treatments-xml/data/03/DF/87/03DF87DBEB67FFAFBAB0F580FBDAFBCF.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="6F1D894802E0DC683EF40CCD75CF2FB3">Cockscomb-shaped twighopper, Cladonotus bhaskari sp. n., a new and rare pygmy grasshopper species from Sri Lanka (Orthoptera: Tetrigidae: Cladonotinae)</mods:title>
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<mods:namePart id="29105BFCF191FB9C084C8D27D3C022D3">Deranja, Maks</mods:namePart>
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<mods:affiliation id="D619E087E850520C6059908F0B75BBA5">University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia &amp; maks. deranja @ gmail. com; https: // orcid. org / 0000 - 0002 - 5710 - 1916 &amp; karmela. adzic @ gmail. com; https: // orcid. org / 0000 - 0001 - 6223 - 4759 &amp; marko 09. pavlovic @ gmail. com; https: // orcid. org / 0000 - 0002 - 2956 - 6548 &amp; skejo. josip @ gmail. com; https: // orcid. org / 0000 - 0002 - 2554 - 4499</mods:affiliation>
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<mods:namePart id="1457A4989DF3A3D3E9EBB8F6E7BF23AA">Pavlović, Marko</mods:namePart>
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<mods:affiliation id="0CC6F351BC22013A76CDBBB38691A394">University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia &amp; maks. deranja @ gmail. com; https: // orcid. org / 0000 - 0002 - 5710 - 1916 &amp; karmela. adzic @ gmail. com; https: // orcid. org / 0000 - 0001 - 6223 - 4759 &amp; marko 09. pavlovic @ gmail. com; https: // orcid. org / 0000 - 0002 - 2956 - 6548 &amp; skejo. josip @ gmail. com; https: // orcid. org / 0000 - 0002 - 2554 - 4499</mods:affiliation>
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<mods:namePart id="EDC95B2EC2B146DAC6C10E15E45EA424">Skejo, Josip</mods:namePart>
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<mods:affiliation id="B770001394A8C511EA54B217B808FC07">University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia &amp; maks. deranja @ gmail. com; https: // orcid. org / 0000 - 0002 - 5710 - 1916 &amp; karmela. adzic @ gmail. com; https: // orcid. org / 0000 - 0001 - 6223 - 4759 &amp; marko 09. pavlovic @ gmail. com; https: // orcid. org / 0000 - 0002 - 2956 - 6548 &amp; skejo. josip @ gmail. com; https: // orcid. org / 0000 - 0002 - 2554 - 4499 &amp; Heinrich-Heine-Universität Düsseldorf, Institut für Molekulare Evolution, Universitätsstrasse 1, D- 40225 Düsseldorf, Germany &amp; skejo. josip @ gmail. com; https: // orcid. org / 0000 - 0002 - 2554 - 4499</mods:affiliation>
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<mods:date id="A367C78E48CDB5202099A0640C9B5297">2020</mods:date>
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<paragraph id="8BC936CDEB67FFAFBAB0F580FBA9FE1F" blockId="4.[151,1143,474,501]" box="[151,1143,474,501]" pageId="4" pageNumber="337">
<heading id="D08181A1EB67FFAFBAB0F580FBA9FE1F" bold="true" box="[151,1143,474,501]" fontSize="11" level="1" pageId="4" pageNumber="337" reason="1">
<emphasis id="B902EADFEB67FFAFBAB0F580FBA9FE1F" bold="true" box="[151,1143,474,501]" pageId="4" pageNumber="337">
I) Decurved Sri-Lankan twighopper,
<taxonomicName id="4C764D4EEB67FFAFB840F580FBA9FE1F" ID-CoL="VPQB" authority="(Saussure, 1862)" baseAuthorityName="Saussure" baseAuthorityYear="1862" box="[615,1143,474,501]" class="Insecta" family="Tetrigidae" genus="Cladonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="humbertianus">
<emphasis id="B902EADFEB67FFAFB840F580FC7DFE1E" bold="true" box="[615,931,474,501]" italics="true" pageId="4" pageNumber="337">Cladonotus humbertianus</emphasis>
(
<bibRefCitation id="EFE74B3CEB67FFAFB995F580FBB1FE1F" author="Saussure, H. de" box="[946,1135,474,500]" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure, 1862</bibRefCitation>
