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<document id="C4E7B7F3C5BC987B30DB0A710CF31CCD" ID-CLB-Dataset="117406" ID-DOI="10.35929/RSZ.0013" ID-GBIF-Dataset="8458d5d1-44ea-4c33-b453-b746ddaa2534" ID-ISSN="0035-418" ID-Zenodo-Dep="5743391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638275126813" checkinUser="carolina" docAuthor="Sabella, Giorgio, Cuccodoro, Giulio &amp; Kurbatov, Sergey A." docDate="2020" docId="03DE232DB040FFECFC0452DFD56AFE49" docLanguage="en" docName="RevueSuisseZool.127.1.129-156.pdf" docOrigin="Revue suisse de Zoologie 127 (1)" docSource="http://dx.doi.org/10.35929/rsz.0013" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Achilia denticornis Jeannel 1962" docType="treatment" docVersion="11" lastPageNumber="139" masterDocId="FFE75B55B049FFE0FFB15060D767FF90" masterDocTitle="A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="156" masterPageNumber="129" pageNumber="137" updateTime="1699471439041" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="EF0829538EDDD535FA70833DD591F863" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9F9F849F8B8B6CF34C266A2BB51E30A8">A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="F9326F12D086FB64C74CE0A07A4B3B8D" type="personal">
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<mods:namePart id="F09C3FD3656D57F5200C40FC7588C03C">Sabella, Giorgio</mods:namePart>
<mods:affiliation id="FD51E83BF6CD29819E0692218CE7704E">Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dell’Università - sezione Biologia Animale, via</mods:affiliation>
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<mods:namePart id="C86F6BF9D325EC3F3AD9E837098EF379">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="754BB591EBA750909E17804F00A64A51">Muséum d’histoire naturelle, CP 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
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<mods:namePart id="41C49706844B2479E1AD067AA3E904A6">Kurbatov, Sergey A.</mods:namePart>
<mods:affiliation id="9888DC75BF28E4AB009A6377BD6D907B">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail:</mods:affiliation>
</mods:name>
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<mods:title id="91A3C07D19E8A8F59A654F808896B00D">Revue suisse de Zoologie</mods:title>
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<mods:part id="022A76C97F7769B95F37E9CC1833DA2F">
<mods:date id="77AB7CBF6EE262A59CDA00642E1F2423">2020</mods:date>
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<treatment id="03DE232DB040FFECFC0452DFD56AFE49" ID-DOI="http://doi.org/10.5281/zenodo.5743399" ID-GBIF-Taxon="198967438" ID-Zenodo-Dep="5743399" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49" httpUri="http://treatment.plazi.org/id/03DE232DB040FFECFC0452DFD56AFE49" lastPageId="12" lastPageNumber="139" pageId="9" pageNumber="137">
<subSubSection id="C36DC1B0B040FFE9FC0452DFD240FD47" box="[949,1319,703,727]" pageId="9" pageNumber="137" type="nomenclature">
<paragraph id="8BC8923BB040FFE9FC0452DFD240FD47" blockId="9.[949,1319,703,759]" box="[949,1319,703,727]" pageId="9" pageNumber="137">
<heading id="D0802557B040FFE9FC0452DFD240FD47" bold="true" box="[949,1319,703,727]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="137" reason="2">
<taxonomicName id="4C77E9B8B040FFE9FC0452DFD240FD47" ID-CoL="977K" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" box="[949,1319,703,727]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FC0452DFD240FD47" bold="true" box="[949,1319,703,727]" pageId="9" pageNumber="137">
<emphasis id="B9034E29B040FFE9FC0452DFD3E1FD47" bold="true" box="[949,1158,703,727]" italics="true" pageId="9" pageNumber="137">Achilia denticornis</emphasis>
<bibRefCitation id="EFE6EFCAB040FFE9FB3C52A0D240FD47" author="Jeannel R." box="[1165,1319,704,727]" pageId="9" pageNumber="137" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B040FFE9FC095280D243FD67" box="[952,1316,736,759]" pageId="9" pageNumber="137" type="description">
<paragraph id="8BC8923BB040FFE9FC095280D243FD67" blockId="9.[949,1319,703,759]" box="[952,1316,736,759]" pageId="9" pageNumber="137">
<figureCitation id="134C8EBEB040FFE9FC095280D375FD67" box="[952,1042,736,759]" captionStart="Figs 7-11" captionStartId="11.[151,192,1996,2017]" captionTargetBox="[171,1406,1288,1952]" captionTargetId="figure-530@11.[152,1437,1264,1985]" captionTargetPageId="11" captionText="Figs 7-11. Aedeagi (7, 9) and parameres in lateral view (8, 10-11) of Achilia species. (7) A. denticornis. (8) A. denticornis, same specimen. (9) A. bicornis, specimen from Chiloé, Mocopulli. (10) A. bicornis, specimen from Osorno, Bahia Mansa. (11) A. bicornis, specimen from Cautín, Huerquehue National Park." figureDoi="http://doi.org/10.5281/zenodo.5743395" httpUri="https://zenodo.org/record/5743395/files/figure.png" pageId="9" pageNumber="137">Figs 7-8</figureCitation>
,
<figureCitation id="134C8EBEB040FFE9FBAC5280D35DFD67" box="[1053,1082,736,759]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="9" pageNumber="137">16</figureCitation>
,
<figureCitation id="134C8EBEB040FFE9FBF45280D305FD67" box="[1093,1122,736,759]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="9" pageNumber="137">21</figureCitation>
,
<figureCitation id="134C8EBEB040FFE9FBDC5280D3D5FD67" box="[1133,1202,736,759]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="9" pageNumber="137">38, 44</figureCitation>
,
<figureCitation id="134C8EBEB040FFE9FB0C5280D399FD67" box="[1213,1278,736,759]" captionStart="Figs 71-74" captionStartId="21.[151,192,2053,2074]" captionTargetBox="[151,1435,866,2011]" captionTargetId="figure-383@21.[151,1436,866,2014]" captionTargetPageId="21" captionText="Figs 71-74. Achilia denticornis. Male head in (71) dorsal, (72) lateral, (73) semilateral, and (74) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743421" httpUri="https://zenodo.org/record/5743421/files/figure.png" pageId="9" pageNumber="137">71-74</figureCitation>
,
<figureCitation id="134C8EBEB040FFE9FAB85280D243FD67" box="[1289,1316,736,759]" captionStart="Fig" captionStartId="25.[151,186,1909,1930]" captionTargetBox="[236,716,257,1880]" captionTargetId="figure-90@25.[236,687,254,1880]" captionTargetPageId="25" captionText="Fig. 83. Distribution map of Achilia grandiceps group. (u green diamonds) A. grandiceps. (red squares) A. delamarei. (p blue triangles) A. denticornis. (« fuchsia stars) A. jeanneli n. sp. (¤ squares edged in fuchsia) A. franzi n. sp. (q red inverted triangles) A. elguetai n. sp." figureDoi="http://doi.org/10.5281/zenodo.5743427" httpUri="https://zenodo.org/record/5743427/files/figure.png" pageId="9" pageNumber="137">83</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B040FFE9FC8E536ED3EAFCD3" pageId="9" pageNumber="137" type="reference_group">
<paragraph id="8BC8923BB040FFE9FC8E536ED3EAFCD3" blockId="9.[831,1437,782,835]" pageId="9" pageNumber="137">
<treatmentCitationGroup id="AB67B515B040FFE9FC8E536ED3EAFCD3" pageId="9" pageNumber="137">
