308 lines
38 KiB
XML
308 lines
38 KiB
XML
<document id="3B55FC0C77C5D8CBCB939D91A8E33027" ID-DOI="10.11646/zootaxa.4759.3.6" ID-GBIF-Dataset="a4b4f815-6b23-4430-8970-056cf3b60b80" ID-ISSN="1175-5326" ID-Zenodo-Dep="3741262" ID-ZooBank="7F3C0290-C75B-4814-BE7D-87762F4C4B6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586156741884" checkinUser="plazi" docAuthor="Mainda, Tobias" docDate="2020" docId="03D5D118FFFCFFCEB990FA1DE5FFC38A" docLanguage="en" docName="zootaxa.4759.3.6.pdf" docOrigin="Zootaxa 4759 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenus praedator Mainda 2020, sp. nov." docType="treatment" docUuid="E2C83390-CA78-4B2F-8A0E-CD52759D6FD8" docUuidSource="ZooBank" docVersion="6" lastPageNumber="410" masterDocId="FFECA960FFFDFFCBB907FFD0E179C531" masterDocTitle="Stenus praedator sp. nov. from the Philippines and a new species group in Stenus Latreille, 1797 (Coleoptera: Staphylinidae, Steninae)" masterLastPageNumber="412" masterPageNumber="405" pageNumber="406" updateTime="1698735779919" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4733FF750824B7E4B85E7E5AF4D5FF49">Stenus praedator sp. nov. from the Philippines and a new species group in Stenus Latreille, 1797 (Coleoptera: Staphylinidae, Steninae)</mods:title>
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<mods:name id="100741F1348A4C439A5634A854D18B88" type="personal">
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<mods:roleTerm id="D065E207D59C098126FB24A7850027AE">Author</mods:roleTerm>
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<mods:namePart id="11198A502EFFA519CF461123B0887755">Mainda, Tobias</mods:namePart>
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<mods:title id="54882EAFD427EADDE08300209BEAD9C9">Zootaxa</mods:title>
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<mods:part id="C8735BDEF7888C8D56697FCC1D5EF551">
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<mods:date id="64A91C99292B05A36E6CC51365CF3874">2020</mods:date>
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<mods:number id="74CD62D3EDDA26E7B72C0C731DE883F5">2020-04-03</mods:number>
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<mods:number id="585C73ECE0EDE5B8CBAFBC5C66CDC689">4759</mods:number>
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<mods:identifier id="049A9CB7117F1D11E486C652DB465714" type="DOI">10.11646/zootaxa.4759.3.6</mods:identifier>
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<treatment id="03D5D118FFFCFFCEB990FA1DE5FFC38A" ID-DOI="http://doi.org/10.5281/zenodo.3810319" ID-GBIF-Taxon="163437746" ID-Zenodo-Dep="3810319" LSID="urn:lsid:zoobank.org:act:E2C83390-CA78-4B2F-8A0E-CD52759D6FD8" httpUri="http://treatment.plazi.org/id/03D5D118FFFCFFCEB990FA1DE5FFC38A" lastPageId="5" lastPageNumber="410" pageId="1" pageNumber="406">
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<emphasis id="B908BC1CFFFCFFCAB990FA1DE0B3C0D6" bold="true" box="[151,458,1485,1512]" pageId="1" pageNumber="406">
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<taxonomicName id="4C7C1B8DFFFCFFCAB990FA1DE01CC0D9" ID-CoL="8YBBP" authority="Mainda, 2020" authorityName="Mainda" authorityYear="2020" box="[151,357,1485,1512]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="406" phylum="Arthropoda" rank="species" species="praedator" status="sp. nov.">
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<emphasis id="B908BC1CFFFCFFCAB990FA1DE01CC0D9" bold="true" box="[151,357,1485,1512]" italics="true" pageId="1" pageNumber="406">Stenus praedator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B0167FFFCFFCAB86AFA1DE0B3C0D6" box="[365,458,1485,1511]" pageId="1" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3663385FFFCFFCAB990FA21E2D1C31F" pageId="1" pageNumber="406" type="description">
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<paragraph id="8BC3600EFFFCFFCAB990FA21E00FC33A" blockId="1.[151,936,1485,1582]" box="[151,374,1521,1547]" pageId="1" pageNumber="406">
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<figureCitation id="13477C8BFFFCFFCAB990FA21E1A7C33A" box="[151,222,1521,1547]" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1972,1997]" captionTargetBox="[161,1414,1162,1945]" captionTargetId="figure@2.[158,1428,1161,1947]" captionTargetPageId="2" captionText="FIGURES 4–6. Holotypes of S. inflaticollis (photo: Field Museum of Natural History 2019) (4), S. ernstjuengeri (photo: Museum of Comparative Science 2019) (5) and S. praedator sp. nov. (6)." figureDoi="http://doi.org/10.5281/zenodo.3741266" httpUri="https://zenodo.org/record/3741266/files/figure.png" pageId="1" pageNumber="406">Figs 6</figureCitation>
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,
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<figureCitation id="13477C8BFFFCFFCAB9EDFA21E183C33A" box="[234,250,1521,1547]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1969,1994]" captionTargetBox="[159,1436,494,1934]" captionTargetId="figure@3.[151,1436,489,1944]" captionTargetPageId="3" captionText="FIGURES 7–9. Aedeagi of S. praedator sp. nov. (7), S. ernstjuengeri (8) and S. inflaticollis, holotype, from Puthz (1998) (9)." figureDoi="http://doi.org/10.5281/zenodo.3741268" httpUri="https://zenodo.org/record/3741268/files/figure.png" pageId="1" pageNumber="406">7</figureCitation>
