198 lines
39 KiB
XML
198 lines
39 KiB
XML
<document id="EE19157B40B732521CEA0E1C1466D46C" ID-DOI="10.5281/zenodo.181357" ID-GBIF-Dataset="7b6b3cdb-b35e-4c7a-a7a8-8e1925b61fec" ID-ISSN="1175-5326" ID-Zenodo-Dep="181357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832480416" checkinUser="plazi" docAuthor="Liang, Ai-Ping & Wang, Rong-Rong" docDate="2008" docId="03E0878CFF913D47FF5EF9BDBABA6381" docLanguage="en" docName="zt01732p044.pdf" docOrigin="Zootaxa 1732" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Sogana robustocarina Liang & Wang, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="FFD9FFF4FF9A3D4BFFC9FF8FB951676B" masterDocTitle="A review of the Oriental planthopper genus Sogana Matsumura 1914 (Hemiptera: Fulgoromorpha: Tropiduchidae) with descriptions of three new species" masterLastPageNumber="44" masterPageNumber="29" pageNumber="40" updateTime="1698223042257" updateUser="plazi">
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<mods:mods id="E93CE8E4BB648521CF0791D4DB80F40B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A541B4721BE3CBB71D9A8300A7066D9B">
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<mods:title id="A59282F1E09E1C06278681EFB40C72F9">A review of the Oriental planthopper genus Sogana Matsumura 1914 (Hemiptera: Fulgoromorpha: Tropiduchidae) with descriptions of three new species</mods:title>
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<mods:name id="9B2BCA80D4574EB84D8942A925AC1027" type="personal">
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<mods:namePart id="0D6D87CE2B672984B93359CE06C0EFEA">Liang, Ai-Ping</mods:namePart>
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<mods:roleTerm id="73805EEC646110949C8B65A02A2DF4D8">Author</mods:roleTerm>
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<mods:namePart id="5FF0CD4D6B326BB48476646072C6720D">Wang, Rong-Rong</mods:namePart>
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<mods:typeOfResource id="FEB954E3C3E9059A0BED7DE750B3D424">text</mods:typeOfResource>
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<mods:title id="9084114C99132E7EDE5F0FEC0FA18A69">Zootaxa</mods:title>
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<mods:part id="09E55617375201F85E3CDD345FA2B295">
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<mods:date id="A1068F3956CAB02DD9E16A8FF54BD4DB">2008</mods:date>
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<mods:number id="E14A34DABD38A6A9D3C790129EDC9E46">1732</mods:number>
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<mods:start id="5E7F11E0E4E8842D7111D237A368263A">29</mods:start>
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<mods:end id="888C359FB85D6638F414056BD7AC49A6">44</mods:end>
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<mods:identifier id="DBD2872EE2DDFE1B0BF3BEE00F18D11A" type="DOI">10.5281/zenodo.181357</mods:identifier>
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<mods:identifier id="818E9D5C1C8D94BFC93C6F3958810390" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E0878CFF913D47FF5EF9BDBABA6381" ID-DOI="http://doi.org/10.5281/zenodo.6231120" ID-GBIF-Taxon="119358138" ID-Zenodo-Dep="6231120" LSID="urn:lsid:plazi:treatment:03E0878CFF913D47FF5EF9BDBABA6381" httpUri="http://treatment.plazi.org/id/03E0878CFF913D47FF5EF9BDBABA6381" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
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<subSubSection id="C3536511FF913D40FF5EF9BDBB0B6104" pageId="11" pageNumber="40" type="nomenclature">
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<paragraph id="8BF6369AFF913D40FF5EF9BDBB596127" blockId="11.[151,602,1586,1647]" box="[151,520,1586,1612]" pageId="11" pageNumber="40">
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<heading id="D0BE81F6FF913D40FF5EF9BDBB596127" bold="true" box="[151,520,1586,1612]" fontSize="11" level="1" pageId="11" pageNumber="40" reason="1">
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<emphasis id="B93DEA88FF913D40FF5EF9BDBB596127" bold="true" box="[151,520,1586,1612]" pageId="11" pageNumber="40">
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<taxonomicName id="4C494D19FF913D40FF5EF9BDB8F36127" ID-CoL="4XYY2" box="[151,418,1586,1612]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="robustocarina" status="sp. nov.">
