" box="[184,314,420,445]" pageId="4" pageNumber="71" value="1996-12-17">
<collectingDate id="EFB3E99FE553FFCC88919DAF672B7D8E" box="[184,314,420,445]" pageId="4" pageNumber="71" value="1996-12-17">17.xii.1996</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89539DAE64447D8E" box="[378,597,420,445]" pageId="4" pageNumber="71">B.J. Van Rensburg.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8A4E9DAE649D7D8E" box="[615,652,421,445]" pageId="4" pageNumber="71" type="male">9♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8AB39DAE64AD7D8E" box="[666,700,421,445]" pageId="4" pageNumber="71" type="female">8♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8AEC9DAF62137D8E" bold="true" box="[709,1026,420,445]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8AEC9DAF65057D8E" box="[709,788,420,445]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="4" pageNumber="71">UPSA</collectionCode>
): 
<materialsCitation id="3B213CEAE553FFCC8B039DAF62137D8E" ID-GBIF-Occurrence="1290984949" box="[810,1026,420,445]" collectingDate="1995-07-17" collectionCode="UPSA" collectorName="B. J. Van Rensburg." country="South Africa" location="Thembe Elephant Park" pageId="4" pageNumber="71" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8B039DAF62137D8E" box="[810,1026,420,445]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</materialsCitation>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC8C729DAF63717D8E" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8C729DAF63717D8E" box="[1115,1376,420,445]" country="South Africa" name="Thembe Elephant Park" pageId="4" pageNumber="71">Thembe Elephant Park</location>
27[◦] 01[’] S; 32[◦] 24[’] E, 
<date id="FFF71077E553FFCC89B29DC0640C7DD7" box="[411,541,459,484]" pageId="4" pageNumber="71" value="1995-07-17">
<collectingDate id="EFB3E99FE553FFCC89B29DC0640C7DD7" box="[411,541,459,484]" pageId="4" pageNumber="71" value="1995-07-17">17.vii.1995</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8A769DC7652C7DD7" box="[607,829,459,484]" pageId="4" pageNumber="71">B.J. Van Rensburg.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8B679DC765657DD7" box="[846,884,460,484]" pageId="4" pageNumber="71" type="male">2♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8BAA9DC765B47DD7" box="[899,933,460,484]" pageId="4" pageNumber="71" type="female">3♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B869DC062137DD7" bold="true" box="[943,1026,459,484]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B869DC065EB7DD7" box="[943,1018,459,484]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="4" pageNumber="71">UPSA</collectionCode>
)
</emphasis>
: 
<materialsCitation id="3B213CEAE553FFCC8C3A9DC063847E38" ID-GBIF-Occurrence="1290984941" collectingDate="2008-10-15" collectingDateMax="2008-10-30" collectingDateMin="2008-10-15" collectionCode="SANC, TMSA" collectorName="B. J. Van Rensburg." country="South Africa" location="Thembe Elephant Park" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8C3A9DC062FD7DD7" bold="true" box="[1043,1260,459,484]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8C3A9DC062FD7DD7" box="[1043,1260,459,484]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN 
<location id="8E96606CE553FFCC8D699DC0672B7E38" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8D699DC0672B7E38" country="South Africa" name="Thembe Elephant Park" pageId="4" pageNumber="71">Thembe Elephant Park</location>
26[◦] 55[’] S; 32[◦] 23[’] E, 
<date id="FFF71077E553FFCC8AB19DF965217E38" box="[664,816,498,523]" pageId="4" pageNumber="71" value="2008-10-15" valueMax="2008-10-30" valueMin="2008-10-15">
<collectingDate id="EFB3E99FE553FFCC8AB19DF965217E38" box="[664,816,498,523]" pageId="4" pageNumber="71" value="2008-10-15" valueMax="2008-10-30" valueMin="2008-10-15">15-30.x.2008</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8B519DF8624B7E38" box="[888,1114,498,523]" pageId="4" pageNumber="71">B.J. Van Rensburg.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8C479DF862837E38" box="[1134,1170,499,523]" pageId="4" pageNumber="71" type="male">2♂</specimenCount>
<emphasis id="B93DEAA5E553FFCC8CB79DF9631D7E38" bold="true" box="[1182,1292,498,523]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8CB79DF962FD7E38" box="[1182,1260,498,523]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
, 
<specimenCount id="9D4FFD3EE553FFCC8CD79DF9631D7E38" box="[1278,1292,498,523]" pageId="4" pageNumber="71" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="B93DEAA5E553FFCC8D139DF963847E38" bold="true" box="[1338,1429,498,523]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8D139DF9639C7E38" box="[1338,1421,498,523]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC88BE9E1266FD7E6B" ID-GBIF-Occurrence="1290984934" collectingDate="1971-08-25" collectionCode="SANC" collectorName="Bornemissza &amp; Insley." country="South Africa" location="Umfolozi" pageId="4" pageNumber="71" specimenCount="11" specimenCount-female="10" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC88BE9E12677C7E01" bold="true" box="[151,365,537,562]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC88BE9E12677C7E01" box="[151,365,537,562]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC89EB9E12643F7E01" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC89EB9E12643F7E01" box="[450,558,537,562]" country="South Africa" name="Umfolozi" pageId="4" pageNumber="71">Umfolozi</location>
28[◦] 15[’] S; 31[◦] 55[’] E, 
<date id="FFF71077E553FFCC8B4C9E1265E17E01" box="[869,1008,537,562]" pageId="4" pageNumber="71" value="1971-08-25">
<collectingDate id="EFB3E99FE553FFCC8B4C9E1265E17E01" box="[869,1008,537,562]" pageId="4" pageNumber="71" value="1971-08-25">25.viii.1971</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8C059E1262AC7E01" box="[1068,1213,537,562]" pageId="4" pageNumber="71">Bornemissza</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8CC99E12633B7E01" box="[1248,1322,537,562]" pageId="4" pageNumber="71">Insley.</collectorName>