<treatmentCitation id="0AD6B42AB040FFE9FC8E536ED3D2FCB3" author="Jeannel R." box="[831,1205,782,803]" page="421" pageId="9" pageNumber="137" year="1962">
<taxonomicName id="4C77E9B8B040FFE9FC8E536ED3D2FCB3" ID-CoL="977K" authority="Jeannel, 1962: 421" authorityName="Jeannel" authorityPageNumber="421" authorityYear="1962" box="[831,1205,782,803]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FC8E536ED490FCB3" box="[831,1015,782,803]" italics="true" pageId="9" pageNumber="137">Achilia denticornis</emphasis>
<bibRefCitation id="EFE6EFCAB040FFE9FC4F536ED3D2FCB3" author="Jeannel R." box="[1022,1205,782,803]" pageId="9" pageNumber="137" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 421</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 423 figs 183 (head and antennae), 184 (aedeagus).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B040FFEAFC8E533BD5C3FECD" lastPageId="10" lastPageNumber="138" pageId="9" pageNumber="137" type="materials_examined">
<paragraph id="8BC8923BB040FFE9FC8E533BD39FFB01" blockId="9.[831,1437,859,1169]" pageId="9" pageNumber="137">
<materialsCitation id="3B1F9866B040FFE9FC8E533BD242FC62" ID-GBIF-Occurrence="3886337465" collectingDate="1958-10-03" collectionCode="MHNS" country="Chile" location="Region Los Lagos" pageId="9" pageNumber="137" specimenCount="28" specimenCount-male="1" stateProvince="Region Los Lagos" typeStatus="holotype">
<emphasis id="B9034E29B040FFE9FC8E533BD337FCE2" bold="true" box="[831,1104,859,882]" pageId="9" pageNumber="137">
Type material (
<specimenCount id="9D7159B2B040FFE9FC47533BD325FCE2" box="[1014,1090,859,882]" pageId="9" pageNumber="137" type="generic">27 ex.</specimenCount>
):
</emphasis>
SOUTHERN
<collectingCountry id="F360D2ABB040FFE9FB44533BD226FCE2" box="[1269,1345,859,882]" name="Chile" pageId="9" pageNumber="137" value="Chile">CHILI</collectingCountry>
:
<collectingRegion id="49B35CD9B040FFE9FAFE533BD4D8FC02" country="Chile" name="Los Lagos" pageId="9" pageNumber="137">Región Los Lagos</collectingRegion>
:
<collectingRegion id="49B35CD9B040FFE9FC62531BD304FC02" box="[979,1123,891,914]" pageId="9" pageNumber="137">Chiloé prov</collectingRegion>
.:
<collectionCode id="ED660AFEB040FFE9FBCA531BD3ABFC02" box="[1147,1228,891,914]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="9" pageNumber="137">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FB50531BD274FC02" box="[1249,1299,891,914]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B040FFE9FA9A531BD2F7FC02" box="[1323,1424,891,914]" pageId="9" pageNumber="137" type="holotype">holotype</typeStatus>
); labels verbatim “Type / Chepu,
<date id="FFC9B4FBB040FFE9FB1053FBD273FC22" box="[1185,1300,923,946]" pageId="9" pageNumber="137" value="1958-10-03">03.X.1958</date>
, G. Kuschel /
<taxonomicName id="4C77E9B8B040FFE9FCE653DBD351FC42" box="[855,1078,955,978]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FCE653DBD351FC42" box="[855,1078,955,978]" italics="true" pageId="9" pageNumber="137">Achillia denticornis</emphasis>
</taxonomicName>
/
<taxonomicName id="4C77E9B8B040FFE9FBEF53DBD3BFFC42" box="[1118,1240,955,978]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FBEF53DBD3BFFC42" box="[1118,1240,955,978]" italics="true" pageId="9" pageNumber="137">denticornis</emphasis>
</taxonomicName>
(handwritten by Jeannel) / CHILE, M.N.H.N., Typo, n. 1845”
</materialsCitation>
.
<materialsCitation id="3B1F9866B040FFE9FA9E53BBD31FFBA1" ID-GBIF-Occurrence="3886337495" collectingDate="1958-10-03" collectionCode="MNHN" collectorName="Kuschel" country="Chile" pageId="9" pageNumber="137" specimenCount="24" specimenCount-female="11" specimenCount-male="13" stateProvince="Chepu" typeStatus="paratype">
–
<collectionCode id="ED660AFEB040FFE9FAF253BBD2FFFC62" box="[1347,1432,987,1010]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="137">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FC8E539BD410FB82" box="[831,887,1019,1042]" pageId="9" pageNumber="137" type="male">13 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FC1F539BD482FB82" box="[942,997,1019,1042]" pageId="9" pageNumber="137" type="female">11 ♀</specimenCount>
(
<typeStatus id="54CC2C99B040FFE9FC44539BD306FB82" box="[1013,1121,1019,1042]" pageId="9" pageNumber="137" type="paratype">paratypes</typeStatus>
); labels verbatim “Paratype / Chepu,
<date id="FFC9B4FBB040FFE9FC22547AD361FBA1" box="[915,1030,1050,1073]" pageId="9" pageNumber="137" value="1958-10-03">
<collectingDate id="EF8D4D13B040FFE9FC22547AD361FBA1" box="[915,1030,1050,1073]" pageId="9" pageNumber="137" value="1958-10-03">03.X.1958</collectingDate>
</date>
,
<collectorName id="2682F7EDB040FFE9FBA3547AD308FBA1" box="[1042,1135,1050,1073]" pageId="9" pageNumber="137">Kuschel</collectorName>
”
</materialsCitation>
.
<materialsCitation id="3B1F9866B040FFE9FB32547AD4C0FBE1" ID-GBIF-Occurrence="3886337305" collectingDate="1958-10-04" collectionCode="MNHN" country="Chile" location="Kuschel" pageId="9" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="Kuschel" typeStatus="paratype">
–
<collectionCode id="ED660AFEB040FFE9FB26547AD38BFBA1" box="[1175,1260,1050,1073]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="137">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FB46547AD247FBA1" box="[1271,1312,1050,1073]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B040FFE9FA9E547AD2F6FBA1" box="[1327,1425,1050,1073]" pageId="9" pageNumber="137" type="paratype">paratype</typeStatus>
); labels verbatim “Paratype / Chepu,
<date id="FFC9B4FBB040FFE9FA93545AD2F0FBC1" box="[1314,1431,1082,1105]" pageId="9" pageNumber="137" value="1958-10-04">
<collectingDate id="EF8D4D13B040FFE9FA93545AD2F0FBC1" box="[1314,1431,1082,1105]" pageId="9" pageNumber="137" value="1958-10-04">04.X.1958</collectingDate>
</date>
,
<collectingRegion id="49B35CD9B040FFE9FC8E543AD4FAFBE1" box="[831,925,1114,1137]" pageId="9" pageNumber="137">Kuschel</collectingRegion>
”
</materialsCitation>
.
<materialsCitation id="3B1F9866B040FFE9FC07543AD393FB01" ID-GBIF-Occurrence="3886337354" collectingDate="1958-10-11" collectionCode="MNHN" country="Chile" pageId="9" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="Chepu" typeStatus="paratype">
–
<collectionCode id="ED660AFEB040FFE9FC61543AD342FBE1" box="[976,1061,1114,1137]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="137">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FB87543AD304FBE1" box="[1078,1123,1114,1137]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B040FFE9FBC6543AD3BDFBE1" box="[1143,1242,1114,1137]" pageId="9" pageNumber="137" type="paratype">paratype</typeStatus>
); labels verbatim “Paratype / Chepu,
<date id="FFC9B4FBB040FFE9FBA1541AD3E5FB01" box="[1040,1154,1146,1169]" pageId="9" pageNumber="137" value="1958-10-11">
<collectingDate id="EF8D4D13B040FFE9FBA1541AD3E5FB01" box="[1040,1154,1146,1169]" pageId="9" pageNumber="137" value="1958-10-11">11.X.1958</collectingDate>
</date>
, Kuschel”
</materialsCitation>
.