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<emphasis id="B93DEA88FF913D40FF5EF9BDB8F36127" bold="true" box="[151,418,1586,1612]" italics="true" pageId="11" pageNumber="40">Sogana robustocarina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57F3FF913D40FE63F9BDBB596127" box="[426,520,1586,1612]" pageId="11" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF6369AFF913D40FF5EF9DABB0B6104" blockId="11.[151,602,1586,1647]" box="[151,602,1621,1647]" pageId="11" pageNumber="40">
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(
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<figureCitation id="13722A1FFF913D40FF56F9DAB9A26104" box="[159,243,1621,1647]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="11" pageNumber="40">Figs. 6</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FF36F9DAB8266104" box="[255,375,1621,1647]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">16–18, 22</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FE4AF9DAB8F26104" box="[387,419,1621,1647]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="11" pageNumber="40">27</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FE66F9DAB8AB6104" box="[431,506,1621,1647]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="11" pageNumber="40">40–43</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FDCEF9DABB766104" box="[519,551,1621,1647]" captionStart="FIGURES 48 – 51" captionStartId="14.[151,269,872,896]" captionTargetBox="[153,1421,202,871]" captionTargetId="figure@14.[151,1436,152,886]" captionTargetPageId="14" captionText="FIGURES 48 – 51. Sogana species: aedeagus (left view); same (right view). 48 – 49. S. pseudohopponis sp. nov. (holotype). 50 – 51. S. robustocarina sp. nov. (holotype)." httpUri="https://zenodo.org/record/181363/files/figure.png" pageId="11" pageNumber="40">50</figureCitation>
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, 51)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3536511FF913D47FF5EF92AB9876431" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40" type="description">
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<paragraph id="8BF6369AFF913D40FF5EF92ABDF161D4" blockId="11.[151,1436,1700,2007]" box="[151,1184,1700,1727]" pageId="11" pageNumber="40">
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<emphasis id="B93DEA88FF913D40FF5EF92AB87E61D4" bold="true" box="[151,303,1701,1727]" pageId="11" pageNumber="40">Description.</emphasis>
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Length (from apex of vertex to tip of fore wings): ď 10.5 mm, Ψ 10.5 mm.
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</paragraph>
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<paragraph id="8BF6369AFF913D40FF0CF942BAAC60EC" blockId="11.[151,1436,1700,2007]" pageId="11" pageNumber="40">
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General color tawny brown. Vertex (
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<figureCitation id="13722A1FFF913D40FDB2F942BB81618C" box="[635,720,1741,1767]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="11" pageNumber="40">Figs. 6</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FD17F942BA79618C" box="[734,808,1741,1767]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">16–18</figureCitation>
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) with blackish, I-shaped stripes at apex; central disc suffused with blackish, lateral margins black; lateral areas and all carinae on vertex, pronotum and mesonotum yellowish brown. Frons (
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<figureCitation id="13722A1FFF913D40FE3CF892BB1B605C" box="[501,586,1821,1847]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">Fig. 17</figureCitation>
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) with 6 pairs of carmine transverse bands, carinae on frons and clypeus orange to reddish. Gena (
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<figureCitation id="13722A1FFF913D40FE74F8CABB436034" box="[445,530,1861,1887]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">Fig. 18</figureCitation>
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) with a black patch. Ocelli (
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<figureCitation id="13722A1FFF913D40FC95F8CABAE16034" box="[860,944,1861,1887]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">Fig. 18</figureCitation>
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) suffused with red. Legs brown, trochanters fuscous, apex of femora reddish, postfemora with long fuscous stripes.