)
</taxonomicName>
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<subSubSection id="C36C6546EB67FFAFBAB0F645FD10FDDC" box="[151,718,543,567]" pageId="4" pageNumber="337" type="synonymic_list">
<paragraph id="8BC936CDEB67FFAFBAB0F645FD10FDDC" blockId="4.[151,1437,543,628]" box="[151,718,543,567]" pageId="4" pageNumber="337">
<taxonomicName id="4C764D4EEB67FFAFBAB0F645FD17FDDC" authority=": Saussure, 1862: 478" authorityName=": Saussure" authorityPageNumber="478" authorityYear="1862" box="[151,713,543,567]" class="Insecta" family="Tetrigidae" genus="Tettix" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="humbertianus" subGenus="Cladonotus">
<emphasis id="B902EADFEB67FFAFBAB0F645FE32FDDC" box="[151,492,543,567]" italics="true" pageId="4" pageNumber="337">Tettix (Cladonotus) humbertianus</emphasis>
:
<bibRefCitation id="EFE74B3CEB67FFAFBBDDF645FD17FDDC" author="Saussure, H. de" box="[506,713,543,567]" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure, 1862: 478</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C36C6546EB67FFAFBAB0F667FCB4FD9F" pageId="4" pageNumber="337" type="reference_group">
<paragraph id="8BC936CDEB67FFAFBAB0F667FCB4FD9F" blockId="4.[151,1437,543,628]" pageId="4" pageNumber="337">
<taxonomicName id="4C764D4EEB67FFAFBAB0F667FDBEFDBE" ID-CoL="VPQB" authority=": Walker 1871: 842" authorityName="Walker" authorityPageNumber="842" authorityYear="1871" box="[151,608,573,597]" class="Insecta" family="Tetrigidae" genus="Cladonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="humbertianus">
<emphasis id="B902EADFEB67FFAFBAB0F667FE43FDBE" box="[151,413,573,597]" italics="true" pageId="4" pageNumber="337">Cladonotus humbertianus</emphasis>
:
<bibRefCitation id="EFE74B3CEB67FFAFBB8DF667FDBEFDBE" author="Walker, F." box="[426,608,573,597]" pageId="4" pageNumber="337" refId="ref7086" refString="Walker, F. (1871) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum. Vol. 5. Department of Zoology, British Museum (Natural History), London, pp. 811 - 850." type="book" year="1871">Walker 1871: 842</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFE74B3CEB67FFAFB84DF667FCFAFDBE" author="Bolivar, I." box="[618,804,573,597]" pageId="4" pageNumber="337" pagination="175 - 313" refId="ref5734" refString="Bolivar, I. (1887) Essai sur les Acridiens de la tribu des Tettigidae. Annales de la Societe Entomologique de Belgique, 31, 175 - 313, pls. 4 - 5" type="journal article" year="1887">Bolívar 1887: 209</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFB909F667FC2AFDBE" author="Hancock, J. L." box="[814,1012,573,597]" pageId="4" pageNumber="337" pagination="97 - 157" refId="ref5995" refString="Hancock, J. L. (1904) The Tettigidae of Ceylon. Spolia Zeylanica, 2, 97 - 157." type="journal article" year="1904">Hancock 1904: 113</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFB9D8F667FB87FDBE" author="Hancock, J. L." box="[1023,1113,573,597]" pageId="4" pageNumber="337" pagination="1 - 79" refId="ref6018" refString="Hancock, J. L. (1907) Orthoptera Fam. Acridiidae. Subfam. Tetriginae. Genera Insectorum, 48, 1 - 79, pls. 1 - 4." type="journal article" year="1907">1907: 16</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFBE43F667FB61FDBE" author="Hancock, J. L." box="[1124,1215,573,597]" pageId="4" pageNumber="337" pagination="55 - 137" refId="ref6051" refString="Hancock, J. L. (1915) V. Indian Tetriginae (Acrydiinae). Records of the Indian Museum, 11, 55 - 137." type="journal article" year="1915">1915: 62</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFBEEEF667FA86FDBE" author="Kirby, W. F." box="[1225,1368,573,597]" pageId="4" pageNumber="337" refId="ref6255" refString="Kirby, W. F. (1910) A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae). The Trustees of the British Museum, London, 674 pp." type="book" year="1910">Kirby 1910: 9</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFBF44F667FF3CFD9F" author="Kirby, W. F." pageId="4" pageNumber="337" refId="ref6290" refString="Kirby, W. F. (1914) Fauna of British India, including Ceylon and Burma. Orthoptera (Acrididae). Taylor &amp; Francis, London, 276 pp. https: // doi. org / 10.5962 / bhl. title. 