<emphasis id="B93DEAA5E553FFCC8D189E12632B7E01" bold="true" box="[1329,1338,537,562]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8D139E12634C7E02" box="[1338,1373,537,561]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8D429E12638A7E02" box="[1387,1435,537,561]" pageId="4" pageNumber="71" type="female">10♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC88BE9E3466FD7E6B" bold="true" box="[151,236,575,600]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC88BE9E3466F47E6B" box="[151,229,575,600]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC88D59E3467E37E4C" ID-GBIF-Occurrence="1290984939" collectingDate="1978-10-27" collectionCode="SANC" collectorName="Bornemissza." country="South Africa" location="Dune Forest" pageId="4" pageNumber="71" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC88D59E3467C27E6B" bold="true" box="[252,467,575,600]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC88D59E3467C27E6B" box="[252,467,575,600]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN St Lucia Est. Nat. Reserve, 
<location id="8E96606CE553FFCC8B759E4B65F67E64" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8B759E4B65F67E64" box="[860,999,576,599]" country="South Africa" name="Dune Forest" pageId="4" pageNumber="71">Dune Forest</location>
28[◦] 13[’] S; 32[◦] 21[’] E, 
<date id="FFF71077E553FFCC8D0C9E4B63867E6B" box="[1317,1431,576,600]" pageId="4" pageNumber="71" value="1978-10-27">
<collectingDate id="EFB3E99FE553FFCC8D0C9E4B63867E6B" box="[1317,1431,576,600]" pageId="4" pageNumber="71" value="1978-10-27">27.x.1978</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC88E09E6D67707E4C" box="[201,353,614,639]" pageId="4" pageNumber="71">Bornemissza.</collectorName>
<emphasis id="B93DEAA5E553FFCC89419E6D67607E4C" bold="true" box="[360,369,614,639]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC895B9E6C67857E4C" box="[370,404,615,639]" pageId="4" pageNumber="71" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89B59E6D67E37E4C" bold="true" box="[412,498,614,639]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC89B59E6D67FB7E4C" box="[412,490,614,639]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8A2B9E6D649B7E95" ID-GBIF-Occurrence="1290984942" collectingDate="1990-02-08" collectingDateMax="1990-02-11" collectingDateMin="1990-02-08" collectionCode="SANC" collectorName="B. Grobbelaar." country="South Africa" location="Ntl Kosi Bay" municipality="Ntl Kosi Bay Nature Res" pageId="4" pageNumber="71" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8A2B9E6D64C87E4C" bold="true" box="[514,729,614,639]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8A2B9E6D64C87E4C" box="[514,729,614,639]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN 
<collectingMunicipality id="6B92ACCDE553FFCC8B009E6D62547E4C" box="[809,1093,614,640]" pageId="4" pageNumber="71">
<location id="8E96606CE553FFCC8B009E6D65AE7EB3" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8B009E6D65AE7EB3" box="[809,959,614,640]" country="South Africa" municipality="Ntl Kosi Bay Nature Res" name="Ntl Kosi Bay" pageId="4" pageNumber="71">Ntl Kosi Bay</location>
Nature Res
</collectingMunicipality>
26[◦] 34[’] S; 32[◦] 28[’] E, 
<date id="FFF71077E553FFCC8DAD9E6C67187E95" pageId="4" pageNumber="71" value="1990-02-08" valueMax="1990-02-11" valueMin="1990-02-08">
<collectingDate id="EFB3E99FE553FFCC8DAD9E6C67187E95" pageId="4" pageNumber="71" value="1990-02-08" valueMax="1990-02-11" valueMin="1990-02-08">8- 11.ii.1990</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89649E8567E47E95" box="[333,501,653,678]" pageId="4" pageNumber="71">B. Grobbelaar.</collectorName>
<emphasis id="B93DEAA5E553FFCC89D79E8664167E95" bold="true" box="[510,519,653,678]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8A2E9E85643B7E95" box="[519,554,654,678]" pageId="4" pageNumber="71" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8A1D9E86649B7E95" bold="true" box="[564,650,653,678]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8A1D9E8664937E95" box="[564,642,653,678]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8AB59E8665487EFE" ID-GBIF-Occurrence="1290984944" collectingDate="1978-10-27" collectionCode="SANC" collectorName="Bornemissza." country="South Africa" location="Near Richards Bay" pageId="4" pageNumber="71" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8AB59E8665647E95" bold="true" box="[668,885,653,678]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8AB59E8665647E95" box="[668,885,653,678]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC8BF99E8562BD7E95" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8BF99E8562BD7E95" box="[976,1196,653,678]" country="South Africa" name="Near Richards Bay" pageId="4" pageNumber="71">Near Richards Bay</location>
, 28[◦] 36[’] 52[’’] S; 32[◦] 17[’] 34[’’] E, 
<date id="FFF71077E553FFCC89559EBE67FF7EFE" box="[380,494,693,717]" pageId="4" pageNumber="71" value="1978-10-27">
<collectingDate id="EFB3E99FE553FFCC89559EBE67FF7EFE" box="[380,494,693,717]" pageId="4" pageNumber="71" value="1978-10-27">27.x.1978</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8A079EBF64D77EFE" box="[558,710,692,717]" pageId="4" pageNumber="71">Bornemissza.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8AF19EBE64EA7EFE" box="[728,763,693,717]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B2A9EBF65487EFE" bold="true" box="[771,857,692,717]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B2A9EBF65407EFE" box="[771,849,692,717]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8B409EBF65DA7EC7" ID-GBIF-Occurrence="1290984935" collectingDate="1996-02" collectionCode="TMSA" collectorName="R Stals." country="South Africa" location="Thembe Elephant Park" pageId="4" pageNumber="71" specimenCount="11" specimenCount-female="6" specimenCount-male="5" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8B409EBF62517EFE" bold="true" box="[873,1088,692,717]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8B409EBF62517EFE" box="[873,1088,692,717]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN, 
<location id="8E96606CE553FFCC8CB19EBF638D7EFE" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8CB19EBF638D7EFE" box="[1176,1436,692,717]" country="South Africa" name="Thembe Elephant Park" pageId="4" pageNumber="71">Thembe Elephant Park</location>
27[◦] 01[’] S; 32[◦] 24[’] E, 01-49. 