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</paragraph>
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<paragraph id="8BF6369AFF913D47FF0CF81ABB9067B1" blockId="11.[151,1436,1700,2007]" lastBlockId="12.[151,1437,152,1258]" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
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Vertex (
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<figureCitation id="13722A1FFF913D40FEEAF81AB82760C4" box="[291,374,1941,1967]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="11" pageNumber="40">Figs. 6</figureCitation>
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,
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<figureCitation id="13722A1FFF913D40FE4DF81AB8F360C4" box="[388,418,1941,1967]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">16</figureCitation>
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) with anterior margin broadly convex in dorsal view, lateral areas convex and ridged; median carina percurrent and broadly thickened, diverging medially. Fore wings (
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<figureCitation id="13722A1FFF913D40FBA9F832BDE660BC" box="[1120,1207,1981,2007]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="11" pageNumber="40">Fig. 22</figureCitation>
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) with 14–15 apical cells (sometimes the left and right fore wings are different) and 5 subapical cells. Spines of hind legs: hind tibia laterally 3, distally 6, basitarsus distally 6.
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</paragraph>
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<paragraph id="8BF6369AFF963D47FF0CFF67BB656611" blockId="12.[151,1437,152,1258]" pageId="12" pageNumber="41">
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<emphasis id="B93DEA88FF963D47FF0CFF67B80B666A" box="[197,346,232,257]" italics="true" pageId="12" pageNumber="41">Proportions.</emphasis>
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Vertex with ratio of median length to the widest breadth 1.11:1, frons with ratio of median length to the widest breadth 1.67:1, pronotum with ratio of width to median length 4.22:1, pronotum and mesonotum together medially about 1.4 times as long as median length of vertex, fore wings with ratio of length to maximum breadth 3.31:1.
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</paragraph>
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<paragraph id="8BF6369AFF963D47FF0CFE07B9876431" blockId="12.[151,1437,152,1258]" pageId="12" pageNumber="41">
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Male genitalia (
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<figureCitation id="13722A1FFF963D47FEB4FE07B8B166C9" box="[381,480,392,418]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="12" pageNumber="41">Figs. 27</figureCitation>
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,
|
||
<figureCitation id="13722A1FFF963D47FE22FE07BB6666C9" box="[491,567,392,418]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="12" pageNumber="41">40–43</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FD8AFE07BBDC66C9" box="[579,653,392,418]" captionStart="FIGURES 48 – 51" captionStartId="14.[151,269,872,896]" captionTargetBox="[153,1421,202,871]" captionTargetId="figure@14.[151,1436,152,886]" captionTargetPageId="14" captionText="FIGURES 48 – 51. Sogana species: aedeagus (left view); same (right view). 48 – 49. S. pseudohopponis sp. nov. (holotype). 50 – 51. S. robustocarina sp. nov. (holotype)." httpUri="https://zenodo.org/record/181363/files/figure.png" pageId="12" pageNumber="41">50, 51</figureCitation>
|
||
) with pygofer very narrow and high, wider dorsally than ventrally, with dorsal posterior margin angularly produced posteriorly in lateral view (
|
||
<figureCitation id="13722A1FFF963D47FBE0FE3FBDD266A1" box="[1065,1155,432,458]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="12" pageNumber="41">Fig. 27</figureCitation>
|
||
). Anal tube with apical margin oblique concave in dorsal view (
|
||