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, 13, 18, 19
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</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C3663385FFFCFFCAB990F98DE079C236" pageId="1" pageNumber="406" type="materials_examined">
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<paragraph id="8BC3600EFFFCFFCAB990F98DE079C236" blockId="1.[151,1437,1628,2015]" pageId="1" pageNumber="406">
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<emphasis id="B908BC1CFFFCFFCAB990F98DE0C7C346" bold="true" box="[151,446,1628,1655]" pageId="1" pageNumber="406">Type material examined.</emphasis>
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<materialsCitation id="3B146A53FFFCFFCAB8C2F98DE079C236" ID-GBIF-Occurrence="2595809819" collectingDate="2018-03-28" collectingDateMax="2018-04-02" collectingDateMin="2018-03-28" collectorName="A. Shavrin" country="Philippines" county="Mount Hamiguitan Range Wildlife Sanctuary" elevation="500" latitude="7.75" location="Mindanao Isl." longLatPrecision="1303" longitude="125.38333" municipality="Mindanao" pageId="1" pageNumber="406" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54C7DEACFFFCFFCAB8C2F98DE354C346" box="[453,557,1629,1655]" pageId="1" pageNumber="406" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D7AAB87FFFCFFCABB34F98DE332C346" box="[563,587,1629,1655]" pageId="1" pageNumber="406" type="male">♂</specimenCount>
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: “
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<collectingCountry id="F36B209EFFFCFFCABB67F98DE39CC346" box="[608,741,1629,1655]" name="Philippines" pageId="1" pageNumber="406">Philippines</collectingCountry>
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:
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<location id="8EA336D5FFFCFFCABBE8F98DE2F4C346" LSID="urn:lsid:plazi:treatment:03D5D118FFFCFFCEB990FA1DE5FFC38A:8EA336D5FFFCFFCABBE8F98DE2F4C346" box="[751,909,1629,1655]" country="Philippines" county="Mount Hamiguitan Range Wildlife Sanctuary" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Mindanao" name="Mindanao Isl." pageId="1" pageNumber="406">Mindanao Isl.</location>
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,
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<location id="8EA336D5FFFCFFCABA9FF98DE5FCC346" LSID="urn:lsid:plazi:treatment:03D5D118FFFCFFCEB990FA1DE5FFC38A:8EA336D5FFFCFFCABA9FF98DE5FCC346" box="[920,1157,1629,1655]" country="Philippines" county="Mount Hamiguitan Range Wildlife Sanctuary" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Mindanao" name="Barangay Baganihan" pageId="1" pageNumber="406">Barangay Baganihan</location>
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,
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<location id="8EA336D5FFFCFFCABD88F98DE43EC346" LSID="urn:lsid:plazi:treatment:03D5D118FFFCFFCEB990FA1DE5FFC38A:8EA336D5FFFCFFCABD88F98DE43EC346" box="[1167,1351,1629,1655]" country="Philippines" county="Mount Hamiguitan Range Wildlife Sanctuary" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Mindanao" name="Marilog District" pageId="1" pageNumber="406">Marilog District</location>
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,
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<location id="8EA336D5FFFCFFCABC55F98DE1A4C3AA" LSID="urn:lsid:plazi:treatment:03D5D118FFFCFFCEB990FA1DE5FFC38A:8EA336D5FFFCFFCABC55F98DE1A4C3AA" country="Philippines" county="Mount Hamiguitan Range Wildlife Sanctuary" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Mindanao" name="Eagles Ridge" pageId="1" pageNumber="406">Eagles Ridge</location>
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,
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<geoCoordinate id="EE4806C9FFFCFFCAB9EDF950E039C3AA" box="[234,320,1664,1691]" degrees="7" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="406" precision="925" value="7.75">7°45’N</geoCoordinate>
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,
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<geoCoordinate id="EE4806C9FFFCFFCAB849F950E0C5C3AA" box="[334,444,1664,1691]" degrees="125" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="406" precision="925" value="125.38333">125°23’E</geoCoordinate>
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, 26.–
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<date id="FFC246CEFFFCFFCAB8FFF951E314C3AA" box="[504,621,1665,1691]" pageId="1" pageNumber="406" value="2018-03-28">
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<collectingDate id="EF86BF26FFFCFFCAB8FFF951E314C3AA" box="[504,621,1665,1691]" pageId="1" pageNumber="406" value="2018-03-28">28.3.2018</collectingDate>
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</date>
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, secondary broad-leaved forest, sifted from wet litter near small rocks, leg.