<date id="FFF71077E553FFCC8A3F9ED064767EC7" box="[534,615,731,756]" pageId="4" pageNumber="71" value="1996-02">
<collectingDate id="EFB3E99FE553FFCC8A3F9ED064767EC7" box="[534,615,731,756]" pageId="4" pageNumber="71" value="1996-02">ii.1996</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8A8F9ED764EF7EC7" box="[678,766,731,756]" pageId="4" pageNumber="71">R Stals.</collectorName>
<emphasis id="B93DEAA5E553FFCC8B2F9ED0651E7EC7" bold="true" box="[774,783,731,756]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8B269ED065247EC7" box="[783,821,731,756]" pageId="4" pageNumber="71" type="male">5♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8B689ED065757EC7" box="[833,868,731,756]" pageId="4" pageNumber="71" type="female">6♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B459ED065DA7EC7" bold="true" box="[876,971,731,756]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B459ED065D37EC7" box="[876,962,731,756]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8BF19ED063387F28" ID-GBIF-Occurrence="1290984946" collectingDate="2002-11-14" collectionCode="TMSA" collectorName="Burguer &amp; Harrison &amp; Muller." country="South Africa" location="Ntl Kosi Bay" municipality="Ntl Kosi Bay Nature Res" pageId="4" pageNumber="71" specimenCount="15" specimenCount-female="8" specimenCount-male="7" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8BF19ED062BE7EC7" box="[984,1199,731,756]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
, KZN, 
<collectingMunicipality id="6B92ACCDE553FFCC8D2F9ED067077F29" pageId="4" pageNumber="71">
<location id="8E96606CE553FFCC8D2F9ED0638A7EC7" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8D2F9ED0638A7EC7" box="[1286,1435,731,756]" country="South Africa" municipality="Ntl Kosi Bay Nature Res" name="Ntl Kosi Bay" pageId="4" pageNumber="71">Ntl Kosi Bay</location>
Nature Res
</collectingMunicipality>
26[◦] 34[’] S; 32[◦] 28[’] E, 
<date id="FFF71077E553FFCC8A749F0964C67F28" box="[605,727,770,795]" pageId="4" pageNumber="71" value="2002-11-14">
<collectingDate id="EFB3E99FE553FFCC8A749F0964C67F28" box="[605,727,770,795]" pageId="4" pageNumber="71" value="2002-11-14">14.xi.2002</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8B319F0865647F28" box="[792,885,771,795]" pageId="4" pageNumber="71">Burguer</collectorName>
, 
<collectorName id="26BC5361E553FFCC8BAB9F0965F57F28" box="[898,996,770,795]" pageId="4" pageNumber="71">Harrison</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8C259F09624F7F28" box="[1036,1118,770,795]" pageId="4" pageNumber="71">Muller.</collectorName>
<emphasis id="B93DEAA5E553FFCC8C4F9F09627E7F28" bold="true" box="[1126,1135,770,795]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8C599F0862827F29" box="[1136,1171,771,794]" pageId="4" pageNumber="71" type="male">7♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8C8A9F0962D47F29" box="[1187,1221,770,794]" pageId="4" pageNumber="71" type="female">8♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8CE69F0963387F28" bold="true" box="[1231,1321,770,795]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8CE69F0963337F28" box="[1231,1314,770,795]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8D139F0966E07F5B" ID-GBIF-Occurrence="1290984930" collectingDate="1992-11-23" collectionCode="TMSA" collectorName="Endrody-Younga." country="South Africa" location="Sodwana Bay" municipality="Zululand" pageId="4" pageNumber="71" specimenCount="16" specimenCount-female="8" specimenCount-male="8" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8D139F0967157F72" bold="true" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8D139F0967157F72" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, 
<collectingMunicipality id="6B92ACCDE553FFCC893D9F23676D7F72" box="[276,380,808,833]" pageId="4" pageNumber="71">Zululand</collectingMunicipality>
, 
<location id="8E96606CE553FFCC89A09F2364377F72" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC89A09F2364377F72" box="[393,550,808,833]" country="South Africa" municipality="Zululand" name="Sodwana Bay" pageId="4" pageNumber="71">Sodwana Bay</location>
<quantity id="4CB19B52E553FFCC8A199F2364757F72" box="[560,612,808,833]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="71" unit="km" value="5.0">5km</quantity>
, 27[◦] 21[’] S; 32[◦] 23[’] E, 
<date id="FFF71077E553FFCC8B989F23623A7F72" box="[945,1067,808,833]" pageId="4" pageNumber="71" value="1992-11-23">
<collectingDate id="EFB3E99FE553FFCC8B989F23623A7F72" box="[945,1067,808,833]" pageId="4" pageNumber="71" value="1992-11-23">23.xi.1992</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8C459F2363257F72" box="[1132,1332,808,833]" pageId="4" pageNumber="71">Endrody-Younga.</collectorName>
<emphasis id="B93DEAA5E553FFCC8D159F2363547F72" bold="true" box="[1340,1349,808,833]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8D6C9F2263797F72" box="[1349,1384,809,833]" pageId="4" pageNumber="71" type="male">8♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8D509F22638A7F72" box="[1401,1435,809,833]" pageId="4" pageNumber="71" type="female">8♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC88BE9F4466E07F5B" bold="true" box="[151,241,847,872]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC88BE9F4466FB7F5B" box="[151,234,847,872]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC892D9F4464537FBC" ID-GBIF-Occurrence="1290984951" collectingDate="1992-12-01" collectionCode="TMSA" collectorName="N Zululand &amp; Endrody-Younga." country="South Africa" location="Ndumu Game Reserve" pageId="4" pageNumber="71" specimenCount="72" specimenCount-female="42" specimenCount-male="30" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC892D9F4467F17F5B" bold="true" box="[260,480,847,872]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC892D9F4467F17F5B" box="[260,480,847,872]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, 