<figureCitation id="13722A1FFF963D47FDA7FE57BB946699" box="[622,709,472,498]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="12" pageNumber="41">Fig. 40</figureCitation>
|
||
), more or less bent ventrad in lateral view (
|
||
<figureCitation id="13722A1FFF963D47FB0BFE57BC486699" box="[1218,1305,472,498]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="12" pageNumber="41">Fig. 27</figureCitation>
|
||
). Gonostylus (
|
||
<figureCitation id="13722A1FFF963D47FF03FD8FB87D6571" box="[202,300,512,538]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="12" pageNumber="41">Figs. 27</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FEF0FD8FB8D46571" box="[313,389,512,538]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="12" pageNumber="41">42, 43</figureCitation>
|
||
) 3.31 times as long as broad in lateral view. Aedeagus (
|
||
<figureCitation id="13722A1FFF963D47FBD0FD8FBD2D6571" box="[1049,1148,512,538]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="12" pageNumber="41">Figs. 41</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FB40FD8FBD856571" box="[1161,1236,512,538]" captionStart="FIGURES 48 – 51" captionStartId="14.[151,269,872,896]" captionTargetBox="[153,1421,202,871]" captionTargetId="figure@14.[151,1436,152,886]" captionTargetPageId="14" captionText="FIGURES 48 – 51. Sogana species: aedeagus (left view); same (right view). 48 – 49. S. pseudohopponis sp. nov. (holotype). 50 – 51. S. robustocarina sp. nov. (holotype)." httpUri="https://zenodo.org/record/181363/files/figure.png" pageId="12" pageNumber="41">50, 51</figureCitation>
|
||
) with three stout processes: one anterodorsally directed process arising from left side at apical fifth, which is longer than threequarters of aedeagus, reaching approximately to base of aedeagus, with distinctly teeth along margin and a sharp protrusion on basal margin; the other anterodorsally directed process arising from dorsal margin at apical fifth, with 5–8 stout spines on ventral margin; the third process forked basally, arising below the left process on right side and directed posteriorly; endosoma membranous, distinctly expanded, with 2 processes: the dorsal one stubby, the ventral one slender and cuspidal; phallotrema long and narrow, exposed subapically on the left side (
|
||
<figureCitation id="13722A1FFF963D47FEE6FC97B8ED6459" box="[303,444,792,818]" captionStart="FIGURES 48 – 51" captionStartId="14.[151,269,872,896]" captionTargetBox="[153,1421,202,871]" captionTargetId="figure@14.[151,1436,152,886]" captionTargetPageId="14" captionText="FIGURES 48 – 51. Sogana species: aedeagus (left view); same (right view). 48 – 49. S. pseudohopponis sp. nov. (holotype). 50 – 51. S. robustocarina sp. nov. (holotype)." httpUri="https://zenodo.org/record/181363/files/figure.png" pageId="12" pageNumber="41">Figs. 50, 51</figureCitation>
|
||
); periandrium developed, ventrally protruding into a very short, triangular process at apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536511FF963D47FF0FFCE8BC3564B9" pageId="12" pageNumber="41" type="materials_examined">
|
||
<paragraph id="8BF6369AFF963D47FF0FFCE8BC3564B9" blockId="12.[151,1437,152,1258]" pageId="12" pageNumber="41">
|
||
<emphasis id="B93DEA88FF963D47FF0FFCE8B89464EA" bold="true" box="[198,453,871,897]" pageId="12" pageNumber="41">Material examined.</emphasis>
|
||
<typeStatus id="54F28838FF963D47FE1EFCE7BB1564E9" box="[471,580,872,898]" pageId="12" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||
ď, [
|
||
<collectingCountry id="F35E760AFF963D47FD43FCE7BA7C64E9" box="[650,813,872,898]" name="Malaysia" pageId="12" pageNumber="41">MALAYSIA</collectingCountry>
|
||
:] British N. Borneo, Kalabakan, Primary forest,
|
||
<date id="FFF7105AFF963D47FF5EFC1FB84D64C1" box="[151,284,912,938]" pageId="12" pageNumber="41" value="1958-11-15">15.XI.1958</date>
|
||
(T. C. Maa) (
|
||
<collectionCode id="ED58AE5FFF963D47FE09FC1FBB4964C1" LSID="urn:lsid:biocol.org:col:34697" box="[448,536,912,938]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="12" pageNumber="41">BPBM</collectionCode>
|
||
).