109305" type="book" year="1914">1914: 17</bibRefCitation>
, Sandrasagara 1950: 136,
<bibRefCitation id="EFE74B3CEB67FFAFBBCAF606FD78FD9F" author="Blackith, R. E." box="[493,678,604,628]" pageId="4" pageNumber="337" refId="ref5694" refString="Blackith, R. E. (1992) Tetrigidae (Insecta; Orthoptera) of Southeast Asia: Annotated catalogue with partial translated keys and bibliography. JAPAGA Press, Dublin, 248 pp., key LI-LIV." type="book" year="1992">Blackith 1992: 22</bibRefCitation>
,
<bibRefCitation id="EFE74B3CEB67FFAFB897F606FCB8FD9F" author="Hollier, J." box="[688,870,604,628]" pageId="4" pageNumber="337" pagination="203 - 219" refId="ref6115" refString="Hollier, J. (2013) An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Museum d'histoire naturelle de Geneve. Part 4: The Acridomorpha excluding the su- perfamily Acridoidea. Revue Suisse de Zoologie, 120 (2), 203 - 219." type="journal article" year="2013">Hollier 2013: 211</bibRefCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BC936CDEB67FFAFBAB0F6F8FDC0FCEE" blockId="4.[151,1437,674,1061]" pageId="4" pageNumber="337">
<emphasis id="B902EADFEB67FFAFBAB0F6F8FEF7FD57" bold="true" box="[151,297,674,700]" pageId="4" pageNumber="337">Distribution</emphasis>
.
<collectingCountry id="F361765DEB67FFAFBB19F6F8FE68FD57" box="[318,438,674,700]" name="Sri Lanka" pageId="4" pageNumber="337">Sri Lanka</collectingCountry>
: Peradeniya (see
<tableCitation id="C6F40376EB67FFAFB8ACF6F8FD37FD57" box="[651,745,674,700]" captionStart="TABLE 1" captionStartId="1.[152,245,1541,1565]" captionText="TABLE 1. Type material of the twighopper specimens (genus Cladonotus) with the accompanying data. Coordinates inside the brackets were approximately assigned by the authors (HT—holotype, ST—syntype)." pageId="4" pageNumber="337">
<emphasis id="B902EADFEB67FFAFB8ACF6F8FD37FD57" bold="true" box="[651,745,674,700]" pageId="4" pageNumber="337">Table 1</emphasis>
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), Trincomalee (
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,
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) (
<bibRefCitation id="EFE74B3CEB67FFAFBEC6F6F8FA4EFD56" author="Saussure, H. de" box="[1249,1424,674,701]" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure 1862</bibRefCitation>
), Kandy (for coordinates see
<tableCitation id="C6F40376EB67FFAFBBF2F69CFDF3FD0B" box="[469,557,710,736]" captionStart="TABLE 1" captionStartId="1.[152,245,1541,1565]" captionText="TABLE 1. Type material of the twighopper specimens (genus Cladonotus) with the accompanying data. Coordinates inside the brackets were approximately assigned by the authors (HT—holotype, ST—syntype)." pageId="4" pageNumber="337">
<emphasis id="B902EADFEB67FFAFBBF2F69CFDF3FD0B" bold="true" box="[469,557,710,736]" pageId="4" pageNumber="337">Table 1</emphasis>
</tableCitation>
), Gammaduwa: Mavusakanda (= probably Mousakanda Estate) (
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,
<geoCoordinate id="EE42500AEB67FFAFBAB0F6B0FEFFFCEE" box="[151,289,746,773]" degrees="80.696309" direction="east" orientation="longitude" pageId="4" pageNumber="337" precision="1" value="80.69631">E80.696309</geoCoordinate>
) (Sandrasagara 1950).
</paragraph>
<paragraph id="8BC936CDEB67FFAFBAE0F754FE66FCC3" blockId="4.[151,1437,674,1061]" box="[199,440,782,809]" pageId="4" pageNumber="337">
<emphasis id="B902EADFEB67FFAFBAE0F754FEFDFCC3" bold="true" box="[199,291,782,808]" pageId="4" pageNumber="337">Habitat</emphasis>
. Rainforests.
</paragraph>
</subSubSection>
<subSubSection id="C36C6546EB67FFAFBAE0F768FBDAFBCF" pageId="4" pageNumber="337" type="discussion">
<paragraph id="8BC936CDEB67FFAFBAE0F768FBDAFBCF" blockId="4.[151,1437,674,1061]" pageId="4" pageNumber="337">
<emphasis id="B902EADFEB67FFAFBAE0F768FED4FCA7" bold="true" box="[199,266,818,844]" pageId="4" pageNumber="337">Notes</emphasis>
. The decurved Sri-Lankan twighopper has been reported once (Sandrasagara 1950) since the description (
<bibRefCitation id="EFE74B3CEB67FFAFBAB8F70CFE99FC9A" author="Saussure, H. de" box="[159,327,854,881]" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure 1862</bibRefCitation>
).
<typeStatus id="54CD886FEB67FFAFBB7CF70CFE1CFC9B" box="[347,450,854,880]" pageId="4" pageNumber="337" type="syntype">Syntypes</typeStatus>
were collected more than 150 years ago, around Peradeniya and Trincomalee (
<bibRefCitation id="EFE74B3CEB67FFAFBF10F70CFF0DFC7E" author="Saussure, H. de" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure 1862</bibRefCitation>
). In the original description,
<bibRefCitation id="EFE74B3CEB67FFAFB807F720FD09FC7E" author="Saussure, H. de" box="[544,727,890,917]" pageId="4" pageNumber="337" pagination="469 - 494" refId="ref6615" refString="Saussure, H. de ([1862] 1861) Etudes sur quelques orthopteres du Musee de Geneve. Annales de la Societe Entomologique de France, 4 (1), 469 - 494." type="journal article" year="1862">Saussure (1862)</bibRefCitation>
placed this species in the genus
<taxonomicName id="4C764D4EEB67FFAFBE75F721FB38FC7F" authority="Charp." authorityName="Charp." box="[1106,1254,890,916]" class="Insecta" family="Tetrigidae" genus="Tettix" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADFEB67FFAFBE75F721FB51FC7F" box="[1106,1167,891,916]" italics="true" pageId="4" pageNumber="337">Tettix</emphasis>
Charp.
</taxonomicName>
, newly founded subgenus
<taxonomicName id="4C764D4EEB67FFAFBB20F7C4FE56FC53" authorityName=": Walker" authorityYear="1871" box="[263,392,926,952]" class="Insecta" family="Tetrigidae" genus="Cladonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADFEB67FFAFBB20F7C4FE56FC53" box="[263,392,926,952]" italics="true" pageId="4" pageNumber="337">Cladonotus</emphasis>
</taxonomicName>
. Because the original combination was
<emphasis id="B902EADFEB67FFAFB969F7C5FB0EFC53" box="[846,1232,926,952]" italics="true" pageId="4" pageNumber="337">
<taxonomicName id="4C764D4EEB67FFAFB969F7C5FB12FC53" authorityName=": Saussure" authorityYear="1862" box="[846,1228,926,952]" class="Insecta" family="Tetrigidae" genus="Tettix" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="humbertianus" subGenus="Cladonotus">Tettix (Cladonotus) humbertianus</taxonomicName>
,
</emphasis>
the author should be written in brackets according to Article 51.3. of the International Code of the Zoological Nomenclature (
<bibRefCitation id="EFE74B3CEB67FFAFBF7EF798FF0DFBEB" author="ICZN &amp; International Commission on the Zoological Nomenclature" pageId="4" pageNumber="337" refId="ref6180" refString="ICZN [International Commission on the Zoological Nomenclature] (1999) International code of zoological nomenclature. International Trust for Zoological Nomenclature, London, 304 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). Besides the type material (MHNG), there are a few specimens of this species in the Colombo Museum,
<collectingCountry id="F361765DEB67FFAFBF5BF7BCFF03FBCF" name="Sri Lanka" pageId="4" pageNumber="337">Sri Lanka</collectingCountry>
(Sandrasagara 1950), which should be studied and digitalized in future.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>