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<collectorName id="268905D8FFFCFFCAB9CEF975E031C38E" box="[201,328,1701,1727]" pageId="1" pageNumber="406">A. Shavrin</collectorName>
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” [ZMM].
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<specimenCount id="9D7AAB87FFFCFFCAB8C4F975E327C38E" box="[451,606,1701,1727]" pageId="1" pageNumber="406" type="female" typeStatus="Paratype">1 Paratype ♀</specimenCount>
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: “
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<collectingCountry id="F36B209EFFFCFFCABB70F975E385C38E" box="[631,764,1701,1727]" name="Philippines" pageId="1" pageNumber="406">Philippines</collectingCountry>
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:
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<collectingMunicipality id="6BA7FA74FFFCFFCABA0DF975E207C38E" box="[778,894,1701,1727]" pageId="1" pageNumber="406">Mindanao</collectingMunicipality>
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,
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<collectingCounty id="62A21882FFFCFFCABA8AF975E4E1C38E" box="[909,1432,1701,1727]" pageId="1" pageNumber="406">Mount Hamiguitan Range Wildlife Sanctuary</collectingCounty>
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,
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<geoCoordinate id="EE4806C9FFFCFFCAB990F919E05EC3D2" box="[151,295,1737,1763]" degrees="6" direction="north" minutes="43" orientation="latitude" pageId="1" pageNumber="406" precision="1" seconds="48.0" value="6.73">6°43’48.0”N</geoCoordinate>
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<geoCoordinate id="EE4806C9FFFCFFCAB829F919E0A3C3D2" box="[302,474,1737,1763]" degrees="126" direction="east" minutes="08" orientation="longitude" pageId="1" pageNumber="406" precision="1" seconds="24.0" value="126.14">126°08’24.0”E</geoCoordinate>
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,
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<quantity id="4C84CDEBFFFCFFCAB8E2F918E357C3D2" box="[485,558,1736,1763]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="406" unit="m" value="500.0">
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<elevation id="0051873DFFFCFFCAB8E2F918E357C3D2" box="[485,558,1736,1763]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="406" unit="m" value="500.0">500 m</elevation>
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</quantity>
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,
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<date id="FFC246CEFFFCFFCABB3DF919E387C3D2" box="[570,766,1737,1763]" pageId="1" pageNumber="406" value="2018-03-30" valueMax="2018-04-02" valueMin="2018-03-30">
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<collectingDate id="EF86BF26FFFCFFCABB3DF919E387C3D2" box="[570,766,1737,1763]" pageId="1" pageNumber="406" value="2018-03-30" valueMax="2018-04-02" valueMin="2018-03-30">30.III.–2.IV.2018</collectingDate>
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</date>
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, sifted from wet litter and debris near stream, leg.
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||
<collectorName id="268905D8FFFCFFCABC37F919E1D6C236" pageId="1" pageNumber="406">A. Shavrin</collectorName>
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” [cM].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3663385FFFCFFCEB9C0F8C0E2F1C08E" lastPageId="5" lastPageNumber="410" pageId="1" pageNumber="406" type="description">
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<paragraph id="8BC3600EFFFCFFCAB9C0F8C0E37EC242" blockId="1.[151,1437,1628,2015]" pageId="1" pageNumber="406">
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<emphasis id="B908BC1CFFFCFFCAB9C0F8C0E020C21A" bold="true" box="[199,345,1808,1835]" pageId="1" pageNumber="406">Description.</emphasis>
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Measurements of the HT (in mm): BL: 7.5, DE: 0.55, FBL: 3.6, EL: 1.23, EW: 1.15, HW: 1.25, PL: 1.27, PW: 0.9, SL: 0.8. Measurements of the PT (in mm): BL: 7.4, DE: 0.4, FBL: 3.5, EL: 1.22, EW: 1.17, HW: 1.25, PL: 1.27, PW: 0.9, SL: 0.8.
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</paragraph>
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||
<paragraph id="8BC3600EFFFCFFCAB9C0F8ADE294C28A" blockId="1.[151,1437,1628,2015]" pageId="1" pageNumber="406">
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||
Habitus as in
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||
<figureCitation id="13477C8BFFFCFFCAB858F8ADE0DAC2A6" box="[351,419,1917,1943]" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1972,1997]" captionTargetBox="[161,1414,1162,1945]" captionTargetId="figure@2.[158,1428,1161,1947]" captionTargetPageId="2" captionText="FIGURES 4–6. Holotypes of S. inflaticollis (photo: Field Museum of Natural History 2019) (4), S. ernstjuengeri (photo: Museum of Comparative Science 2019) (5) and S. praedator sp. nov. (6)." figureDoi="http://doi.org/10.5281/zenodo.3741266" httpUri="https://zenodo.org/record/3741266/files/figure.png" pageId="1" pageNumber="406">Fig. 6</figureCitation>
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||
. Micropterous, body black, shining, with weak metallic hue, completely without microsculpture; maxillary palpi, legs and antennae yellowish; labrum reddish-yellow.