</paragraph>
<paragraph id="8BC8923BB040FFEAFC8E54C9D5C3FECD" blockId="9.[831,1437,1193,2077]" lastBlockId="10.[151,756,258,349]" lastPageId="10" lastPageNumber="138" pageId="9" pageNumber="137">
<materialsCitation id="3B1F9866B040FFE9FC8E54C9D4A1FAD0" ID-GBIF-Occurrence="3886337457" collectingDate="1958-10-03" collectionCode="MHNS" collectorName="G. Kuschel" country="Chile" location="Chepu" pageId="9" pageNumber="137" specimenCount="66" specimenCount-female="14" specimenCount-male="4" stateProvince="Region Los Lagos" typeStatus="paratype">
<emphasis id="B9034E29B040FFE9FC8E54C9D39BFB50" bold="true" box="[831,1276,1193,1216]" pageId="9" pageNumber="137">
Additional material examined (
<specimenCount id="9D7159B2B040FFE9FB1C54C9D393FB50" box="[1197,1268,1193,1216]" pageId="9" pageNumber="137" type="generic">48 ex.</specimenCount>
)
</emphasis>
: SOUTHERN
<collectingCountry id="F360D2ABB040FFE9FC8E54A9D4ECFB70" box="[831,907,1225,1248]" name="Chile" pageId="9" pageNumber="137">CHILI</collectingCountry>
:
<collectingRegion id="49B35CD9B040FFE9FC2854A9D30DFB70" box="[921,1130,1225,1248]" country="Chile" name="Los Lagos" pageId="9" pageNumber="137">Región Los Lagos</collectingRegion>
:
<collectingRegion id="49B35CD9B040FFE9FBC654A9D267FB70" box="[1143,1280,1225,1248]" pageId="9" pageNumber="137">Chiloé prov</collectingRegion>
.:
<collectionCode id="ED660AFEB040FFE9FAA054A9D205FB70" box="[1297,1378,1225,1248]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="9" pageNumber="137">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FAC154A9D2FBFB70" box="[1392,1436,1225,1248]" pageId="9" pageNumber="137" type="male">4 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FCDE5489D4C1FA90" box="[879,934,1257,1280]" pageId="9" pageNumber="137" type="female">14 ♀</specimenCount>
(mislabelled as
<typeStatus id="54CC2C99B040FFE9FBEB5489D3A7FA90" box="[1114,1216,1257,1280]" pageId="9" pageNumber="137" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C77E9B8B040FFE9FB595489D2E7FA90" box="[1256,1408,1257,1280]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FB595489D2E7FA90" box="[1256,1408,1257,1280]" italics="true" pageId="9" pageNumber="137">A. denticornis</emphasis>
</taxonomicName>
n. 1846-47, n. 1849 and n. 2053-67);
<location id="8EA8C4E0B040FFE9FB7D5569D272FAB0" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FB7D5569D272FAB0" box="[1228,1301,1289,1312]" country="Chile" name="Chepu" pageId="9" pageNumber="137" stateProvince="Region Los Lagos">Chepu</location>
;
<date id="FFC9B4FBB040FFE9FA955569D2FFFAB0" box="[1316,1432,1289,1312]" pageId="9" pageNumber="137" value="1958-10-03">
<collectingDate id="EF8D4D13B040FFE9FA955569D2FFFAB0" box="[1316,1432,1289,1312]" pageId="9" pageNumber="137" value="1958-10-03">03.X.1958</collectingDate>
</date>
;
<collectorName id="2682F7EDB040FFE9FC8E5549D4A5FAD0" box="[831,962,1321,1344]" pageId="9" pageNumber="137">G. Kuschel</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FC605549D3CEFAEF" ID-GBIF-Occurrence="3886337391" collectingDate="1958-10-04" collectionCode="MHNS" collectorName="G. Kuschel" location="Chepu" pageId="9" pageNumber="137" specimenCount="5" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
–
<collectionCode id="ED660AFEB040FFE9FC5A5549D35BFAD0" box="[1003,1084,1321,1344]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="9" pageNumber="137">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FBFC5549D31DFAD0" box="[1101,1146,1321,1344]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FB0B5549D38FFAD0" box="[1210,1256,1321,1344]" pageId="9" pageNumber="137" type="female">4 ♀</specimenCount>
(mislabelled as
<typeStatus id="54CC2C99B040FFE9FC8E5528D4C0FACF" box="[831,935,1352,1375]" pageId="9" pageNumber="137" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C77E9B8B040FFE9FC6F5528D3E7FACF" box="[990,1152,1352,1375]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="137" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B040FFE9FC6F5528D3E7FACF" box="[990,1152,1352,1375]" italics="true" pageId="9" pageNumber="137">A. denticornis</emphasis>
</taxonomicName>
n. 1848 and 2068-71);
<location id="8EA8C4E0B040FFE9FC8E5508D4EEFAEF" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FC8E5508D4EEFAEF" box="[831,905,1384,1407]" country="Chile" name="Chepu" pageId="9" pageNumber="137" stateProvince="Los Lagos">Chepu</location>
;
<date id="FFC9B4FBB040FFE9FC2A5508D377FAEF" box="[923,1040,1384,1407]" pageId="9" pageNumber="137" value="1958-10-04">
<collectingDate id="EF8D4D13B040FFE9FC2A5508D377FAEF" box="[923,1040,1384,1407]" pageId="9" pageNumber="137" value="1958-10-04">04.X.1958</collectingDate>
</date>
;
<collectorName id="2682F7EDB040FFE9FB935508D3C1FAEF" box="[1058,1190,1384,1407]" pageId="9" pageNumber="137">G. Kuschel</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FB075508D2FAFA0F" ID-GBIF-Occurrence="3886337503" collectingDate="1983-02-02" collectionCode="MHNG" collectorName="T. Cekalovic" location="Mocopulli" municipality="Isla Chiloe" pageId="9" pageNumber="137" specimenCount="7" specimenCount-female="5" specimenCount-male="2">
–
<collectionCode id="ED660AFEB040FFE9FB605508D241FAEF" box="[1233,1318,1384,1407]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="137" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FA885508D20FFAEF" box="[1337,1384,1384,1407]" pageId="9" pageNumber="137" type="male">2 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FC8E55E8D40AFA0F" box="[831,877,1416,1439]" pageId="9" pageNumber="137" type="female">5 ♀</specimenCount>
;
<collectingMunicipality id="6BAC0841B040FFE9FC3055E8D49AFA0F" box="[897,1021,1416,1439]" pageId="9" pageNumber="137">Isla Chiloé</collectingMunicipality>
,
<location id="8EA8C4E0B040FFE9FBBF55E8D3E6FA0F" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FBBF55E8D3E6FA0F" box="[1038,1153,1416,1439]" country="Chile" municipality="Isla Chiloe" name="Mocopulli" pageId="9" pageNumber="137" stateProvince="Los Lagos">Mocopulli</location>
;
<date id="FFC9B4FBB040FFE9FB2355E8D390FA0F" box="[1170,1271,1416,1439]" pageId="9" pageNumber="137" value="1983-02-02">
<collectingDate id="EF8D4D13B040FFE9FB2355E8D390FA0F" box="[1170,1271,1416,1439]" pageId="9" pageNumber="137" value="1983-02-02">2.II.1983</collectingDate>
</date>
;
<collectorName id="2682F7EDB040FFE9FAB655E8D2FEFA0F" box="[1287,1433,1416,1439]" pageId="9" pageNumber="137">T. Cekalovic</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FC8E55C8D4BCFA4F" ID-GBIF-Occurrence="3886337543" collectingDate="1983-02-19" collectionCode="MHNG" collectorName="T. Cekalovic" location="Piruquina" municipality="Isla Chiloe" pageId="9" pageNumber="137" specimenCount="1" specimenCount-female="1">
–