<collectorName id="26BC5361E553FFCC89DB9F5B646B7F5B" box="[498,634,847,872]" pageId="4" pageNumber="71">N Zululand</collectorName>
, 
<location id="8E96606CE553FFCC8AA39F4465827F54" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8AA39F4465827F54" box="[650,915,847,872]" country="South Africa" name="Ndumu Game Reserve" pageId="4" pageNumber="71">Ndumu Game Reserve</location>
26[◦] 32[’] S; 32[◦] 10[’] E, 
<date id="FFF71077E553FFCC8CC49F4463707F5B" box="[1261,1377,847,872]" pageId="4" pageNumber="71" value="1992-12-01">
<collectingDate id="EFB3E99FE553FFCC8CC49F4463707F5B" box="[1261,1377,847,872]" pageId="4" pageNumber="71" value="1992-12-01">1.xii.1992</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC88BE9F7D674F7FBC" box="[151,350,886,911]" pageId="4" pageNumber="71">Endrody-Younga.</collectorName>
<emphasis id="B93DEAA5E553FFCC894C9F7D677F7FBC" bold="true" box="[357,366,886,911]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC89469F7C67B27FBC" box="[367,419,887,911]" pageId="4" pageNumber="71" type="male">30♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC89869F7C67F17FBC" box="[431,480,887,911]" pageId="4" pageNumber="71" type="female">42♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89C19F7D64537FBC" bold="true" box="[488,578,886,911]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC89C19F7D642A7FBC" box="[488,571,886,911]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8A7B9F7D64F37F85" ID-GBIF-Occurrence="1290984940" collectingDate="1992-12-12" collectionCode="TMSA" collectorName="N Zululand &amp; Endrodi-Younga." country="South Africa" location="Lake Bangazi" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8A7B9F7D65387FBC" bold="true" box="[594,809,886,911]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8A7B9F7D65387FBC" box="[594,809,886,911]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, 
<collectorName id="26BC5361E553FFCC8B119F7C65AD7FBC" box="[824,956,886,911]" pageId="4" pageNumber="71">N Zululand</collectorName>
, 
<location id="8E96606CE553FFCC8BE19F7D62757FBC" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8BE19F7D62757FBC" box="[968,1124,886,911]" country="South Africa" name="Lake Bangazi" pageId="4" pageNumber="71">Lake Bangazi</location>
28[◦] 04[’] S; 32[◦] 18[’] E, 
<date id="FFF71077E553FFCC88BE9F9667087F85" box="[151,281,925,950]" pageId="4" pageNumber="71" value="1992-12-12">
<collectingDate id="EFB3E99FE553FFCC88BE9F9667087F85" box="[151,281,925,950]" pageId="4" pageNumber="71" value="1992-12-12">12.xii.1992</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89719F9664087F85" box="[344,537,925,950]" pageId="4" pageNumber="71">Endrodi-Younga.</collectorName>
<emphasis id="B93DEAA5E553FFCC8A089F96643B7F85" bold="true" box="[545,554,925,950]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8A039F95645F7F85" box="[554,590,926,950]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8A759F95646E7F85" box="[604,639,926,950]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8AAE9F9664F37F85" bold="true" box="[647,738,925,950]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8AAE9F9664CB7F85" box="[647,730,925,950]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<emphasis id="B93DEAA5E553FFCC8ADB9F9665D87F85" bold="true" box="[754,969,925,950]" pageId="4" pageNumber="71">SOUTH AFRICA</emphasis>
, Zululand, 
<materialsCitation id="3B213CEAE553FFCC8C649F9565C67FEE" ID-GBIF-Occurrence="1290984926" collectingDate="1992-11-23" collectionCode="TMSA" collectorName="Endrody-Younga." country="Saint Lucia" location="St. Lucia" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F35E7627E553FFCC8C649F9562A77F85" box="[1101,1206,925,950]" name="Saint Lucia" pageId="4" pageNumber="71">St. Lúcia</collectingCountry>
. Mission Rock 28[◦] 13[’] S; 32[◦] 21[’] E, 
<date id="FFF71077E553FFCC89B89FCF641B7FEE" box="[401,522,964,989]" pageId="4" pageNumber="71" value="1992-11-23">
<collectingDate id="EFB3E99FE553FFCC89B89FCF641B7FEE" box="[401,522,964,989]" pageId="4" pageNumber="71" value="1992-11-23">23.xi.1992</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC8A609FCF65017FEE" box="[585,784,964,989]" pageId="4" pageNumber="71">Endrody-Younga.</collectorName>
<emphasis id="B93DEAA5E553FFCC8B3E9FCF65317FEE" bold="true" box="[791,800,964,989]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8B099FCF65547FEF" box="[800,837,964,988]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8B7B9FCF65647FEF" box="[850,885,964,988]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B549FCF65C67FEE" bold="true" box="[893,983,964,989]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B549FCF65C17FEE" box="[893,976,964,989]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8BCF9FCF629E7830" ID-GBIF-Occurrence="1290984933" collectingDate="1990-01-23" collectionCode="TMSA" collectorName="J. Klimaszewski." country="South Africa" location="Forest" municipality="Natal" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Cape" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8BCF9FCF62AC7FEE" bold="true" box="[998,1213,964,989]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8BCF9FCF62AC7FEE" box="[998,1213,964,989]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, 