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</paragraph>
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<paragraph id="8BC3600EFFFCFFC9B9C0F815E216C42E" blockId="1.[151,1437,1628,2015]" lastBlockId="2.[151,1436,153,287]" lastPageId="2" lastPageNumber="407" pageId="1" pageNumber="406">Median portion of head shiny, about twice as wide as lateral portions and with tiny puncture in its anterior centre. Anterior inner eye-margin with three to five punctures; median portion separated from lateral portions by a series of punctures, diameter of punctures about as wide as basal cross-section of antennomere III. Clypeus and labrum with dense silvery pubescence. Antennae long and slender, extending to posterior margin of pronotum when reflexed; antennomere X about twice as long as antennomere XI.</paragraph>
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||
<caption id="DF033086FFFFFFC9B990FBE6E50FC142" ID-DOI="http://doi.org/10.5281/zenodo.3741264" ID-Zenodo-Dep="3741264" httpUri="https://zenodo.org/record/3741264/files/figure.png" pageId="2" pageNumber="407" startId="2.[151,264,1078,1103]" targetBox="[152,1422,311,1052]" targetPageId="2">
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<paragraph id="8BC3600EFFFFFFC9B990FBE6E50FC142" blockId="2.[151,1436,1078,1139]" pageId="2" pageNumber="407">
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||
<emphasis id="B908BC1CFFFFFFC9B990FBE6E040C17E" bold="true" box="[151,313,1078,1103]" pageId="2" pageNumber="407">FIGURES 1–3.</emphasis>
|
||
Holotypes of
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||
<taxonomicName id="4C7C1B8DFFFFFFC9B8C1FBE7E32CC17F" authorityName="L. Benick" authorityYear="1929" box="[454,597,1078,1102]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="heterocerus">
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||
<emphasis id="B908BC1CFFFFFFC9B8C1FBE7E32CC17F" box="[454,597,1078,1102]" italics="true" pageId="2" pageNumber="407">S. heterocerus</emphasis>
|
||
</taxonomicName>
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||
(photo: Field Museum of Natural History 2019) (1),
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||
<taxonomicName id="4C7C1B8DFFFFFFC9BD61FBE7E46CC17F" authorityName="Puthz" authorityYear="1974" box="[1126,1301,1078,1102]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="thaumatocerus">
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||
<emphasis id="B908BC1CFFFFFFC9BD61FBE7E46CC17F" box="[1126,1301,1078,1102]" italics="true" pageId="2" pageNumber="407">S. thaumatocerus</emphasis>
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||
</taxonomicName>
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||
(photo: Field Museum of Natural History 2019) (2),
|
||
<taxonomicName id="4C7C1B8DFFFFFFC9BB25FB8BE303C143" authorityName="Puthz" authorityYear="2013" box="[546,634,1115,1138]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="quatei">
|
||
<emphasis id="B908BC1CFFFFFFC9BB25FB8BE303C143" box="[546,634,1115,1138]" italics="true" pageId="2" pageNumber="407">S. quatei</emphasis>
|
||
</taxonomicName>
|
||
(photo: Bishop Museum, James Boone 2019) (3).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF033086FFFFFFC9B990F864E224C2C0" ID-DOI="http://doi.org/10.5281/zenodo.3741266" ID-Zenodo-Dep="3741266" httpUri="https://zenodo.org/record/3741266/files/figure.png" pageId="2" pageNumber="407" startId="2.[151,264,1972,1997]" targetBox="[161,1414,1162,1945]" targetPageId="2">
|
||
<paragraph id="8BC3600EFFFFFFC9B990F864E224C2C0" blockId="2.[151,1437,1972,2033]" pageId="2" pageNumber="407">
|
||
<emphasis id="B908BC1CFFFFFFC9B990F864E042C2FD" bold="true" box="[151,315,1972,1997]" pageId="2" pageNumber="407">FIGURES 4–6.</emphasis>
|
||
Holotypes of
|
||
<taxonomicName id="4C7C1B8DFFFFFFC9B8CAF865E320C2FD" authorityName="Puthz" authorityYear="1998" box="[461,601,1972,1996]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="inflaticollis">
|
||
<emphasis id="B908BC1CFFFFFFC9B8CAF865E320C2FD" box="[461,601,1972,1996]" italics="true" pageId="2" pageNumber="407">S. inflaticollis</emphasis>
|
||
</taxonomicName>
|
||
(photo: Field Museum of Natural History 2019) (4),
|
||
<taxonomicName id="4C7C1B8DFFFFFFC9BD7EF865E463C2FD" authorityName="Puthz" authorityYear="1984" box="[1145,1306,1973,1996]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="ernstjuengeri">
|
||
<emphasis id="B908BC1CFFFFFFC9BD7EF865E463C2FD" box="[1145,1306,1973,1996]" italics="true" pageId="2" pageNumber="407">S. ernstjuengeri</emphasis>
|
||
</taxonomicName>
|
||
(photo: Museum of Comparative Science 2019) (5) and
|
||