<collectionCode id="ED660AFEB040FFE9FCEC55C8D4D4FA2F" box="[861,947,1448,1471]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="137" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FC7755C8D490FA2F" box="[966,1015,1448,1471]" pageId="9" pageNumber="137" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BAC0841B040FFE9FBBF55C8D3F7FA2F" box="[1038,1168,1448,1471]" pageId="9" pageNumber="137">Isla Chiloé</collectingMunicipality>
,
<location id="8EA8C4E0B040FFE9FB1555C8D276FA2F" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FB1555C8D276FA2F" box="[1188,1297,1448,1471]" country="Chile" municipality="Isla Chiloe" name="Piruquina" pageId="9" pageNumber="137" stateProvince="Los Lagos">Piruquina</location>
;
<date id="FFC9B4FBB040FFE9FA9555C8D2FFFA2F" box="[1316,1432,1448,1471]" pageId="9" pageNumber="137" value="1983-02-19">
<collectingDate id="EF8D4D13B040FFE9FA9555C8D2FFFA2F" box="[1316,1432,1448,1471]" pageId="9" pageNumber="137" value="1983-02-19">19.II.1983</collectingDate>
</date>
;
<collectorName id="2682F7EDB040FFE9FC8E55A8D4B0FA4F" box="[831,983,1480,1503]" pageId="9" pageNumber="137">T. Cekalovic</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FC5D55A8D4B2F90E" ID-GBIF-Occurrence="3886337446" collectedFrom="leaf &amp; log" collectingDate="1997-01-27" collectingMethod="berlese" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" elevation="310" latitude="-41.165833" location="Vicente Perez Rosales National Park" longLatPrecision="11" longitude="-72.513336" municipality="Llanquihue prov" pageId="9" pageNumber="137" specimenCode="FMHD #97-38" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
–
<collectingMunicipality id="6BAC0841B040FFE9FBBB55A8D3A9FA4F" box="[1034,1230,1480,1503]" pageId="9" pageNumber="137">Llanquihue prov</collectingMunicipality>
.:
<collectionCode id="ED660AFEB040FFE9FB5655A8D252FA4F" box="[1255,1333,1480,1503]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMNH</collectionCode>
(
<specimenCode id="DBD13A40B040FFE9FAFC55A8D4E8FA6F" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMHD #97-38</specimenCode>
);
<specimenCount id="9D7159B2B040FFE9FC145588D4B7FA6F" box="[933,976,1512,1535]" pageId="9" pageNumber="137" type="male">2 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FBB85588D353FA6F" box="[1033,1076,1512,1535]" pageId="9" pageNumber="137" type="female">1 ♀</specimenCount>
;
<location id="8EA8C4E0B040FFE9FBF55588D414F98F" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FBF55588D414F98F" country="Chile" latitude="-41.165833" longLatPrecision="11" longitude="-72.513336" municipality="Llanquihue prov" name="Vicente Perez Rosales National Park" pageId="9" pageNumber="137" stateProvince="Los Lagos">Vicente Perez Rosales National Park</location>
,
<location id="8EA8C4E0B040FFE9FC325668D3C6F98F" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FC325668D3C6F98F" box="[899,1185,1544,1567]" latitude="-41.165833" longLatPrecision="11" longitude="-72.513336" municipality="Llanquihue prov" name="SW slope Volcan Osorno" pageId="9" pageNumber="137">SW slope Volcan Osorno</location>
,
<locationDeviation id="9A7A8B19B040FFE9FB015668D2FFF98F" box="[1200,1432,1544,1567]" pageId="9" pageNumber="137">km 4 to La Burbuja</locationDeviation>
;
<geoCoordinate id="EE43F4FCB040FFE9FC8E5647D4A4F9AE" box="[831,963,1575,1598]" degrees="41" direction="south" minutes="09.95" orientation="latitude" pageId="9" pageNumber="137" precision="9" value="-41.165833">41° 09.95’S</geoCoordinate>
<geoCoordinate id="EE43F4FCB040FFE9FC7D5647D33BF9AE" box="[972,1116,1575,1598]" degrees="72" direction="west" minutes="30.80" orientation="longitude" pageId="9" pageNumber="137" precision="9" value="-72.513336">72° 30.80’W</geoCoordinate>
;
<quantity id="4C8F3FDEB040FFE9FBD85647D3D4F9AE" box="[1129,1203,1575,1598]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="137" unit="m" value="310.0">
<elevation id="005A7508B040FFE9FBD85647D3D4F9AE" box="[1129,1203,1575,1598]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="137" unit="m" value="310.0">310 m</elevation>
</quantity>
;
<date id="FFC9B4FBB040FFE9FB705647D24DF9AE" box="[1217,1322,1575,1598]" pageId="9" pageNumber="137" value="1997-01-27">
<collectingDate id="EF8D4D13B040FFE9FB705647D24DF9AE" box="[1217,1322,1575,1598]" pageId="9" pageNumber="137" value="1997-01-27">27.I.1997</collectingDate>
</date>
; site 1007 secondary valdivian rainforest w/
<emphasis id="B9034E29B040FFE9FB085627D35CF9EE" italics="true" pageId="9" pageNumber="137">
<taxonomicName id="4C77E9B8B040FFE9FB085627D2FBF9CE" box="[1209,1436,1607,1630]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="137" phylum="Tracheophyta" rank="species" species="dombeyi">Nothofagus dombeyi</taxonomicName>
- Eucryphia cordifolia,
</emphasis>
<collectingMethod id="5236EA2CB040FFE9FBF45607D3F4F9EE" box="[1093,1171,1639,1662]" pageId="9" pageNumber="137">berlese</collectingMethod>
,
<collectedFrom id="386BD1C6B040FFE9FB105607D270F9EE" box="[1185,1303,1639,1662]" pageId="9" pageNumber="137">leaf &amp; log</collectedFrom>
;
<collectorName id="2682F7EDB040FFE9FA955607D4B6F90E" pageId="9" pageNumber="137">A. Newton &amp; M. Thayer</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FC6D56E7D3FFF88E" ID-GBIF-Occurrence="3886337447" collectedFrom="leaf &amp; log litter" collectingDate="1997-01-04" collectingMethod="berlese" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" elevation="385" latitude="-41.5105" location="near SE end," longLatPrecision="11" longitude="-72.399666" municipality="Lago Chapo" pageId="9" pageNumber="137" specimenCode="FMHD #97-16" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
–
<collectionCode id="ED660AFEB040FFE9FC4356E7D358F90E" box="[1010,1087,1671,1694]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMNH</collectionCode>
(
<specimenCode id="DBD13A40B040FFE9FBFF56E7D392F90E" box="[1102,1269,1671,1694]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMHD #97-16</specimenCode>
);
<specimenCount id="9D7159B2B040FFE9FABB56E7D253F90E" box="[1290,1332,1671,1694]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FADA56E7D2F2F90E" box="[1387,1429,1671,1694]" pageId="9" pageNumber="137" type="female">2 ♀</specimenCount>
;
<collectingMunicipality id="6BAC0841B040FFE9FC8E56C7D4A1F92E" box="[831,966,1703,1726]" country="Chile" name="Lago Chapo" pageId="9" pageNumber="137" stateProvince="Los Lagos">Lago Chapo</collectingMunicipality>
,
<location id="8EA8C4E0B040FFE9FC6056C7D33DF92E" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FC6056C7D33DF92E" box="[977,1114,1703,1726]" latitude="-41.5105" longLatPrecision="11" longitude="-72.399666" municipality="Lago Chapo" name="near SE end," pageId="9" pageNumber="137">near SE end,</location>
<locationDeviation id="9A7A8B19B040FFE9FBD056C7D2FFF92E" box="[1121,1432,1703,1726]" pageId="9" pageNumber="137">km 9.9 on road from Rollizo</locationDeviation>
;
<geoCoordinate id="EE43F4FCB040FFE9FC8E56A7D4A4F94E" box="[831,963,1735,1758]" degrees="41" direction="south" minutes="30.63" orientation="latitude" pageId="9" pageNumber="137" precision="9" value="-41.5105">41° 30.63’S</geoCoordinate>