|
||
<typeStatus id="54F28838FF963D47FDE5FC1FBBF364C1" box="[556,674,912,938]" pageId="12" pageNumber="41" type="paratype">Paratypes</typeStatus>
|
||
. [
|
||
<collectingCountry id="F35E760AFF963D47FD7DFC1FBA0664C1" box="[692,855,912,938]" name="Malaysia" pageId="12" pageNumber="41">MALAYSIA</collectingCountry>
|
||
:] 1ď, British N. Borneo, Keningan,
|
||
<date id="FFF7105AFF963D47FAC9FC1FBCCD64C1" box="[1280,1436,912,938]" pageId="12" pageNumber="41" value="1959-01-12" valueMax="1959-01-17" valueMin="1959-01-12">12–17.I.1959</date>
|
||
(T. C. Maa); 1Ψ, British N. Borneo, Kalabakan, Primary forest,
|
||
<date id="FFF7105AFF963D47FC4BFC37BD5664B9" box="[898,1031,952,978]" pageId="12" pageNumber="41" value="1958-11-10">10.XI.1958</date>
|
||
(T. C. Maa) (both in
|
||
<collectionCode id="ED58AE5FFF963D47FAC8FC37BC0964B9" LSID="urn:lsid:biocol.org:col:34697" box="[1281,1368,952,978]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="12" pageNumber="41">BPBM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536511FF963D47FF0FFC50BC7F6491" box="[198,1326,991,1018]" pageId="12" pageNumber="41" type="etymology">
|
||
<paragraph id="8BF6369AFF963D47FF0FFC50BC7F6491" blockId="12.[151,1437,152,1258]" box="[198,1326,991,1018]" pageId="12" pageNumber="41">
|
||
<emphasis id="B93DEA88FF963D47FF0FFC50B8026492" bold="true" box="[198,339,991,1017]" pageId="12" pageNumber="41">Etymology.</emphasis>
|
||
This new species is named for the presence of a robust median carina on the vertex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536511FF963D47FF0FFB88BB036349" box="[198,594,1031,1058]" pageId="12" pageNumber="41" type="distribution">
|
||
<paragraph id="8BF6369AFF963D47FF0FFB88BB036349" blockId="12.[151,1437,152,1258]" box="[198,594,1031,1058]" pageId="12" pageNumber="41">
|
||
<emphasis id="B93DEA88FF963D47FF0FFB88B836634A" bold="true" box="[198,359,1031,1057]" pageId="12" pageNumber="41">Distribution.</emphasis>
|
||
<collectingCountry id="F35E760AFF963D47FEA7FB87B88B6349" box="[366,474,1032,1058]" name="Malaysia" pageId="12" pageNumber="41">Malaysia</collectingCountry>
|
||
(Borneo).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536511FF963D47FF0FFBA0BABA6381" pageId="12" pageNumber="41" type="discussion">
|
||
<paragraph id="8BF6369AFF963D47FF0FFBA0BABA6381" blockId="12.[151,1437,152,1258]" pageId="12" pageNumber="41">
|
||
<emphasis id="B93DEA88FF963D47FF0FFBA0B86E6322" bold="true" box="[198,319,1071,1097]" pageId="12" pageNumber="41">Remarks.</emphasis>
|
||
This species is externally similar to
|
||
<taxonomicName id="4C494D19FF963D47FD39FBBFBD646321" authority="Melichar, 1914" authorityName="Melichar" authorityYear="1914" box="[752,1077,1072,1098]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="stimulata">
|
||
<emphasis id="B93DEA88FF963D47FD39FBBFBA2A6322" box="[752,891,1072,1097]" italics="true" pageId="12" pageNumber="41">S. stimulata</emphasis>
|
||
<bibRefCitation id="EFD84B6BFF963D47FC4AFBBFBD646321" author="Melichar" box="[899,1077,1072,1098]" pageId="12" pageNumber="44" refString="Melichar, L. (1914) Monographie der Tropiduchinen (Homoptera). Verhandlungen des Naturforschenden Vereins in Brunn, 53, 1 - 145." type="journal article" year="1914">Melichar, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
from Mentawei Islands,
|
||
<collectingCountry id="F35E760AFF963D47FA94FBBFB9826319" name="Indonesia" pageId="12" pageNumber="41">Indonesia</collectingCountry>
|
||
in having the vertex with lateral areas between sublateral carinae and lateral carinae not distinctly depressed (shallowly depressed in
|
||