<collectingMunicipality id="6B92ACCDE553FFCC8CE59FCF631A7FEE" box="[1228,1291,964,989]" pageId="4" pageNumber="71">Natal</collectingMunicipality>
, 
<collectingRegion id="498DF855E553FFCC8D3E9FCF63407FEF" box="[1303,1361,964,988]" country="South Africa" name="Western Cape" pageId="4" pageNumber="71">Cape</collectingRegion>
Vidal, 
<location id="8E96606CE553FFCC88BE9FE066CC7831" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC88BE9FE066CC7831" box="[151,221,1003,1026]" country="South Africa" municipality="Natal" name="Forest" pageId="4" pageNumber="71" stateProvince="Cape">Forest</location>
litter 28[◦] 8[’] S; 32[◦] 33[’] E, 
<collectingDate id="EFB3E99FE553FFCC8A7D9FE164D07830" box="[596,705,1002,1027]" pageId="4" pageNumber="71" value="1990-01-23">
<date id="FFF71077E553FFCC8A7D9FE164D07830" box="[596,705,1002,1027]" pageId="4" pageNumber="71" value="1990-01-23">23.i.1990</date>
</collectingDate>
, leg: 
<collectorName id="26BC5361E553FFCC8B2B9FE065D27830" box="[770,963,1002,1027]" pageId="4" pageNumber="71">J. Klimaszewski.</collectorName>
<emphasis id="B93DEAA5E553FFCC8BE29FE165C47830" bold="true" box="[971,981,1002,1027]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC8BFC9FE065E87830" box="[981,1017,1003,1027]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8C219FE0623A7830" box="[1032,1067,1003,1027]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8C1D9FE1629E7830" bold="true" box="[1076,1167,1002,1027]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8C1D9FE162967830" box="[1076,1159,1002,1027]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8CB69FE166FD7862" ID-GBIF-Occurrence="1290984932" collectingDate="1992-11-20" collectionCode="SANC" collectorName="N Zululand &amp; Endrody-Younga." country="South Africa" location="SOUTH AFRICA" pageId="4" pageNumber="71" specimenCount="42" specimenCount-female="30" specimenCount-male="12" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8CB69FE163697830" bold="true" box="[1183,1400,1002,1027]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8CB69FE163697830" box="[1183,1400,1002,1027]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, 
<collectorName id="26BC5361E553FFCC8DA19FE066EE7819" pageId="4" pageNumber="71">N Zululand</collectorName>
, Hluhluwe Game Reserve. 28[◦] 03[’] S; 32[◦] 02[’] E, 
<date id="FFF71077E553FFCC8BA4981A62197819" box="[909,1032,1041,1066]" pageId="4" pageNumber="71" value="1992-11-20">
<collectingDate id="EFB3E99FE553FFCC8BA4981A62197819" box="[909,1032,1041,1066]" pageId="4" pageNumber="71" value="1992-11-20">20.xi.1992</collectingDate>
</date>
. leg: 
<collectorName id="26BC5361E553FFCC8C65981A63057819" box="[1100,1300,1041,1066]" pageId="4" pageNumber="71">Endrody-Younga.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8D0E9819634A7819" box="[1319,1371,1042,1066]" pageId="4" pageNumber="71" type="male">12♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8D429819638A7819" box="[1387,1435,1042,1066]" pageId="4" pageNumber="71" type="female">30♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC88BE983366FD7862" bold="true" box="[151,236,1080,1105]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC88BE983366F47862" box="[151,229,1080,1105]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
; 
<materialsCitation id="3B213CEAE553FFCC88D29833640B784B" ID-GBIF-Occurrence="1290984927" collectingDate="1979-01-23" collectingDateMax="1979-01-24" collectingDateMin="1979-01-23" collectionCode="TMSA" collectorName="NH Aschenborn." country="South Africa" location="Forest" pageId="4" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC88D2983367C17862" bold="true" box="[251,464,1080,1105]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC88D2983367C17862" box="[251,464,1080,1105]" name="South Africa" pageId="4" pageNumber="71">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN St Lucia Est. Nat. Reserve, 
<location id="8E96606CE553FFCC8B7C9832658A7863" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8B7C9832658A7863" box="[853,923,1081,1104]" country="South Africa" name="Forest" pageId="4" pageNumber="71">Forest</location>
28[◦] 13[’] S; 32[◦] 21[’] E, 
<date id="FFF71077E553FFCC8CFC983363777862" box="[1237,1382,1080,1105]" pageId="4" pageNumber="71" value="1979-01-24" valueMax="1979-01-23" valueMin="1979-01-24">
<collectingDate id="EFB3E99FE553FFCC8CFC983363777862" box="[1237,1382,1080,1105]" pageId="4" pageNumber="71" value="1979-01-24" valueMax="1979-01-23" valueMin="1979-01-24">24-23.i.1979</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC88BE986B6744784B" box="[151,341,1119,1144]" pageId="4" pageNumber="71">NH Aschenborn.</collectorName>
<emphasis id="B93DEAA5E553FFCC897598546774784B" bold="true" box="[348,357,1119,1144]" pageId="4" pageNumber="71">(</emphasis>
<specimenCount id="9D4FFD3EE553FFCC894C985467987844" box="[357,393,1119,1143]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC89BF985467A87844" box="[406,441,1119,1143]" pageId="4" pageNumber="71" type="female">2♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89969854640B784B" bold="true" box="[447,538,1119,1144]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC899698546403784B" box="[447,530,1119,1144]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="71">TMSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8A009854640B78AD" ID-GBIF-Occurrence="1290984936" collectingDate="1974-05-04" collectionCode="UPSA" collectorName="A. Strydom." country="Mozambique" location="Pomene" pageId="4" pageNumber="71" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Inhambane" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8A00985464E4784B" bold="true" box="[553,757,1119,1144]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8A00985464E4784B" box="[553,757,1119,1144]" name="Mozambique" pageId="4" pageNumber="71">MOZAMBIQUE</collectingCountry>
</emphasis>
, 
<collectingRegion id="498DF855E553FFCC8B2A98546592784B" box="[771,899,1119,1144]" country="Mozambique" name="Inhambane" pageId="4" pageNumber="71">Inhambane</collectingRegion>
, 
<location id="8E96606CE553FFCC8BA4986B65FB7844" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8BA4986B65FB7844" box="[909,1002,1120,1143]" country="Mozambique" name="Pomene" pageId="4" pageNumber="71" stateProvince="Inhambane">Pomene</location>
, 22[◦] 35[’] S; 35[◦] 21[’] E, 
<date id="FFF71077E553FFCC8D0E985463867844" box="[1319,1431,1119,1143]" pageId="4" pageNumber="71" value="1974-05-04">
<collectingDate id="EFB3E99FE553FFCC8D0E985463867844" box="[1319,1431,1119,1143]" pageId="4" pageNumber="71" value="1974-05-04">04.v.1974</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC88E3988D674778AD" box="[202,342,1157,1182]" pageId="4" pageNumber="71">A. Strydom.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8941988E679B78AD" box="[360,394,1157,1182]" pageId="4" pageNumber="71" type="male">5♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC89B2988E67AF78AD" box="[411,446,1157,1182]" pageId="4" pageNumber="71" type="female">5♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC89EE988E640B78AD" bold="true" box="[455,538,1157,1182]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC89EE988E640378AD" box="[455,530,1157,1182]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="4" pageNumber="71">UPSA</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8A02988E656478F6" ID-GBIF-Occurrence="1290984928" collectingDate="2007-11-10" collectingDateMax="2007-11-16" collectingDateMin="2007-11-10" collectionCode="SANC" collectorName="W. Strumpher &amp; C. Deschodt." country="Mozambique" location="Maputo" pageId="4" pageNumber="71" specimenCount="17" specimenCount-female="10" specimenCount-male="7" stateProvince="Maputo" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8A02988E64E678AD" bold="true" box="[555,759,1157,1182]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8A02988E64E678AD" box="[555,759,1157,1182]" name="Mozambique" pageId="4" pageNumber="71">MOZAMBIQUE</collectingCountry>
</emphasis>
, 
<collectingRegion id="498DF855E553FFCC8B2E988C654F78AD" box="[775,862,1159,1182]" country="Mozambique" name="Maputo" pageId="4" pageNumber="71">Maputo</collectingRegion>
Elephant Reserve 26[◦] 39[’] S; 32[◦] 43[’] E, 
<date id="FFF71077E553FFCC8D5F988D670078F6" pageId="4" pageNumber="71" value="2007-11-10" valueMax="2007-11-16" valueMin="2007-11-10">
<collectingDate id="EFB3E99FE553FFCC8D5F988D670078F6" pageId="4" pageNumber="71" value="2007-11-10" valueMax="2007-11-16" valueMin="2007-11-10">10- 16.xi.2007</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC896698A667FD78F6" box="[335,492,1196,1221]" pageId="4" pageNumber="71">W. Strümpher</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8A3898A664B578F6" box="[529,676,1196,1221]" pageId="4" pageNumber="71">C. Deschodt.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8A9D98A664C678F6" box="[692,727,1197,1221]" pageId="4" pageNumber="71" type="male">7♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8ACF98A6650678F6" box="[742,791,1197,1221]" pageId="4" pageNumber="71" type="female">10♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B3698A7656478F6" bold="true" box="[799,885,1196,1221]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B3698A7657C78F6" box="[799,877,1196,1221]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8BAD98A7659878DF" ID-GBIF-Occurrence="1290984931" collectingDate="1972-01-19" collectionCode="SANC" collectorName="Bornemissza &amp; Kirk." country="Mozambique" location="Dondo" pageId="4" pageNumber="71" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sofala" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8BAD98A7624178F6" bold="true" box="[900,1104,1196,1221]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8BAD98A7624178F6" box="[900,1104,1196,1221]" name="Mozambique" pageId="4" pageNumber="71">MOZAMBIQUE</collectingCountry>
</emphasis>
, 
<collectingRegion id="498DF855E553FFCC8C7798A762B878F6" box="[1118,1193,1196,1221]" country="Mozambique" name="Sofala" pageId="4" pageNumber="71">Sofala</collectingRegion>
, 
<location id="8E96606CE553FFCC8C9C98A7631078F6" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8C9C98A7631078F6" box="[1205,1281,1196,1221]" country="Mozambique" name="Dondo" pageId="4" pageNumber="71" stateProvince="Sofala">Dondo</location>
19[◦] 37[’] S; 34[◦] 45[’] E. 
<date id="FFF71077E553FFCC891998D8678C78DF" box="[304,413,1235,1260]" pageId="4" pageNumber="71" value="1972-01-19">
<collectingDate id="EFB3E99FE553FFCC891998D8678C78DF" box="[304,413,1235,1260]" pageId="4" pageNumber="71" value="1972-01-19">19.i.1972</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89F098D8647B78DF" box="[473,618,1235,1260]" pageId="4" pageNumber="71">Bornemissza</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8AA498D864D978DF" box="[653,712,1235,1260]" pageId="4" pageNumber="71">Kirk.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8AF198DF64EA78DF" box="[728,763,1236,1260]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
, 
<specimenCount id="9D4FFD3EE553FFCC8B2098DF653D78DF" box="[777,812,1236,1260]" pageId="4" pageNumber="71" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E553FFCC8B1A98D8659878DF" bold="true" box="[819,905,1235,1260]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8B1A98D8659078DF" box="[819,897,1235,1260]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B213CEAE553FFCC8BBE98D8655C7920" ID-GBIF-Occurrence="1290984954" collectingDate="1972-01-19" collectionCode="SANC" collectorName="Bornemissza &amp; Kirk." country="Mozambique" location="Beira" pageId="4" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Sofala" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8BBE98D8627278DF" bold="true" box="[919,1123,1235,1260]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8BBE98D8627278DF" box="[919,1123,1235,1260]" name="Mozambique" pageId="4" pageNumber="71">MOZAMBIQUE</collectingCountry>
</emphasis>
, 
<collectingRegion id="498DF855E553FFCC8C5898D862AA78DF" box="[1137,1211,1235,1260]" country="Mozambique" name="Sofala" pageId="4" pageNumber="71">Sofala</collectingRegion>
, 
<location id="8E96606CE553FFCC8CEE98D8631578DF" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8CEE98D8631578DF" box="[1223,1284,1235,1260]" country="Mozambique" name="Beira" pageId="4" pageNumber="71" stateProvince="Sofala">Beira</location>
19[◦] 50[’] S; 34[◦] 51[’] E 
<date id="FFF71077E553FFCC890198F167857920" box="[296,404,1274,1299]" pageId="4" pageNumber="71" value="1972-01-19">
<collectingDate id="EFB3E99FE553FFCC890198F167857920" box="[296,404,1274,1299]" pageId="4" pageNumber="71" value="1972-01-19">19.i.1972</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89F998F164707920" box="[464,609,1274,1299]" pageId="4" pageNumber="71">Bornemissza</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8AAD98F164AF7920" box="[644,702,1274,1299]" pageId="4" pageNumber="71">Kirk.</collectorName>
(
<specimenCount id="9D4FFD3EE553FFCC8AE798F064E07920" box="[718,753,1275,1299]" pageId="4" pageNumber="71" type="male">1♂</specimenCount>
<emphasis id="B93DEAA5E553FFCC8ADE98F1655C7920" bold="true" box="[759,845,1274,1299]" pageId="4" pageNumber="71">
<collectionCode id="ED58AE72E553FFCC8ADE98F165547920" box="[759,837,1274,1299]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="71">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
; 
<materialsCitation id="3B213CEAE553FFCC8B7298F164FC7909" ID-GBIF-Occurrence="1290984952" collectingDate="1972-01-11" collectingDateMax="1972-01-28" collectingDateMin="1972-01-11" collectorName="Bornemissza &amp; Kirk." country="Mozambique" location="Gorongosa" pageId="4" pageNumber="71" specimenCount="1" stateProvince="Sofala" typeStatus="holotype">
<emphasis id="B93DEAA5E553FFCC8B7298F162367920" bold="true" box="[859,1063,1274,1299]" pageId="4" pageNumber="71">
<collectingCountry id="F35E7627E553FFCC8B7298F162367920" box="[859,1063,1274,1299]" name="Mozambique" pageId="4" pageNumber="71">MOZAMBIQUE</collectingCountry>
</emphasis>
, 
<collectingRegion id="498DF855E553FFCC8C1D98F1626E7920" box="[1076,1151,1274,1299]" country="Mozambique" name="Sofala" pageId="4" pageNumber="71">Sofala</collectingRegion>
, 
<location id="8E96606CE553FFCC8CA398F063147920" LSID="urn:lsid:plazi:treatment:03E087A1E554FFCC88BE9C9C62A87A69:8E96606CE553FFCC8CA398F063147920" box="[1162,1285,1275,1299]" country="Mozambique" name="Gorongosa" pageId="4" pageNumber="71" stateProvince="Sofala">Gorongosa</location>
18[◦] 57[’] S; 34[◦] 10[’] E, 
<date id="FFF71077E553FFCC8919992A67D17909" box="[304,448,1313,1338]" pageId="4" pageNumber="71" value="1972-01-11" valueMax="1972-01-28" valueMin="1972-01-11">
<collectingDate id="EFB3E99FE553FFCC8919992A67D17909" box="[304,448,1313,1338]" pageId="4" pageNumber="71" value="1972-01-11" valueMax="1972-01-28" valueMin="1972-01-11">11-28.i.1972</collectingDate>
</date>
, leg: 
<collectorName id="26BC5361E553FFCC89D4992A649F7909" box="[509,654,1313,1338]" pageId="4" pageNumber="71">Bornemissza</collectorName>
&amp; 
<collectorName id="26BC5361E553FFCC8A98992A64FC7909" box="[689,749,1313,1338]" pageId="4" pageNumber="71">Kirk.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BF636B7E553FFCC88EE994E66F179DD" blockId="4.[151,1436,151,1626]" pageId="4" pageNumber="71">
<emphasis id="B93DEAA5E553FFCC88EE994E673B796D" bold="true" box="[199,298,1349,1374]" pageId="4" pageNumber="71">Habitat.</emphasis>
This new species is confined to dense coastal woodlands and forest from northeast 
<collectingCountry id="F35E7627E553FFCC8CFD994E6375796D" box="[1236,1380,1349,1374]" name="South Africa" pageId="4" pageNumber="71">South Africa</collectingCountry>
to at least Central 
<collectingCountry id="F35E7627E553FFCC8915996267C179B1" box="[316,464,1385,1410]" name="Mozambique" pageId="4" pageNumber="71">Mozambique</collectingCountry>
(
<collectingRegion id="498DF855E553FFCC89C19962647879B1" box="[488,617,1385,1410]" country="Mozambique" name="Inhambane" pageId="4" pageNumber="71">Inhambane</collectingRegion>
, Pomene, Gorongosa, Dondo, Beira) (
<figureCitation id="13722A32E553FFCC8C689962629C79B1" box="[1089,1165,1385,1410]" captionStart="FIGURE 4" captionStartId="6.[151,250,1720,1742]" captionTargetBox="[408,1246,841,1677]" captionTargetId="figure@6.[340,1247,841,1698]" captionTargetPageId="6" captionText="FIGURE 4. Map of southern Africa showing the geographic distribution of the new species: S. umbraphilus, S. oralensis and S. neobornemisszanus." httpUri="https://zenodo.org/record/266255/files/figure.png" pageId="4" pageNumber="71">
<emphasis id="B93DEAA5E553FFCC8C689962629C79B1" bold="true" box="[1089,1165,1385,1410]" pageId="4" pageNumber="71">Fig. 4</emphasis>
</figureCitation>
). Across a vegetation restoration gradient at Richards Bay, 
<collectingRegion id="498DF855E553FFCC8A1E998664FB7995" box="[567,746,1421,1446]" country="South Africa" name="KwaZulu-Natal" pageId="4" pageNumber="71">KwaZulu-Natal</collectingRegion>
, 
<collectingCountry id="F35E7627E553FFCC8ADC998665967995" box="[757,903,1421,1446]" name="South Africa" pageId="4" pageNumber="71">South Africa</collectingCountry>
, 
<taxonomicName id="4C494D34E553FFCC8BBB998562047996" box="[914,1045,1421,1445]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="oralensis">
<emphasis id="B93DEAA5E553FFCC8BBB998562047996" box="[914,1045,1421,1445]" italics="true" pageId="4" pageNumber="71">S. oralensis</emphasis>
</taxonomicName>
was absent from grassland cleared of forest, but present in younger and older regenerating woodland (19, 6) plus natural dune forest (14) (
<bibRefCitation id="EFD84B46E553FFCC8D3399BA66C579DD" author="Davis" pageId="4" pageNumber="71" refString="Davis, A. L. V., Van, A., Arde, R. J., Scholtz, C. H. &amp; Delport J. H. (2002) Increasing representation of localized dung beetles across a chronosequence of regenerating vegetation and natural dune forest in South Africa. Global Ecology &amp; Biogeography (2002), 11, 191 - 209. http: // dx. doi. org / 10.1046 / j. 1466 - 822 X. 2002.00283. x" type="journal article" year="2002">
Davis 
<emphasis id="B93DEAA5E553FFCC8D4D99B9638479FA" box="[1380,1429,1457,1481]" italics="true" pageId="4" pageNumber="71">et al</emphasis>
. 2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C353653CE553FFCC88EE99F262A87A69" pageId="4" pageNumber="71" type="discussion">
<paragraph id="8BF636B7E553FFCC88EE99F262A87A69" blockId="4.[151,1436,151,1626]" pageId="4" pageNumber="71">
<emphasis id="B93DEAA5E553FFCC88EE99F2672A7A21" bold="true" box="[199,315,1529,1554]" pageId="4" pageNumber="71">Remarks.</emphasis>
Both species (
<taxonomicName id="4C494D34E553FFCC89CF99F264837A22" box="[486,658,1529,1553]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="umbraphilus" status="sp. nov.">
<emphasis id="B93DEAA5E553FFCC89CF99F264837A22" box="[486,658,1529,1553]" italics="true" pageId="4" pageNumber="71">S. umbraphilus</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E553FFCC8AB399F264E77A21" bold="true" box="[666,758,1529,1554]" pageId="4" pageNumber="71">
<taxonomicNameLabel id="A20E57DEE553FFCC8AB399F264E77A21" box="[666,758,1529,1554]" pageId="4" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C494D34E553FFCC8B0699F265A57A22" box="[815,948,1529,1553]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E553FFCC8B0699F265A57A22" box="[815,948,1529,1553]" italics="true" pageId="4" pageNumber="71">S. oralensis</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E553FFCC8B9499F262097A21" bold="true" box="[957,1048,1529,1554]" pageId="4" pageNumber="71">
<taxonomicNameLabel id="A20E57DEE553FFCC8B9499F262097A21" box="[957,1048,1529,1554]" pageId="4" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) exhibit little sexual dimorphism, but may be separated by meta- and mesotibia curved; the mid-ventral narrowing of the last abdominal sternite in males; compared to the lack of narrowing, and meta- and mesotibia almost straight in females,
</paragraph>
</subSubSection>
</treatment>
</document>