<geoCoordinate id="EE43F4FCB040FFE9FC7F56A7D339F94E" box="[974,1118,1735,1758]" degrees="72" direction="west" minutes="23.98" orientation="longitude" pageId="9" pageNumber="137" precision="9" value="-72.399666">72° 23.98’W</geoCoordinate>
;
<quantity id="4C8F3FDEB040FFE9FBDC56A7D3D0F94E" box="[1133,1207,1735,1758]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="9" pageNumber="137" unit="m" value="385.0">
<elevation id="005A7508B040FFE9FBDC56A7D3D0F94E" box="[1133,1207,1735,1758]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="9" pageNumber="137" unit="m" value="385.0">385 m</elevation>
</quantity>
;
<date id="FFC9B4FBB040FFE9FB7456A7D248F94E" box="[1221,1327,1735,1758]" pageId="9" pageNumber="137" value="1997-01-04">
<collectingDate id="EF8D4D13B040FFE9FB7456A7D248F94E" box="[1221,1327,1735,1758]" pageId="9" pageNumber="137" value="1997-01-04">04.I.1997</collectingDate>
</date>
; site 989, valdivian rainforest on steep slope,
<collectingMethod id="5236EA2CB040FFE9FB7C5687D27CF96E" box="[1229,1307,1767,1790]" pageId="9" pageNumber="137">berlese</collectingMethod>
,
<collectedFrom id="386BD1C6B040FFE9FA985687D412F88E" pageId="9" pageNumber="137">leaf &amp; log litter</collectedFrom>
;
<collectorName id="2682F7EDB040FFE9FC305767D3F2F88E" box="[897,1173,1799,1822]" pageId="9" pageNumber="137">A. Newton &amp; M. Thayer</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FB105767D4F5F8CD" ID-GBIF-Occurrence="3886337336" collectingDate="2013-12-13" collectingMethod="car net" collectionCode="PCTS" latitude="-41.2333" location="Aguas Calientes" longitude="-72.45" pageId="9" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="Osorno prov.">
–
<collectingRegion id="49B35CD9B040FFE9FB065767D22EF88E" box="[1207,1353,1799,1822]" pageId="9" pageNumber="137">Osorno prov.</collectingRegion>
:
<collectionCode id="ED660AFEB040FFE9FAE45767D2FFF88E" box="[1365,1432,1799,1822]" pageId="9" pageNumber="137">PCTS</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FC8E5746D40EF8AD" box="[831,873,1830,1853]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
;
<location id="8EA8C4E0B040FFE9FCC75746D34FF8AD" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FCC75746D34FF8AD" box="[886,1064,1830,1853]" country="Chile" longitude="-72.45" name="Aguas Calientes" pageId="9" pageNumber="137" stateProvince="Osorno prov.">Aguas Calientes</location>
, 40° 74’S 72° 27’W;
<date id="FFC9B4FBB040FFE9FAA35746D2FFF8AD" box="[1298,1432,1830,1853]" pageId="9" pageNumber="137" value="2013-12-13">
<collectingDate id="EF8D4D13B040FFE9FAA35746D2FFF8AD" box="[1298,1432,1830,1853]" pageId="9" pageNumber="137" value="2013-12-13">13.XII.2013</collectingDate>
</date>
;
<collectingMethod id="5236EA2CB040FFE9FC8E5726D4E9F8CD" box="[831,910,1862,1885]" pageId="9" pageNumber="137">car net</collectingMethod>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FC2A5726D279F8ED" ID-GBIF-Occurrence="3886337338" collectedFrom="litter layer" collectingDate="2013-12-14" collectionCode="MHNG" latitude="-41.2333" location="Aguas Calientes" longitude="-72.45" pageId="9" pageNumber="137" specimenCount="8" specimenCount-female="7" specimenCount-male="1">
–
<collectionCode id="ED660AFEB040FFE9FC035726D360F8CD" box="[946,1031,1862,1885]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="137" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B040FFE9FBA45726D327F8CD" box="[1045,1088,1862,1885]" pageId="9" pageNumber="137" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B040FFE9FBCB5726D3C2F8CD" box="[1146,1189,1862,1885]" pageId="9" pageNumber="137" type="female">7 ♀</specimenCount>
;
<location id="8EA8C4E0B040FFE9FB045726D20EF8CD" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FB045726D20EF8CD" box="[1205,1385,1862,1885]" country="Chile" longitude="-72.45" name="Aguas Calientes" pageId="9" pageNumber="137" stateProvince="Los Lagos">Aguas Calientes</location>
, 40° 74’S 72° 27’W;
<date id="FFC9B4FBB040FFE9FBB65706D3EAF8ED" box="[1031,1165,1894,1917]" pageId="9" pageNumber="137" value="2013-12-14">
<collectingDate id="EF8D4D13B040FFE9FBB65706D3EAF8ED" box="[1031,1165,1894,1917]" pageId="9" pageNumber="137" value="2013-12-14">14.XII.2013</collectingDate>
</date>
;
<collectedFrom id="386BD1C6B040FFE9FB115706D27DF8ED" box="[1184,1306,1894,1917]" pageId="9" pageNumber="137">litter layer</collectedFrom>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFE9FA9A5706D4D0F86D" ID-GBIF-Occurrence="3886337390" collectedFrom="leaf &amp; log litter" collectingDate="1982-12-20" collectingDateMax="1982-12-25" collectingDateMin="1982-12-20" collectingMethod="berlese" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" elevation="470" location="Antillanca road;" municipality="Puyehue National Park" pageId="9" pageNumber="137" specimenCount="1" specimenCount-female="1">
–
<collectionCode id="ED660AFEB040FFE9FAF65706D2FFF8ED" box="[1351,1432,1894,1917]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMNH</collectionCode>
;
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;
<collectingMunicipality id="6BAC0841B040FFE9FCCE57E6D3E5F80D" box="[895,1154,1926,1949]" pageId="9" pageNumber="137">Puyehue National Park</collectingMunicipality>
,
<location id="8EA8C4E0B040FFE9FB3E57E6D226F80D" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FB3E57E6D226F80D" box="[1167,1345,1926,1949]" municipality="Puyehue National Park" name="Antillanca road;" pageId="9" pageNumber="137">Antillanca road;</location>
<quantity id="4C8F3FDEB040FFE9FAFC57E6D2FFF80D" box="[1357,1432,1926,1949]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="9" pageNumber="137" unit="m" value="470.0">
<elevation id="005A7508B040FFE9FAFC57E6D2FFF80D" box="[1357,1432,1926,1949]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="9" pageNumber="137" unit="m" value="470.0">470 m</elevation>
</quantity>
;
<date id="FFC9B4FBB040FFE9FC8E57C6D48EF82D" box="[831,1001,1958,1981]" pageId="9" pageNumber="137" value="1982-12-20" valueMax="1982-12-25" valueMin="1982-12-20">
<collectingDate id="EF8D4D13B040FFE9FC8E57C6D48EF82D" box="[831,1001,1958,1981]" pageId="9" pageNumber="137" value="1982-12-20" valueMax="1982-12-25" valueMin="1982-12-20">20-25.XII.1982</collectingDate>
</date>
;
<location id="8EA8C4E0B040FFE9FC4857C6D3B4F82D" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FC4857C6D3B4F82D" box="[1017,1235,1958,1981]" municipality="Puyehue National Park" name="valdivian rainforest" pageId="9" pageNumber="137">valdivian rainforest</location>
,
<collectedFrom id="386BD1C6B040FFE9FB5357C6D2FEF82D" box="[1250,1433,1958,1981]" pageId="9" pageNumber="137">leaf &amp; log litter</collectedFrom>
,
<collectingMethod id="5236EA2CB040FFE9FC8E57A6D4EAF84D" box="[831,909,1990,2013]" pageId="9" pageNumber="137">berlese</collectingMethod>
, vouchers associated with larvae;
<collectorName id="2682F7EDB040FFE9FAB757A6D4D3F86D" pageId="9" pageNumber="137">A. Newton &amp; M. Thayer</collectorName>
.
</materialsCitation>
<materialsCitation id="3B1F9866B040FFEAFC0E5786D5C3FECD" ID-GBIF-Occurrence="3886337315" collectedFrom="leaf &amp; log litter" collectingDate="2002-12-19" collectingMethod="berlese" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer &amp; D. J. Clarke &amp; M. Chani" elevation="580" lastPageId="10" lastPageNumber="138" latitude="-40.6705" location="Puyehue National Park" longLatPrecision="11" longitude="-72.08683" pageId="9" pageNumber="137" specimenCode="FMHD #2002-90" specimenCount="1" specimenCount-female="1">
–
<collectionCode id="ED660AFEB040FFE9FC655786D346F86D" box="[980,1057,2022,2045]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMNH</collectionCode>
(
<specimenCode id="DBD13A40B040FFE9FB805786D394F86D" box="[1073,1267,2022,2045]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="137" type="Museum">FMHD #2002-90</specimenCode>
);
<specimenCount id="9D7159B2B040FFE9FAB65786D256F86D" box="[1287,1329,2022,2045]" pageId="9" pageNumber="137" type="female">1 ♀</specimenCount>
;
<location id="8EA8C4E0B040FFE9FAF15786D4BAF78D" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FAF15786D4BAF78D" latitude="-40.6705" longLatPrecision="11" longitude="-72.08683" name="Puyehue National Park" pageId="9" pageNumber="137">Puyehue National Park</location>
,
<location id="8EA8C4E0B040FFE9FC5E5866D33DF78D" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B040FFE9FC5E5866D33DF78D" box="[1007,1114,2054,2077]" latitude="-40.6705" longLatPrecision="11" longitude="-72.08683" name="Ruta 215" pageId="9" pageNumber="137">Ruta 215</location>
;
<locationDeviation id="9A7A8B19B040FFE9FBDD5866D2FFF78D" box="[1132,1432,2054,2077]" pageId="9" pageNumber="137">km 4.5 of Aduana station</locationDeviation>
;
<geoCoordinate id="EE43F4FCB043FFEAFF265162D647FE89" box="[151,288,258,281]" degrees="40" direction="south" minutes="40.23" orientation="latitude" pageId="10" pageNumber="138" precision="9" value="-40.6705">40° 40.23’S</geoCoordinate>
<geoCoordinate id="EE43F4FCB043FFEAFE9C5162D6A5FE89" box="[301,450,258,281]" degrees="72" direction="west" minutes="05.21" orientation="longitude" pageId="10" pageNumber="138" precision="9" value="-72.08683">72° 05.21’W</geoCoordinate>
;
<quantity id="4C8F3FDEB043FFEAFE645162D544FE89" box="[469,547,258,281]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="10" pageNumber="138" unit="m" value="580.0">
<elevation id="005A7508B043FFEAFE645162D544FE89" box="[469,547,258,281]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="10" pageNumber="138" unit="m" value="580.0">580 m</elevation>
</quantity>
;
<date id="FFC9B4FBB043FFEAFD845162D5DBFE89" box="[565,700,258,281]" pageId="10" pageNumber="138" value="2002-12-19">
<collectingDate id="EF8D4D13B043FFEAFD845162D5DBFE89" box="[565,700,258,281]" pageId="10" pageNumber="138" value="2002-12-19">19.XII.2002</collectingDate>
</date>
; site 1071,
<location id="8EA8C4E0B043FFEAFF535144D6D9FEAB" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B043FFEAFF535144D6D9FEAB" box="[226,446,292,315]" latitude="-40.6705" longLatPrecision="11" longitude="-72.08683" name="valdivian rainforest" pageId="10" pageNumber="138">valdivian rainforest</location>
,
<collectingMethod id="5236EA2CB043FFEAFE615144D578FEAB" box="[464,543,292,315]" pageId="10" pageNumber="138">berlese</collectingMethod>
,
<collectedFrom id="386BD1C6B043FFEAFD805144D588FEAB" box="[561,751,292,315]" pageId="10" pageNumber="138">leaf &amp; log litter</collectedFrom>
;
<collectorName id="2682F7EDB043FFEAFF265126D668FECD" box="[151,271,326,349]" pageId="10" pageNumber="138">A. Newton</collectorName>
,
<collectorName id="2682F7EDB043FFEAFEAB5126D6ECFECD" box="[282,395,326,349]" pageId="10" pageNumber="138">M. Thayer</collectorName>
,
<collectorName id="2682F7EDB043FFEAFE275126D5F8FECD" box="[406,671,326,349]" pageId="10" pageNumber="138">D. J. Clarke &amp; M. Chani</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B043FFEBFF265117D2E2FE5A" lastPageId="11" lastPageNumber="139" pageId="10" pageNumber="138" type="description">
<paragraph id="8BC8923BB043FFEAFF265117D530FDC9" blockId="10.[151,756,375,635]" pageId="10" pageNumber="138">
<emphasis id="B9034E29B043FFEAFF265117D641FE1E" bold="true" box="[151,294,375,398]" pageId="10" pageNumber="138">Description:</emphasis>
Body
<quantity id="4C8F3FDEB043FFEAFECC5117D57AFE1E" box="[381,541,375,398]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="10" pageNumber="138" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.60-1.70 mm</quantity>
long, dark brown with reddish elytra; antennae and legs reddish; palpi yellowish. Pronotum with disc moderately convex; posterior portion of lateral margins subparallel. First abdominal tergite with diverging basal striae extending to about one-fourth of paratergal length, and separated at base by about one-third of tergal width.
</paragraph>
<paragraph id="8BC8923BB043FFEBFF265204D683FB01" blockId="10.[151,756,375,635]" lastBlockId="11.[151,756,258,1201]" lastPageId="11" lastPageNumber="139" pageId="10" pageNumber="138">
<emphasis id="B9034E29B043FFEAFF265204D7A9FDEB" box="[151,206,612,635]" italics="true" pageId="10" pageNumber="138">Male</emphasis>
: Head as in
<figureCitation id="134C8EBEB043FFEAFEEC5204D6B2FDEB" box="[349,469,612,635]" captionStart="Figs 71-74" captionStartId="21.[151,192,2053,2074]" captionTargetBox="[151,1435,866,2011]" captionTargetId="figure-383@21.[151,1436,866,2014]" captionTargetPageId="21" captionText="Figs 71-74. Achilia denticornis. Male head in (71) dorsal, (72) lateral, (73) semilateral, and (74) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743421" httpUri="https://zenodo.org/record/5743421/files/figure.png" pageId="10" pageNumber="138">Figs 71-74</figureCitation>
, with occipital region and basal half of frons raised as two protuberances, and separated by U-shaped median notch; lateral arms of notch extended anteriorly to form pubescent pointed process; anterior portion of frons very deeply excavated; median apophysis curved and directed backwards, sides rounded, originating at base of clypeus from ridge connecting base of antennal tubercles, tip in dorsal view triangular with rounded and pubescent sides; anterior margin of transverse clypeal ridge pointed at middle. Antennae (
<figureCitation id="134C8EBEB043FFEAFC095243D36FFDAA" box="[952,1032,547,570]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="10" pageNumber="138">Fig. 16</figureCitation>
) with scape and pedicel longer than wide, scape slightly swollen; antennomere III sligthly longer than wide; antennomere IV slightly wider than long; antennomere V with distal third of medial margin strongly widened and forming tooth-like process bearing six long apical bristles; antennomeres VI-VIII distinctly wider than long; antennomere IX transverse with distal third of mesal margin protruding to form small tooth; antennomere X wider than long with protruding mesal margin; antennomere XI elongate, distinctly longer than VII-X combined, with margins denticulate. Metaventrite with distal half not pubescent, shiny, raised at middle on apical two-thirds of length, this surface entirely divided by broad and deep median sulcus. Protibiae (
<figureCitation id="134C8EBEB042FFEBFD28522BD58BFDF2" box="[665,748,587,610]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="11" pageNumber="139">Fig. 38</figureCitation>
) slightly enlarged at middlle, medial margin forming tiny apical spine; mesotrochanters (
<figureCitation id="134C8EBEB042FFEBFDAF52ECD509FD33" box="[542,622,652,675]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="11" pageNumber="139">Fig. 31</figureCitation>
) with small tubercle near distal third of ventral margin; mesofemora (
<figureCitation id="134C8EBEB042FFEBFF2E52AED794FD75" box="[159,243,718,741]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="11" pageNumber="139">Fig. 31</figureCitation>
) with basal third of ventral margin covered by broad, short and thick setae, similar to those of
<taxonomicName id="4C77E9B8B042FFEBFF265370D649FCB7" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[151,302,784,807]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="139" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B042FFEBFF265370D649FCB7" box="[151,302,784,807]" italics="true" pageId="11" pageNumber="139">A. grandiceps</emphasis>
</taxonomicName>
; mesotibiae (
<figureCitation id="134C8EBEB042FFEBFE705370D577FCB7" box="[449,528,784,807]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="11" pageNumber="139">Fig. 44</figureCitation>
) with medial margin forming conspicuous stout spine at middle and with thin apical spine, without recurved setae. Aedeagus (
<figureCitation id="134C8EBEB042FFEBFD235332D58CFCF9" box="[658,747,850,873]" captionStart="Figs 7-11" captionStartId="11.[151,192,1996,2017]" captionTargetBox="[171,1406,1288,1952]" captionTargetId="figure-530@11.[152,1437,1264,1985]" captionTargetPageId="11" captionText="Figs 7-11. Aedeagi (7, 9) and parameres in lateral view (8, 10-11) of Achilia species. (7) A. denticornis. (8) A. denticornis, same specimen. (9) A. bicornis, specimen from Chiloé, Mocopulli. (10) A. bicornis, specimen from Osorno, Bahia Mansa. (11) A. bicornis, specimen from Cautín, Huerquehue National Park." figureDoi="http://doi.org/10.5281/zenodo.5743395" httpUri="https://zenodo.org/record/5743395/files/figure.png" pageId="11" pageNumber="139">Figs 7-8</figureCitation>
)
<quantity id="4C8F3FDEB042FFEBFF265313D654FC1A" box="[151,307,883,906]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="11" pageNumber="139" unit="mm" value="0.305" valueMax="0.31" valueMin="0.3">0.30-0.31 mm</quantity>
long; with suboval dorsal plate, dorsal longitudinal struts slightly divergent. Parameres wide without outer lobe, with enlarged and rounded apex, apex bearing three long and thin bristles. Copulatory pieces consisting of pair of stout lateral sclerites subequal in length, slightly pointed laterally and apically rounded, with pair of long, wide medial sclerites that are basally recurved and sclerotized, these sclerites frayed with numerous long and thin spines.
</paragraph>
<caption id="DF08C2B3B043FFEAFF265789D6CCF78B" ID-DOI="http://doi.org/10.5281/zenodo.5743393" ID-Zenodo-Dep="5743393" httpUri="https://zenodo.org/record/5743393/files/figure.png" pageId="10" pageNumber="138" startId="10.[151,192,2025,2046]" targetBox="[175,1415,715,1976]" targetPageId="10">
<paragraph id="8BC8923BB043FFEAFF265789D6CCF78B" blockId="10.[151,1440,2025,2075]" pageId="10" pageNumber="138">
Figs 1-6. Aedeagi of
<taxonomicName id="4C77E9B8B043FFEAFED75789D6CCF86E" authorityName="Reitter" authorityYear="1890" box="[358,427,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B043FFEAFED75789D6CCF86E" box="[358,427,2025,2046]" italics="true" pageId="10" pageNumber="138">Achilia</emphasis>
</taxonomicName>
species. (1)
<emphasis id="B9034E29B043FFEAFD955789D5D6F86E" box="[548,689,2025,2046]" italics="true" pageId="10" pageNumber="138">
<taxonomicName id="4C77E9B8B043FFEAFD955789D5CAF86E" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[548,685,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="grandiceps">A. grandiceps</taxonomicName>
.
</emphasis>
(2)
<emphasis id="B9034E29B043FFEAFD695789D43DF86E" box="[728,858,2025,2046]" italics="true" pageId="10" pageNumber="138">
<taxonomicName id="4C77E9B8B043FFEAFD695789D430F86E" box="[728,855,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="delamarei">A. delamarei</taxonomicName>
.
</emphasis>
(3)
<taxonomicName id="4C77E9B8B043FFEAFC305789D48EF86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[897,1001,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="jeanneli" status="sp. nov.">
<emphasis id="B9034E29B043FFEAFC305789D48EF86E" box="[897,1001,2025,2046]" italics="true" pageId="10" pageNumber="138">A. jeanneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B043FFEAFC5E5789D345F86E" box="[1007,1058,2025,2046]" pageId="10" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
(4)
<taxonomicName id="4C77E9B8B043FFEAFBFB5789D3FAF86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1098,1181,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B043FFEAFBFB5789D3FAF86E" box="[1098,1181,2025,2046]" italics="true" pageId="10" pageNumber="138">A. franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B043FFEAFB125789D3B0F86E" box="[1187,1239,2025,2046]" pageId="10" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
(5)
<taxonomicName id="4C77E9B8B043FFEAFB4E5789D201F86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1279,1382,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="elguetai" status="sp. nov.">
<emphasis id="B9034E29B043FFEAFB4E5789D201F86E" box="[1279,1382,2025,2046]" italics="true" pageId="10" pageNumber="138">A. elguetai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B043FFEAFADC5789D2C7F86E" box="[1389,1440,2025,2046]" pageId="10" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
(6)
<taxonomicName id="4C77E9B8B043FFEAFEA95866D6C1F78B" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[280,422,2054,2075]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="138" phylum="Arthropoda" rank="species" species="valdiviensis">A. valdiviensis</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC8923BB042FFEBFF2654FAD32DFEA9" blockId="11.[151,756,258,1201]" lastBlockId="11.[831,1436,258,313]" pageId="11" pageNumber="139">
<emphasis id="B9034E29B042FFEBFF2654FAD780FB21" box="[151,231,1178,1201]" italics="true" pageId="11" pageNumber="139">Female</emphasis>
: Similar to male except antennae and head not modified, with frons shallowly depressed behind scarcely impressed frontal sulcus.
</paragraph>
<paragraph id="8BC8923BB042FFEBFC8E5132D2E2FE5A" blockId="11.[831,1437,338,458]" pageId="11" pageNumber="139">
<emphasis id="B9034E29B042FFEBFC8E5132D491FEF9" bold="true" box="[831,1014,338,361]" pageId="11" pageNumber="139">Collecting data:</emphasis>
Collected from October to February in valdivian rainforest at elevations ranging from
<quantity id="4C8F3FDEB042FFEBFA8D5112D2E7FE19" box="[1340,1408,370,393]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="139" unit="m" value="310.0">310 m</quantity>
to
<specimenCount id="9D7159B2B042FFEBFC8E51F3D4EBFE3A" box="[831,908,403,426]" pageId="11" pageNumber="139" type="generic">
<quantity id="4C8F3FDEB042FFEBFC8E51F3D4EFFE3A" box="[831,904,403,426]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="139" unit="m" value="580.0">580 m</quantity>
.
</specimenCount>
All specimens came from sifted samples of leaf and log litter, except for
<specimenCount id="9D7159B2B042FFEBFBF751D3D3C0FE5A" box="[1094,1191,435,458]" pageId="11" pageNumber="139" type="male">one male</specimenCount>
collected by car net.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B042FFEBFC8E5182D2F4FD8A" pageId="11" pageNumber="139" type="distribution">
<paragraph id="8BC8923BB042FFEBFC8E5182D2F4FD8A" blockId="11.[831,1436,482,538]" pageId="11" pageNumber="139">
<emphasis id="B9034E29B042FFEBFC8E5182D4B3FE69" bold="true" box="[831,980,482,505]" pageId="11" pageNumber="139">Distribution:</emphasis>
The species is known only from the Los Lagos Región (southern Chile) (
<figureCitation id="134C8EBEB042FFEBFB295263D38FFD8A" box="[1176,1256,515,538]" captionStart="Fig" captionStartId="25.[151,186,1909,1930]" captionTargetBox="[236,716,257,1880]" captionTargetId="figure-90@25.[236,687,254,1880]" captionTargetPageId="25" captionText="Fig. 83. Distribution map of Achilia grandiceps group. (u green diamonds) A. grandiceps. (red squares) A. delamarei. (p blue triangles) A. denticornis. (« fuchsia stars) A. jeanneli n. sp. (¤ squares edged in fuchsia) A. franzi n. sp. (q red inverted triangles) A. elguetai n. sp." figureDoi="http://doi.org/10.5281/zenodo.5743427" httpUri="https://zenodo.org/record/5743427/files/figure.png" pageId="11" pageNumber="139">Fig. 83</figureCitation>
: blue triangles).
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B042FFECFC8E5252D56AFE49" lastPageId="12" lastPageNumber="140" pageId="11" pageNumber="139" type="discussion">
<paragraph id="8BC8923BB042FFEBFC8E5252D38DFBBF" blockId="11.[831,1437,562,1201]" pageId="11" pageNumber="139">
<emphasis id="B9034E29B042FFEBFC8E5252D4A0FDD9" bold="true" box="[831,967,562,585]" pageId="11" pageNumber="139">Comments:</emphasis>
The males of
<taxonomicName id="4C77E9B8B042FFEBFB035252D207FDD9" box="[1202,1376,562,585]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="139" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B042FFEBFB035252D207FDD9" box="[1202,1376,562,585]" italics="true" pageId="11" pageNumber="139">A. denticornis</emphasis>
</taxonomicName>
are easily distinguished from the other species of the
<taxonomicName id="4C77E9B8B042FFEBFC8E5213D4BEFD1A" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[831,985,627,650]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="139" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B042FFEBFC8E5213D4BEFD1A" box="[831,985,627,650]" italics="true" pageId="11" pageNumber="139">A. grandiceps</emphasis>
</taxonomicName>
group by the shape of their mesotibiae (
<figureCitation id="134C8EBEB042FFEBFCF652F3D4FEFD3A" box="[839,921,659,682]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="11" pageNumber="139">Fig. 44</figureCitation>
). The form of their head (
<figureCitation id="134C8EBEB042FFEBFB0052F3D24EFD3A" box="[1201,1321,659,682]" captionStart="Figs 71-74" captionStartId="21.[151,192,2053,2074]" captionTargetBox="[151,1435,866,2011]" captionTargetId="figure-383@21.[151,1436,866,2014]" captionTargetPageId="21" captionText="Figs 71-74. Achilia denticornis. Male head in (71) dorsal, (72) lateral, (73) semilateral, and (74) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743421" httpUri="https://zenodo.org/record/5743421/files/figure.png" pageId="11" pageNumber="139">Figs 71-74</figureCitation>
), antennae (
<figureCitation id="134C8EBEB042FFEBFCF952D4D4C5FD5B" box="[840,930,692,715]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="11" pageNumber="139">Fig. 16</figureCitation>
), protibiae (
<figureCitation id="134C8EBEB042FFEBFB8D52D4D3FFFD5B" box="[1084,1176,692,715]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="11" pageNumber="139">Fig. 38</figureCitation>
), and aedeagus (
<figureCitation id="134C8EBEB042FFEBFADD52D4D404FD7B" captionStart="Figs 7-11" captionStartId="11.[151,192,1996,2017]" captionTargetBox="[171,1406,1288,1952]" captionTargetId="figure-530@11.[152,1437,1264,1985]" captionTargetPageId="11" captionText="Figs 7-11. Aedeagi (7, 9) and parameres in lateral view (8, 10-11) of Achilia species. (7) A. denticornis. (8) A. denticornis, same specimen. (9) A. bicornis, specimen from Chiloé, Mocopulli. (10) A. bicornis, specimen from Osorno, Bahia Mansa. (11) A. bicornis, specimen from Cautín, Huerquehue National Park." figureDoi="http://doi.org/10.5281/zenodo.5743395" httpUri="https://zenodo.org/record/5743395/files/figure.png" pageId="11" pageNumber="139">Figs 7-8</figureCitation>
) are also diagnostic. The females of this species are characterized by their large head with the frons shallowly depressed behind the barely impressed frontal sulcus, antennomeres III-IV are longer than wide, antennomere V is distinctly longer than wide and is longer than all other antennomeres of the funicle. Medial margin of antennomere V slightly widened in the distal third.
<bibRefCitation id="EFE6EFCAB042FFEBFBA353D7D388FC5E" author="Jeannel R." box="[1042,1263,951,974]" pageId="11" pageNumber="139" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962: 423)</bibRefCitation>
claims that the antennomeres V of the female of
<taxonomicName id="4C77E9B8B042FFEBFB1C53B7D223FC7E" box="[1197,1348,983,1006]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="139" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B042FFEBFB1C53B7D223FC7E" box="[1197,1348,983,1006]" italics="true" pageId="11" pageNumber="139">A. denticornis</emphasis>
</taxonomicName>
possess a small tubercle with two bristles, but we did’nt found this feature in all the females examined.
</paragraph>
<paragraph id="8BC8923BB042FFECFC8E5458D56AFE49" blockId="11.[831,1437,562,1201]" lastBlockId="12.[151,756,258,473]" lastPageId="12" lastPageNumber="140" pageId="11" pageNumber="139">
In the original description
<bibRefCitation id="EFE6EFCAB042FFEBFBEF5458D254FBDF" author="Jeannel R." box="[1118,1331,1080,1103]" pageId="11" pageNumber="139" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962: 423)</bibRefCitation>
mentions that this species was described from
<specimenCount id="9D7159B2B042FFEBFAB35439D2FBFBE0" box="[1282,1436,1113,1136]" pageId="11" pageNumber="139" type="generic">50 specimens</specimenCount>
collected by Kuschel in Chepu on
<date id="FFC9B4FBB042FFEBFB0D5419D244FB00" box="[1212,1315,1145,1168]" pageId="11" pageNumber="139" value="1958-10-03">3.X.1958</date>
.
<collectorName id="2682F7EDB042FFEBFA9E5419D2FFFB00" box="[1327,1432,1145,1168]" pageId="11" pageNumber="139">Howewer</collectorName>
, upon examination of the
<typeStatus id="54CC2C99B042FFEBFBE454FAD3E4FB21" box="[1109,1155,1178,1201]" pageId="11" pageNumber="139">type</typeStatus>
series of
<collectorName id="2682F7EDB042FFEBFB4354FAD22EFB21" box="[1266,1353,1178,1201]" pageId="11" pageNumber="139">Kuschel</collectorName>
for the dates indicated by
<collectorName id="2682F7EDB045FFECFED55162D6D3FE89" box="[356,436,258,281]" pageId="12" pageNumber="140">Jeannel</collectorName>
we found only
<specimenCount id="9D7159B2B045FFECFDD35162D593FE89" box="[610,756,258,281]" pageId="12" pageNumber="140" type="generic">25 specimens</specimenCount>
(
<typeStatus id="54CC2C99B045FFECFF2F5142D79AFEA9" box="[158,253,290,313]" pageId="12" pageNumber="140" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CC2C99B045FFECFE835142D6FAFEA9" box="[306,413,290,313]" pageId="12" pageNumber="140" type="paratype">paratypes</typeStatus>
).
<location id="8EA8C4E0B045FFECFE1F5142D6BAFEA9" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B045FFECFE1F5142D6BAFEA9" box="[430,477,290,313]" name="Two" pageId="12" pageNumber="140">Two</location>
other males of the typical series, that are housed in MNHN, were collected by Kuschel in Chepu and are labeled as
<typeStatus id="54CC2C99B045FFECFD925102D5EBFEE9" box="[547,652,354,377]" pageId="12" pageNumber="140" type="paratype">paratypes</typeStatus>
, but were collected on different dates (i.e.
<specimenCount id="9D7159B2B045FFECFDBD51E2D53DFE09" box="[524,602,386,409]" pageId="12" pageNumber="140" type="male">1 male</specimenCount>
<typeStatus id="54CC2C99B045FFECFDD651E2D5AAFE09" box="[615,717,386,409]" pageId="12" pageNumber="140" type="paratype">paratypes</typeStatus>
on
<date id="FFC9B4FBB045FFECFF2651C2D66CFE29" box="[151,267,418,441]" pageId="12" pageNumber="140" value="1958-10-04">
<collectingDate id="EF8D4D13B045FFECFF2651C2D66CFE29" box="[151,267,418,441]" pageId="12" pageNumber="140" value="1958-10-04">04.X.1958</collectingDate>
</date>
; and
<specimenCount id="9D7159B2B045FFECFEF251C2D6ECFE29" box="[323,395,418,441]" pageId="12" pageNumber="140" type="male">1 male</specimenCount>
<typeStatus id="54CC2C99B045FFECFE2351C2D69FFE29" box="[402,504,418,441]" pageId="12" pageNumber="140" type="paratype">paratypes</typeStatus>
on
<date id="FFC9B4FBB045FFECFD9151C2D5F5FE29" box="[544,658,418,441]" pageId="12" pageNumber="140" value="1958-10-11">
<collectingDate id="EF8D4D13B045FFECFD9151C2D5F5FE29" box="[544,658,418,441]" pageId="12" pageNumber="140" value="1958-10-11">11.X.1958</collectingDate>
</date>
), a detail most likely overlooked by
<location id="8EA8C4E0B045FFECFE0451A2D56FFE49" LSID="urn:lsid:plazi:treatment:03DE232DB040FFECFC0452DFD56AFE49:8EA8C4E0B045FFECFE0451A2D56FFE49" box="[437,520,450,473]" name="Jeannel" pageId="12" pageNumber="140">Jeannel</location>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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