<taxonomicName id="4C7C1B8DFFFFFFC9BB5AF809E3A2C2C1" authority="Mainda, 2020" authorityName="Mainda" authorityYear="2020" box="[605,731,2008,2032]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="407" phylum="Arthropoda" rank="species" species="praedator" status="sp. nov.">
|
||
<emphasis id="B908BC1CFFFFFFC9BB5AF809E3A2C2C1" box="[605,731,2008,2032]" italics="true" pageId="2" pageNumber="407">S. praedator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B908BC1CFFFFFFC9BBE5F809E24BC2C0" bold="true" box="[738,818,2009,2033]" pageId="2" pageNumber="407">
|
||
<taxonomicNameLabel id="A23B0167FFFFFFC9BBE5F809E24BC2C0" box="[738,818,2009,2033]" pageId="2" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(6).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC3600EFFFEFFC8B9C0FF49E5F1C456" blockId="3.[151,1436,153,467]" pageId="3" pageNumber="408">Pronotum (Fig. 18) 1.4 times as long as broad, broadest in the middle, sides towards anterior margin slightly convex, towards posterior margin concavely narrowed. Punctation coarse and fairly dense, except for few little elevated impunctate areas on each side, largest punctures in dorsal middle as large as maximal diameter of antennomere III in cross-section, lateral punctation somewhat less coarse, interstices much narrower than diameter of punctures. Legs slender, metatarsi less than half as long as metatibia; metatarsomere I as long as the combined length of metatarsomeres II–V and longer than metatarsomere V; metatarsomere IV deeply bilobed.</paragraph>
|
||
<paragraph id="8BC3600EFFFEFFC8B9C0FEA1E2BDC4E2" blockId="3.[151,1436,153,467]" pageId="3" pageNumber="408">Elytra 0.92 times narrow as head, 1.4 times as long as broad, humeral angles oblique, sides distinctly divergent posteriorly, suture unmodified. Punctation very coarse, diameter of punctures larger than apical cross-section of antennomere III, interstices mostly narrower than diameter of punctures.</paragraph>
|
||
<caption id="DF033086FFFEFFC8B990F861E4EFC2F8" ID-DOI="http://doi.org/10.5281/zenodo.3741268" ID-Zenodo-Dep="3741268" box="[151,1430,1969,1994]" httpUri="https://zenodo.org/record/3741268/files/figure.png" pageId="3" pageNumber="408" startId="3.[151,264,1969,1994]" targetBox="[159,1436,494,1934]" targetPageId="3">
|
||
<paragraph id="8BC3600EFFFEFFC8B990F861E4EFC2F8" blockId="3.[151,1430,1969,1994]" box="[151,1430,1969,1994]" pageId="3" pageNumber="408">
|
||
<emphasis id="B908BC1CFFFEFFC8B990F861E043C2FB" bold="true" box="[151,314,1969,1994]" pageId="3" pageNumber="408">FIGURES 7–9.</emphasis>
|
||
Aedeagi of
|
||
<taxonomicName id="4C7C1B8DFFFEFFC8B8B1F862E34DC2F8" authority="Mainda, 2020" authorityName="Mainda" authorityYear="2020" box="[438,564,1969,1993]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="408" phylum="Arthropoda" rank="species" species="praedator" status="sp. nov.">
|
||
<emphasis id="B908BC1CFFFEFFC8B8B1F862E34DC2F8" box="[438,564,1969,1993]" italics="true" pageId="3" pageNumber="408">S. praedator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B908BC1CFFFEFFC8BB3CF861E3F2C2F8" bold="true" box="[571,651,1969,1993]" pageId="3" pageNumber="408">
|
||
<taxonomicNameLabel id="A23B0167FFFEFFC8BB3CF861E3F2C2F8" box="[571,651,1969,1993]" pageId="3" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(7),
|
||
<taxonomicName id="4C7C1B8DFFFEFFC8BBBBF862E225C2F8" authorityName="Puthz" authorityYear="1984" box="[700,860,1970,1993]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="408" phylum="Arthropoda" rank="species" species="ernstjuengeri">
|
||
<emphasis id="B908BC1CFFFEFFC8BBBBF862E225C2F8" box="[700,860,1970,1993]" italics="true" pageId="3" pageNumber="408">S. ernstjuengeri</emphasis>
|
||
</taxonomicName>
|
||
(8) and
|
||
<taxonomicName id="4C7C1B8DFFFEFFC8BAB5F862E547C2F8" authorityName="Puthz" authorityYear="1998" box="[946,1086,1969,1993]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="408" phylum="Arthropoda" rank="species" species="inflaticollis">
|
||
<emphasis id="B908BC1CFFFEFFC8BAB5F862E547C2F8" box="[946,1086,1969,1993]" italics="true" pageId="3" pageNumber="408">S. inflaticollis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, from
|
||
<bibRefCitation id="EFED1DFFFFFEFFC8BDEFF861E412C2F8" author="Puthz, V." box="[1256,1387,1969,1994]" pageId="3" pageNumber="408" pagination="115 - 149" refId="ref3307" refString="Puthz, V. (1998) Neue Arten der Gattung Stenus Latreille aus der Orientalis, vorwiegend von den Philippinen (Coleoptera: Staphylinidae). Mitteilungen des internationalen entomologischen Vereins Frankfurt am Main, 23, 115 - 149." type="journal article" year="1998">Puthz (1998)</bibRefCitation>
|
||