<taxonomicName id="4C494D19FF963D47FDFAFB0FBBEF63F2" box="[563,702,1152,1177]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="stimulata">
|
||
<emphasis id="B93DEA88FF963D47FDFAFB0FBBEF63F2" box="[563,702,1152,1177]" italics="true" pageId="12" pageNumber="41">S. stimulata</emphasis>
|
||
</taxonomicName>
|
||
; convex in
|
||
<taxonomicName id="4C494D19FF963D47FC82FB0FBD5E63F2" box="[843,1039,1152,1177]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="robustocarina" status="sp. nov.">
|
||
<emphasis id="B93DEA88FF963D47FC82FB0FBD5E63F2" box="[843,1039,1152,1177]" italics="true" pageId="12" pageNumber="41">S. robustocarina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA88FF963D47FBDEFBF0BD2763F2" bold="true" box="[1047,1142,1151,1177]" pageId="12" pageNumber="41">
|
||
<taxonomicNameLabel id="A20E57F3FF963D47FBDEFBF0BD2763F2" box="[1047,1142,1151,1177]" pageId="12" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). It can be distinguished from the latter species by the shape of the median carina of vertex (
|
||
<figureCitation id="13722A1FFF963D47FC78FB27BD5563A9" box="[945,1028,1192,1218]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="12" pageNumber="41">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FBD8FB27BD6063A9" box="[1041,1073,1192,1218]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="12" pageNumber="41">16</figureCitation>
|
||
), pigmentary pattern on vertex (
|
||
<figureCitation id="13722A1FFF963D47FF56FB5FB9A26381" box="[159,243,1232,1258]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="12" pageNumber="41">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FF36FB5FB81A6381" box="[255,331,1232,1258]" captionStart="FIGURES 16 – 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 – 23. Sogana species. 16 – 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 – 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="12" pageNumber="41">16–18</figureCitation>
|
||
), and its distinct aedeagal structure (
|
||
<figureCitation id="13722A1FFF963D47FD3EFB5FBA086381" box="[759,857,1232,1258]" captionStart="FIGURES 24 – 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 – 35. Sogana species. 24 – 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 – 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 – 31. S. clara sp. nov. (holotype). 32 – 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="12" pageNumber="41">Figs. 27</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FCACFB5FBAD46381" box="[869,901,1232,1258]" captionStart="FIGURES 36 – 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 – 47. Sogana species. 36 – 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 – 39. S. pseudohopponis sp. nov. (holotype). 40 – 43. S. robustocarina sp. nov. (holotype). 44 – 47. aedeagus (left view); same (right view). 44 – 45. S. clara sp. nov. (holotype). 46 – 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="12" pageNumber="41">41</figureCitation>
|
||
,
|
||
<figureCitation id="13722A1FFF963D47FC58FB5FBA8F6381" box="[913,990,1232,1258]" captionStart="FIGURES 48 – 51" captionStartId="14.[151,269,872,896]" captionTargetBox="[153,1421,202,871]" captionTargetId="figure@14.[151,1436,152,886]" captionTargetPageId="14" captionText="FIGURES 48 – 51. Sogana species: aedeagus (left view); same (right view). 48 – 49. S. pseudohopponis sp. nov. (holotype). 50 – 51. S. robustocarina sp. nov. (holotype)." httpUri="https://zenodo.org/record/181363/files/figure.png" pageId="12" pageNumber="41">50, 51</figureCitation>
|
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).
|
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</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |