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<document id="47CBA1E4DB9594B15723734A047928D0" ID-CLB-Dataset="25519" ID-DOI="10.11646/zootaxa.4671.2.2" ID-GBIF-Dataset="978ec4b3-8d0e-4f47-8084-610b34b96759" ID-ISSN="1175-5326" ID-Zenodo-Dep="3442344" ID-ZooBank="8DA2FA3F-3629-4D10-92B0-671637D91DD4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568800506143" checkinUser="plazi" docAuthor="Brown, John W." docDate="2019" docId="03E08794FFC9FFF5FF5B7EC8FB06FDDD" docLanguage="en" docName="zootaxa.4671.2.2.pdf" docOrigin="Zootaxa 4671 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Honca Brown 2019, new genus" docType="treatment" docVersion="10" lastPageNumber="204" masterDocId="FFD9FFECFFC1FFFCFFCC7C00FFE1FFC6" masterDocTitle="New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae)" masterLastPageNumber="222" masterPageNumber="195" pageNumber="203" updateTime="1698755155468" updateUser="ExternalLinkService">
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<mods:title id="ACC246161F289621CE2582E9CBE53E0E">New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae)</mods:title>
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<mods:namePart id="590696740062BA032B1A5A64F659F2B5">Brown, John W.</mods:namePart>
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<subSubSection id="C3536509FFC9FFF4FF5B7EC8FE2CFD27" box="[151,461,711,738]" pageId="8" pageNumber="203" type="nomenclature">
<paragraph id="8BF63682FFC9FFF4FF5B7EC8FE2CFD27" blockId="8.[151,461,711,738]" box="[151,461,711,738]" pageId="8" pageNumber="203">
<heading id="D0BE81EEFFC9FFF4FF5B7EC8FE2CFD27" bold="true" box="[151,461,711,738]" fontSize="11" level="1" pageId="8" pageNumber="203" reason="1">
<emphasis id="B93DEA90FFC9FFF4FF5B7EC8FE2CFD27" bold="true" box="[151,461,711,738]" pageId="8" pageNumber="203">
<taxonomicName id="4C494D01FFC9FFF4FF5B7EC8FEDFFD27" ID-CoL="98VP3" authority="Pogue" authorityName="Brown" authorityYear="2019" box="[151,318,711,738]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93DEA90FFC9FFF4FF5B7EC8FF09FD24" bold="true" box="[151,232,712,738]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
Pogue
</taxonomicName>
,
<taxonomicNameLabel id="A20E57EBFFC9FFF4FE877EC7FE2CFD27" box="[331,461,711,737]" pageId="8" pageNumber="203" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BF63682FFC9FFF4FF5B7F06FD14FCD8" blockId="8.[151,757,773,798]" box="[151,757,773,798]" pageId="8" pageNumber="203">
<typeStatus id="54F28820FFC9FFF4FF5B7F06FF28FCD8" box="[151,201,774,798]" pageId="8" pageNumber="203">Type</typeStatus>
species:
<taxonomicName id="4C494D01FFC9FFF4FEEA7F06FDBFFCD8" authority="Busck, 1907" authorityName="Busck" authorityYear="1907" box="[294,606,773,798]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B93DEA90FFC9FFF4FEEA7F06FE36FCD8" box="[294,471,774,798]" italics="true" pageId="8" pageNumber="203">Phalonia grandis</emphasis>
<bibRefCitation id="EFD84B73FFC9FFF4FE117F05FDBFFCD8" author="Busck, A." box="[477,606,773,798]" pageId="8" pageNumber="203" pagination="19 - 36" refId="ref19735" refString="Busck, A. (1907) A review of the tortricid subfamily Phaloniinae with descriptions of new American species. Journal of the New York Entomological Society, 15, 19 - 36." type="journal article" year="1907">Busck, 1907</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
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<paragraph id="8BF63682FFC9FFF4FF5B7F44FD00FC9A" blockId="8.[151,1078,835,890]" box="[151,737,835,860]" pageId="8" pageNumber="203">
<taxonomicName id="4C494D01FFC9FFF4FF5B7F44FDD0FC9D" ID-CoL="98VP3" authority="Pogue and Mickevich, 1990: 322" authorityName="Pogue and Mickevich" authorityPageNumber="322" authorityYear="1990" box="[151,561,835,860]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FF5B7F44FF3AFC9D" box="[151,219,836,859]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
Pogue and Mickevich, 1990: 322
</taxonomicName>
(nomen nudum).
</paragraph>
<paragraph id="8BF63682FFC9FFF4FF5B7F61FBD7FCBC" blockId="8.[151,1078,835,890]" box="[151,1078,865,890]" pageId="8" pageNumber="203">
<treatmentCitationGroup id="AB5911ACFFC9FFF4FF5B7F61FBD7FCBC" box="[151,1078,865,890]" pageId="8" pageNumber="203">
<taxonomicName id="4C494D01FFC9FFF4FF5B7F61FF1AFCBC" ID-CoL="93N3S" authorityName="Guenée" authorityYear="1845" box="[151,251,865,890]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="tribe" tribe="Cochylini">Cochylini</taxonomicName>
New Genus 4 [
<taxonomicName id="4C494D01FFC9FFF4FE567F62FE06FCBC" authorityName="Busck" authorityYear="1907" box="[410,487,866,890]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B93DEA90FFC9FFF4FE567F62FE06FCBC" box="[410,487,866,890]" italics="true" pageId="8" pageNumber="203">grandis</emphasis>
</taxonomicName>
group]:
<treatmentCitation id="0AE81093FFC9FFF4FDF37F62FD12FCBC" author="Brown, J. W." box="[575,755,865,890]" page="208" pageId="8" pageNumber="203" year="2005">
<bibRefCitation id="EFD84B73FFC9FFF4FDF37F62FD12FCBC" author="Brown, J. W." box="[575,755,865,890]" pageId="8" pageNumber="203" refId="ref19539" refString="Brown, J. W. (2005) World catalogue of insects. Vol. 5. Tortricidae (Lepidoptera). Apollo Books, Stenstrup, 741 pp." type="book" year="2005">Brown 2005: 208</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE81093FFC9FFF4FD317F61FBD3FCBC" author="Metzler, E. H. &amp; Brown, J. W." box="[765,1074,865,890]" page="278" pageId="8" pageNumber="203" year="2014">
<bibRefCitation id="EFD84B73FFC9FFF4FD317F61FBD3FCBC" author="Metzler, E. H. &amp; Brown, J. W." box="[765,1074,865,890]" pageId="8" pageNumber="203" pagination="274 - 282" refId="ref20205" refString="Metzler, E. H. &amp; Brown, J. W. (2014) An updated check list the Cochylina (Tortricidae, Tortricinae, Euliini) of North America north of Mexico including Greenland, with comments on classification and identification. Journal of the Lepidopterists' Society, 68, 274 - 282. https: // doi. org / 10.18473 / lepi. v 68 i 4. a 5" type="journal article" year="2014">Metzler and Brown 2014: 278</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BF63682FFC9FFF4FF5B7FA4FD95FB88" blockId="8.[151,1437,931,2038]" pageId="8" pageNumber="203">
<taxonomicName id="4C494D01FFC9FFF4FF5B7FA4FF03FC7B" ID-CoL="98VP3" authorityName="Pogue and Mickevich" authorityYear="1990" box="[151,226,932,957]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FF5B7FA4FF03FC7B" box="[151,226,932,957]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
is described to accommodate the single species
<taxonomicName id="4C494D01FFC9FFF4FCC87FA3FC29FC7B" authorityName="Busck" authorityYear="1907" box="[772,968,931,957]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B93DEA90FFC9FFF4FCC87FA3FC29FC7B" box="[772,968,931,957]" italics="true" pageId="8" pageNumber="203">Phalonia grandis</emphasis>
</taxonomicName>
, which is among the largest Cochylina in the Nearctic region. The forewing pattern (
<figureCitation id="13722A07FFC9FFF4FDB07FC7FD20FC24" box="[636,705,967,994]" captionStart="FIGURES 110" captionStartId="5.[151,264,1832,1857]" captionTargetBox="[194,1394,166,1812]" captionTargetId="figure@5.[192,1396,166,1812]" captionTargetPageId="5" captionText="FIGURES 110. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM)." figureDoi="http://doi.org/10.5281/zenodo.3442348" httpUri="https://zenodo.org/record/3442348/files/figure.png" pageId="8" pageNumber="203">Fig. 7</figureCitation>
) somewhat resembles that of
<taxonomicName id="4C494D01FFC9FFF4FBD87FC8FAC6FC27" authorityName="Kearfott" authorityYear="1908" box="[1044,1319,967,993]" class="Insecta" family="Tortricidae" genus="Henricus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="umbrabasanus">
<emphasis id="B93DEA90FFC9FFF4FBD87FC8FAC6FC27" box="[1044,1319,967,993]" italics="true" pageId="8" pageNumber="203">Henricus umbrabasanus</emphasis>
</taxonomicName>
, however,
<taxonomicName id="4C494D01FFC9FFF4FF5B7FECFEF0FBC3" authorityName="Busck" authorityYear="1907" box="[151,273,1003,1030]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B93DEA90FFC9FFF4FF5B7FECFF4AFBC3" box="[151,171,1004,1029]" italics="true" pageId="8" pageNumber="203">H</emphasis>
.
<emphasis id="B93DEA90FFC9FFF4FF777FEBFEF0FBC3" box="[187,273,1003,1029]" italics="true" pageId="8" pageNumber="203">grandis</emphasis>
</taxonomicName>
is much larger (forewing length
<quantity id="4CB19B67FFC9FFF4FD477FEBFCE5FBC0" box="[651,772,1003,1030]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="8" pageNumber="203" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
vs.
<quantity id="4CB19B67FFC9FFF4FCF97FEBFC42FBC0" box="[821,931,1003,1030]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="8" pageNumber="203" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
) and lacks a pale brown basal patch.
<taxonomicName id="4C494D01FFC9FFF4FA9E7FECFA7CFBC3" ID-CoL="98VP3" authorityName="Pogue and Mickevich" authorityYear="1990" box="[1362,1437,1004,1029]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FA9E7FECFA7CFBC3" box="[1362,1437,1004,1029]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="4C494D01FFC9FFF4FE8D780FFD92FBEF" ID-CoL="922FN" authority="Razowski, 1984" authorityName="Razowski" authorityYear="1984" box="[321,627,1039,1065]" class="Insecta" family="Tortricidae" genus="Aphalonia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FE8D780FFE57FBEF" box="[321,438,1039,1065]" italics="true" pageId="8" pageNumber="203">Aphalonia</emphasis>
<bibRefCitation id="EFD84B73FFC9FFF4FE70780FFD92FBEF" author="Razowski, J." box="[444,627,1039,1065]" pageId="8" pageNumber="203" pagination="275 - 280" refId="ref20800" refString="Razowski, J. (1984) Descriptions of the Neotropical Cochylidii (Lepidoptera, Tortricidae). Annales Zoologici, 38, 275 - 280." type="journal article" year="1984">Razowski, 1984</bibRefCitation>
</taxonomicName>
, a South American genus, in size and forewing maculation, however, the genitalia of the two have little in common.
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC9FFF4FF0B7856FC07FA38" pageId="8" pageNumber="203" type="diagnosis">
<paragraph id="8BF63682FFC9FFF4FF0B7856FC07FA38" blockId="8.[151,1437,931,2038]" pageId="8" pageNumber="203">
<emphasis id="B93DEA90FFC9FFF4FF0B7856FEDBFBB7" bold="true" box="[199,314,1110,1137]" pageId="8" pageNumber="203">Diagnosis</emphasis>
. The shapes of the valva and transtilla of
<taxonomicName id="4C494D01FFC9FFF4FCC67858FCB4FBB7" authorityName="Pogue and Mickevich" authorityYear="1990" box="[778,853,1112,1137]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FCC67858FCB4FBB7" box="[778,853,1112,1137]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
are similar to those of some species of
<taxonomicName id="4C494D01FFC9FFF4FAC77857FAB7FBB7" authorityName="Billberg" authorityYear="1820" box="[1291,1366,1111,1137]" class="Insecta" family="Dryophthoridae" genus="Aethes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FAC77857FAB7FBB7" box="[1291,1366,1111,1137]" italics="true" pageId="8" pageNumber="203">Aethes</emphasis>
</taxonomicName>
, however,
<taxonomicName id="4C494D01FFC9FFF4FF1A787CFEC0FB53" authorityName="Pogue and Mickevich" authorityYear="1990" box="[214,289,1148,1173]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FF1A787CFEC0FB53" box="[214,289,1148,1173]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
lacks the sickle-shaped socii that are characteristic of most species of
<taxonomicName id="4C494D01FFC9FFF4FB89787BFB71FB53" authorityName="Billberg" authorityYear="1820" box="[1093,1168,1147,1173]" class="Insecta" family="Dryophthoridae" genus="Aethes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FB89787BFB71FB53" box="[1093,1168,1147,1173]" italics="true" pageId="8" pageNumber="203">Aethes</emphasis>
</taxonomicName>
. The most conspicuous autapomorphy of
<taxonomicName id="4C494D01FFC9FFF4FEAD78A0FE4DFB7F" authorityName="Pogue and Mickevich" authorityYear="1990" box="[353,428,1184,1209]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FEAD78A0FE4DFB7F" box="[353,428,1184,1209]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
is the highly modified phallus-juxta complex in the male genitalia (
<figureCitation id="13722A07FFC9FFF4FB7F789FFAE8FB7F" box="[1203,1289,1183,1209]" captionStart="FIGURES 1926" captionStartId="17.[151,264,1861,1886]" captionTargetBox="[151,1436,174,1805]" captionTargetId="figure@17.[151,1436,174,1805]" captionTargetPageId="17" captionText="FIGURES 1926. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249." figureDoi="http://doi.org/10.5281/zenodo.3442352" httpUri="https://zenodo.org/record/3442352/files/figure.png" pageId="8" pageNumber="203">Fig. 23</figureCitation>
): the juxta is rather long, slender, subrectangular, and widely bifurcate distally, and is strongly attached to the large, curved phallus. The phallus has a long, flattened process of the right side that is as long as the phallus, and a short, stout, more rounded process on the left side that is about one-fourth the length of the phallus. The latter features are reminiscent of the paired process that subtend the phallus in some species of
<taxonomicName id="4C494D01FFC9FFF4FCA1792FFC59FA8F" authorityName="Billberg" authorityYear="1820" box="[877,952,1327,1353]" class="Insecta" family="Dryophthoridae" genus="Aethes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FCA1792FFC59FA8F" box="[877,952,1327,1353]" italics="true" pageId="8" pageNumber="203">Aethes</emphasis>
</taxonomicName>
. In the female genitalia the spiculate, clubshaped arms of the sterigma (
<figureCitation id="13722A07FFC9FFF4FE2B7953FDDBFAAB" box="[487,570,1363,1389]" captionStart="FIGURES 3338" captionStartId="20.[151,264,1743,1768]" captionTargetBox="[151,1436,166,1723]" captionTargetId="figure@20.[151,1436,166,1723]" captionTargetPageId="20" captionText="FIGURES 3338. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660." figureDoi="http://doi.org/10.5281/zenodo.3442356" httpUri="https://zenodo.org/record/3442356/files/figure.png" pageId="8" pageNumber="203">Fig. 37</figureCitation>
) are somewhat reminiscent of some species of
<taxonomicName id="4C494D01FFC9FFF4FB997954FB32FAAB" authorityName="Brown" authorityYear="2013" box="[1109,1235,1364,1389]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FB997954FB32FAAB" box="[1109,1235,1364,1389]" italics="true" pageId="8" pageNumber="203">Eupinivora</emphasis>
</taxonomicName>
.
<taxonomicName id="4C494D01FFC9FFF4FB2E7954FACCFAAB" authorityName="Pogue and Mickevich" authorityYear="1990" box="[1250,1325,1364,1389]" class="Insecta" family="Tortricidae" genus="Honca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FB2E7954FACCFAAB" box="[1250,1325,1364,1389]" italics="true" pageId="8" pageNumber="203">Honca</emphasis>
</taxonomicName>
is further characterized by the absence of ocelli (usually present in Cochylina), a 3-segmented maxillary palpus (shared with
<taxonomicName id="4C494D01FFC9FFF4FF5B799CFEE0FA73" authorityName="Hubner" authorityYear="1825" box="[151,257,1436,1461]" class="Insecta" family="Tortricidae" genus="Eugnosta" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FF5B799CFEE0FA73" box="[151,257,1436,1461]" italics="true" pageId="8" pageNumber="203">Eugnosta</emphasis>
</taxonomicName>
), a reduced haustellum (shared with
<taxonomicName id="4C494D01FFC9FFF4FD60799CFCF7FA73" authorityName="Hubner" authorityYear="1825" box="[684,790,1436,1461]" class="Insecta" family="Tortricidae" genus="Eugnosta" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FD60799CFCF7FA73" box="[684,790,1436,1461]" italics="true" pageId="8" pageNumber="203">Eugnosta</emphasis>
</taxonomicName>
), four acanthi in the female frenulum (occurring consistently nowhere else in Cochylina), fusion of the socii with the tegumen, and the absence of cornuti in the vesica (shared with
<taxonomicName id="4C494D01FFC9FFF4FEE579E3FE6AFA3B" authorityName="Treitschke" authorityYear="1829" box="[297,395,1507,1533]" class="Insecta" family="Tortricidae" genus="Cochylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FEE579E3FE6AFA3B" box="[297,395,1507,1533]" italics="true" pageId="8" pageNumber="203">Cochylis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C494D01FFC9FFF4FE5579E3FDE7FA3B" authorityName="Walsingham" authorityYear="1897" box="[409,518,1507,1533]" class="Insecta" family="Tortricidae" genus="Thyraylia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC9FFF4FE5579E3FDE7FA3B" box="[409,518,1507,1533]" italics="true" pageId="8" pageNumber="203">Thyraylia</emphasis>
</taxonomicName>
, and
<emphasis id="B93DEA90FFC9FFF4FD8879E3FD3AFA3B" box="[580,731,1507,1533]" italics="true" pageId="8" pageNumber="203">Cochylichroa</emphasis>
, and few other genera).
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC9FFF5FF0B7A06FD91FE31" lastPageId="9" lastPageNumber="204" pageId="8" pageNumber="203" type="description">
<paragraph id="8BF63682FFC9FFF5FF0B7A06FD91FE31" blockId="8.[151,1437,931,2038]" lastBlockId="9.[151,1437,152,539]" lastPageId="9" lastPageNumber="204" pageId="8" pageNumber="203">
<emphasis id="B93DEA90FFC9FFF4FF0B7A06FEB0F9E7" bold="true" box="[199,337,1542,1569]" pageId="8" pageNumber="203">Description</emphasis>
. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus absent; sensory setae of antenna about as long as flagellomere diameter; labial palpus porrect, combined length of all segments ca. 3 times horizontal diameter of compound eye, shorter in female, segment III nearly concealed by scaling of segment II; maxillary palpus 3-segmented. Thorax: Posterior crest absent. Forewing length 11.0
<quantity id="4CB19B67FFC9FFF4FAF07A72FA7CF948" box="[1340,1437,1650,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="203" unit="mm" value="12.5">12.5 mm</quantity>
in male, 11.513.0 mm in female; length 2.42.8 times maximum width; forewing with base of costa straight, gently curved in distal 0.33 to rounded apex, costal fold absent; termen straight, oblique; Sc less than 0.5 wing length; R
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originating beyond middle of discal cell; R
<subScript id="17CD34C7FFC9FFF4FDB47AF1FD61F939" attach="left" box="[632,640,1777,1791]" fontSize="6" pageId="8" pageNumber="203">2</subScript>
nearer to R
<subScript id="17CD34C7FFC9FFF4FCCF7AF1FCEAF939" attach="left" box="[771,779,1777,1791]" fontSize="6" pageId="8" pageNumber="203">3</subScript>
than to R
<subScript id="17CD34C7FFC9FFF4FCB57AF1FC60F939" attach="left" box="[889,897,1777,1791]" fontSize="6" pageId="8" pageNumber="203">1</subScript>
; R
<subScript id="17CD34C7FFC9FFF4FC6F7AF0FC4AF938" attach="left" box="[931,939,1776,1790]" fontSize="6" pageId="8" pageNumber="203">5</subScript>
terminating at costa; M
<subScript id="17CD34C7FFC9FFF4FB7A7AF1FB5FF939" attach="left" box="[1206,1214,1777,1791]" fontSize="6" pageId="8" pageNumber="203">3</subScript>
and CuA
<subScript id="17CD34C7FFC9FFF4FAE67AF1FAD3F939" attach="left" box="[1322,1330,1777,1791]" fontSize="6" pageId="8" pageNumber="203">1</subScript>
separate; CuA
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originating ca. 0.8 length of discal cell; CuP absent; A1+2 stalked about 0.5 of their lengths. Hindwing with costa mostly straight, gently arched in apical 0.33, male without costal roll, termen slightly concave beneath apex; Sc+R
<subScript id="17CD34C7FFC9FFF4FF1A7B5DFF3FF8AD" attach="left" box="[214,222,1885,1899]" fontSize="6" pageId="8" pageNumber="203">1</subScript>
ca. 0.8 wing length; Rs and M
<subScript id="17CD34C7FFC9FFF4FDF37B5DFDA6F8AD" attach="left" box="[575,583,1885,1899]" fontSize="6" pageId="8" pageNumber="203">1</subScript>
stalked to ca. 0.6 length; M
<subScript id="17CD34C7FFC9FFF4FC447B5DFC71F8AD" attach="left" box="[904,912,1885,1899]" fontSize="6" pageId="8" pageNumber="203">3</subScript>
and CuA
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separate; CuA
<subScript id="17CD34C7FFC9FFF4FB637B5DFB56F8AD" attach="left" box="[1199,1207,1885,1899]" fontSize="6" pageId="8" pageNumber="203">2</subScript>
originating ca. 0.66 length of discal cell; frenulum with one acanthus in male, four in female. Abdomen: Unmodified. Male genitalia (
<figureCitation id="13722A07FFC9FFF4FF6C7B93FF13F86B" box="[160,242,1939,1965]" captionStart="FIGURES 1926" captionStartId="17.[151,264,1861,1886]" captionTargetBox="[151,1436,174,1805]" captionTargetId="figure@17.[151,1436,174,1805]" captionTargetPageId="17" captionText="FIGURES 1926. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249." figureDoi="http://doi.org/10.5281/zenodo.3442352" httpUri="https://zenodo.org/record/3442352/files/figure.png" pageId="8" pageNumber="203">Fig. 23</figureCitation>
) with uncus absent; socius prominent, fused to tegumen; transtilla with large, triangular, median process, with strongly sclerotized, pointed tip; valva short, broad, with upturned, well-sclerotized costa and attenuate apex, sacculus a straight sclerotized margin along venter of valva with slightly produced distal tip; vinculum arms con- nected distally by membrane; juxta a complicated plate, narrow, subrectangular, deeply emarginate on distal 0.35 distally resulting in pair of pointed processes, basal end fused to phallus. Phallus large, ca. 0.8 times length of valva, subtended by long, flattened lateral process on right side and shorter, peglike process on left (possibly homologous with “anellus lobes” of
<bibRefCitation id="EFD84B73FFC8FFF5FE617D05FD7FFED9" author="Sabourin, M. &amp; Miller, W. E. &amp; Metzler, E. H. &amp; Vargo, J. T." box="[429,670,261,287]" pageId="9" pageNumber="204" pagination="216 - 233" refId="ref21372" refString="Sabourin, M., Miller, W. E., Metzler, E. H. &amp; Vargo, J. T. (2002) Revised identities and new species of Aethes from midwestern North America. Journal of the Lepidopterists' Society, 56, 216 - 233." type="journal article" year="2002">
Sabourin
<emphasis id="B93DEA90FFC8FFF5FDD07D06FDB1FED9" box="[540,592,261,287]" italics="true" pageId="9" pageNumber="204">et al</emphasis>
. 2002
</bibRefCitation>
), coecum large; vesica without cornuti. Female genitalia (
<figureCitation id="13722A07FFC8FFF5FAF37D05FA75FED9" box="[1343,1428,261,287]" captionStart="FIGURES 3338" captionStartId="20.[151,264,1743,1768]" captionTargetBox="[151,1436,166,1723]" captionTargetId="figure@20.[151,1436,166,1723]" captionTargetPageId="20" captionText="FIGURES 3338. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660." figureDoi="http://doi.org/10.5281/zenodo.3442356" httpUri="https://zenodo.org/record/3442356/files/figure.png" pageId="9" pageNumber="204">Fig. 37</figureCitation>
) with papillae anales weakly curved, sclerotized; apophyses short, stout, apophyses anteriores ca. 1.1 times length of apophyses posteriores; sterigma membranous, median area with a pair of mesially-directed, spiculate, slightly club-shaped processes originating from posterior base of apophyses anteriores, each with 1520 long setae; ostium bursae wide; colliculum absent; ductus bursae short, undifferentiated from corpus bursae; corpus bursae with vertical lines of sclerotization below ostium bursae and with large patch of minute spicules; frail accessory bursae originating anterad of middle of ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC8FFF5FF0B7E01FB06FDDD" box="[199,1255,513,539]" pageId="9" pageNumber="204" type="etymology">
<paragraph id="8BF63682FFC8FFF5FF0B7E01FB06FDDD" blockId="9.[151,1437,152,539]" box="[199,1255,513,539]" pageId="9" pageNumber="204">
<emphasis id="B93DEA90FFC8FFF5FF0B7E01FEA9FDDD" bold="true" box="[199,328,513,539]" pageId="9" pageNumber="204">Etymology</emphasis>
. The generic name is an arbitrary combination of letters and is feminine in gender.
</paragraph>
</subSubSection>
</treatment>
</document>