(9).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF033086FFF9FFCFB990FB92E3B5C14E" ID-DOI="http://doi.org/10.5281/zenodo.3741270" ID-Zenodo-Dep="3741270" httpUri="https://zenodo.org/record/3741270/files/figure.png" pageId="4" pageNumber="409" startId="4.[151,264,1090,1115]" targetBox="[346,1247,188,1058]" targetPageId="4">
|
||
<paragraph id="8BC3600EFFF9FFCFB990FB92E3B5C14E" blockId="4.[151,1436,1090,1151]" pageId="4" pageNumber="409">
|
||
<emphasis id="B908BC1CFFF9FFCFB990FB92E02DC16A" bold="true" box="[151,340,1090,1115]" pageId="4" pageNumber="409">FIGURES 10–12.</emphasis>
|
||
Aedeagi of
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFB8C9FB93E35EC16B" authorityName="Puthz" authorityYear="2013" box="[462,551,1091,1114]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="quatei">
|
||
<emphasis id="B908BC1CFFF9FFCFB8C9FB93E35EC16B" box="[462,551,1091,1114]" italics="true" pageId="4" pageNumber="409">S. quatei</emphasis>
|
||
</taxonomicName>
|
||
, holotype, from
|
||
<bibRefCitation id="EFED1DFFFFF9FFCFBBD7FB92E22AC16A" author="Puthz, V." box="[720,851,1090,1115]" pageId="4" pageNumber="409" pagination="1279 - 1470" refId="ref3377" refString="Puthz, V. (2013) Ubersicht uber die orientalischen Arten der Gattung Stenus Latreille 1797 (Coleoptera, Staphylinidae). Linzer biologische Beitrage, 45, 1279 - 1470." type="journal article" year="2013">Puthz (2013)</bibRefCitation>
|
||
(10),
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFBA97FB93E538C16B" authorityName="Puthz" authorityYear="1974" box="[912,1089,1090,1114]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="thaumatocerus">
|
||
<emphasis id="B908BC1CFFF9FFCFBA97FB93E538C16B" box="[912,1089,1090,1114]" italics="true" pageId="4" pageNumber="409">S. thaumatocerus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, from
|
||
<bibRefCitation id="EFED1DFFFFF9FFCFBDEEFB92E415C16A" author="Puthz, V." box="[1257,1388,1090,1115]" pageId="4" pageNumber="409" pagination="154 - 171" refId="ref3238" refString="Puthz, V. (1974) Beitrage zur Kenntnis der Steninen CXLVII Uber einige Steninen von den Philippinen (Staphylinidae, Coleoptera). Philippia, 2, 154 - 171." type="journal article" year="1974">Puthz (1974)</bibRefCitation>
|
||
(11) and antennomere X of
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFB878FBB7E349C14F" authorityName="Puthz" authorityYear="1974" box="[383,560,1126,1150]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="thaumatocerus">
|
||
<emphasis id="B908BC1CFFF9FFCFB878FBB7E349C14F" box="[383,560,1126,1150]" italics="true" pageId="4" pageNumber="409">S. thaumatocerus</emphasis>
|
||
</taxonomicName>
|
||
, holotype (12).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC3600EFFF9FFCFB990FA25E379C303" blockId="4.[151,1436,1525,1586]" pageId="4" pageNumber="409">
|
||
<emphasis id="B908BC1CFFF9FFCFB990FA25E02DC33F" bold="true" box="[151,340,1525,1550]" pageId="4" pageNumber="409">FIGURES 13–18.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFB8E7FA26E308C33F" authorityName="L. Benick" authorityYear="1929" box="[480,625,1526,1550]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="heterocerus">
|
||
<emphasis id="B908BC1CFFF9FFCFB8E7FA26E308C33F" box="[480,625,1526,1550]" italics="true" pageId="4" pageNumber="409">S. heterocerus</emphasis>
|
||
</taxonomicName>
|
||
(13),
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFBBA9FA26E227C33F" authorityName="Puthz" authorityYear="1974" box="[686,862,1526,1550]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="thaumatocerus">
|
||
<emphasis id="B908BC1CFFF9FFCFBBA9FA26E227C33F" box="[686,862,1526,1550]" italics="true" pageId="4" pageNumber="409">S. thaumatocerus</emphasis>
|
||
</taxonomicName>
|
||
(14),
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFBA9CFA26E28DC33C" authorityName="Puthz" authorityYear="2013" box="[923,1012,1526,1549]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="quatei">
|
||
<emphasis id="B908BC1CFFF9FFCFBA9CFA26E28DC33C" box="[923,1012,1526,1549]" italics="true" pageId="4" pageNumber="409">S. quatei</emphasis>
|
||
</taxonomicName>
|
||
(15),
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFBD36FA26E5C7C33F" authorityName="Puthz" authorityYear="1998" box="[1073,1214,1526,1550]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="inflaticollis">
|
||
<emphasis id="B908BC1CFFF9FFCFBD36FA26E5C7C33F" box="[1073,1214,1526,1550]" italics="true" pageId="4" pageNumber="409">S. inflaticollis</emphasis>
|
||
</taxonomicName>
|
||
(16),
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFBDFBFA26E4E5C33C" authorityName="Puthz" authorityYear="1984" box="[1276,1436,1526,1549]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="ernstjuengeri">
|
||
<emphasis id="B908BC1CFFF9FFCFBDFBFA26E4E5C33C" box="[1276,1436,1526,1549]" italics="true" pageId="4" pageNumber="409">S. ernstjuengeri</emphasis>
|
||
</taxonomicName>
|
||
(17) and
|
||
<taxonomicName id="4C7C1B8DFFF9FFCFB9F4F9CAE008C303" authority="Mainda, 2020" authorityName="Mainda" authorityYear="2020" box="[243,369,1562,1586]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="409" phylum="Arthropoda" rank="species" species="praedator" status="sp. nov.">
|
||
<emphasis id="B908BC1CFFF9FFCFB9F4F9CAE008C303" box="[243,369,1562,1586]" italics="true" pageId="4" pageNumber="409">S. praedator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B908BC1CFFF9FFCFB87FF9CAE0B1C303" bold="true" box="[376,456,1562,1586]" pageId="4" pageNumber="409">
|
||
<taxonomicNameLabel id="A23B0167FFF9FFCFB87FF9CAE0B1C303" box="[376,456,1562,1586]" pageId="4" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(18).
|
||
</paragraph>
|
||
<caption id="DF033086FFF8FFCEB990FC7EE5ABC6F6" ID-DOI="http://doi.org/10.5281/zenodo.3741272" ID-Zenodo-Dep="3741272" box="[151,1234,942,967]" httpUri="https://zenodo.org/record/3741272/files/figure.png" pageId="5" pageNumber="410" startId="5.[151,250,942,967]" targetBox="[151,1436,181,916]" targetPageId="5">
|
||
<paragraph id="8BC3600EFFF8FFCEB990FC7EE5ABC6F6" blockId="5.[151,1234,942,967]" box="[151,1234,942,967]" pageId="5" pageNumber="410">
|
||
<emphasis id="B908BC1CFFF8FFCEB990FC7EE059C6F6" bold="true" box="[151,288,942,967]" pageId="5" pageNumber="410">FIGURE 19.</emphasis>
|
||
sternite VIII, tergite VIII, sternite IX and tergite X (left to right) of
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBAC9FC7FE502C6F7" authority="Mainda, 2020" authorityName="Mainda" authorityYear="2020" box="[974,1147,942,966]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="praedator" status="sp. nov.">
|
||
<emphasis id="B908BC1CFFF8FFCEBAC9FC7FE502C6F7" box="[974,1147,942,966]" italics="true" pageId="5" pageNumber="410">Stenus praedator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B0167FFF8FFCEBD85FC7FE5ABC6F6" box="[1154,1234,943,967]" pageId="5" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC3600EFFF8FFCEB9C0FC25E391C1AE" blockId="5.[151,1437,1013,1723]" pageId="5" pageNumber="410">Abdomen cylindrical; with short pubescence; basal impressions of tergites I–III very deep; punctation of basal impressions of tergites III and IV coarse; interstices much wider than diameter of punctures, their size about as large as an ommatidium at medial eye-margin up to the middle cross-section of antennomere V; punctation of tergites V–VII finer and sparser, interstices as wide as on tergite IV; punctation of tergite VIII denser than that of tergites III–VII, interstices wider than diameter of punctures.</paragraph>
|
||
<paragraph id="8BC3600EFFF8FFCEB9C0FB79E354C046" blockId="5.[151,1437,1013,1723]" pageId="5" pageNumber="410">
|
||
<emphasis id="B908BC1CFFF8FFCEB9C0FB79E072C1F2" bold="true" box="[199,267,1193,1219]" pageId="5" pageNumber="410">Male.</emphasis>
|
||
Femora slightly dilated. Metaventrite coarsely and densely punctured. Sternites III–VI unmodified, sternite VII ventrally flattened and medially more densely punctured than laterally, near posterior margin shallowly impressed; sternite VIII with a rounded notch in posterior sixth; sternite IX serrate apicolaterally; tergite X with convex posterior margin (
|
||
<figureCitation id="13477C8BFFF8FFCEB8BEFAC5E375C01E" box="[441,524,1301,1327]" captionStart="FIGURE 19" captionStartId="5.[151,250,942,967]" captionTargetBox="[151,1436,181,916]" captionTargetId="figure@5.[151,1436,181,916]" captionTargetPageId="5" captionText="FIGURE 19. sternite VIII, tergite VIII, sternite IX and tergite X (left to right) of Stenus praedator sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3741272" httpUri="https://zenodo.org/record/3741272/files/figure.png" pageId="5" pageNumber="410">Fig. 19</figureCitation>
|
||
). Apical portion of aedeagus (
|
||
<figureCitation id="13477C8BFFF8FFCEBA59FAC5E2DBC01E" box="[862,930,1301,1327]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1969,1994]" captionTargetBox="[159,1436,494,1934]" captionTargetId="figure@3.[151,1436,489,1944]" captionTargetPageId="3" captionText="FIGURES 7–9. Aedeagi of S. praedator sp. nov. (7), S. ernstjuengeri (8) and S. inflaticollis, holotype, from Puthz (1998) (9)." figureDoi="http://doi.org/10.5281/zenodo.3741268" httpUri="https://zenodo.org/record/3741268/files/figure.png" pageId="5" pageNumber="410">Fig. 7</figureCitation>
|
||
) acutely narrowed, expulsion-clasp with two “horns” (lower arrow); internal sac with long flagellum (upper arrow); parameres slightly extending beyond apex of median lobe, with six setae apically.
|
||
</paragraph>
|
||
<paragraph id="8BC3600EFFF8FFCEB9C0FA51E2F1C08E" blockId="5.[151,1437,1013,1723]" pageId="5" pageNumber="410">
|
||
<emphasis id="B908BC1CFFF8FFCEB9C0FA51E05CC0AA" bold="true" box="[199,293,1409,1435]" pageId="5" pageNumber="410">Female.</emphasis>
|
||
Legs unmodified. Median pubescence of sternite VII denser than in lateral portions; sternite VIII broadly rounded at posterior margin; valvifer serrate apicolaterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3663385FFF8FFCEB9C0FA18E069C3A6" pageId="5" pageNumber="410" type="diagnosis">
|
||
<paragraph id="8BC3600EFFF8FFCEB9C0FA18E069C3A6" blockId="5.[151,1437,1013,1723]" pageId="5" pageNumber="410">
|
||
<emphasis id="B908BC1CFFF8FFCEB9C0FA18E0B4C0D2" bold="true" box="[199,461,1480,1507]" pageId="5" pageNumber="410">Differential diagnosis.</emphasis>
|
||
The species is distinguished from the similar
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBAC0FA19E50EC0D2" authorityName="Puthz" authorityYear="1984" box="[967,1143,1481,1507]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="ernstjuengeri">
|
||
<emphasis id="B908BC1CFFF8FFCEBAC0FA19E50EC0D2" box="[967,1143,1481,1507]" italics="true" pageId="5" pageNumber="410">S. ernstjuengeri</emphasis>
|
||
</taxonomicName>
|
||
by its male sexual characters, the structure of the pronotum, and the sparser punctation of the abdomen. It is separated from
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBDDBFA3DE403C336" authorityName="L. Benick" authorityYear="1929" box="[1244,1402,1517,1543]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="heterocerus">
|
||
<emphasis id="B908BC1CFFF8FFCEBDDBFA3DE403C336" box="[1244,1402,1517,1543]" italics="true" pageId="5" pageNumber="410">S. heterocerus</emphasis>
|
||
</taxonomicName>
|
||
by its denser and coarser punctation and from
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBB78F9C1E238C31A" authorityName="Puthz" authorityYear="1974" box="[639,833,1553,1579]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="thaumatocerus">
|
||
<emphasis id="B908BC1CFFF8FFCEBB78F9C1E238C31A" box="[639,833,1553,1579]" italics="true" pageId="5" pageNumber="410">S. thaumatocerus</emphasis>
|
||
</taxonomicName>
|
||
by the male sexual characters and coarser punctation of the pronotum and elytra. It differs from
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBB7DF9E5E26FC37E" authorityName="Puthz" authorityYear="1998" box="[634,790,1589,1615]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="inflaticollis">
|
||
<emphasis id="B908BC1CFFF8FFCEBB7DF9E5E26FC37E" box="[634,790,1589,1615]" italics="true" pageId="5" pageNumber="410">S. inflaticollis</emphasis>
|
||
</taxonomicName>
|
||
by the punctation of the whole body and the male sexual characters and from
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEB885F989E09FC342" authorityName="Puthz" authorityYear="2013" box="[386,486,1625,1651]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="species" species="quatei">
|
||
<emphasis id="B908BC1CFFF8FFCEB885F989E09FC342" box="[386,486,1625,1651]" italics="true" pageId="5" pageNumber="410">S. quatei</emphasis>
|
||
</taxonomicName>
|
||
by denser punctation of the frons and the pronotum, as well as by the male sexual characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3663385FFF8FFCEB9C0F971E5FFC38A" box="[199,1158,1697,1723]" pageId="5" pageNumber="410" type="etymology">
|
||
<paragraph id="8BC3600EFFF8FFCEB9C0F971E5FFC38A" blockId="5.[151,1437,1013,1723]" box="[199,1158,1697,1723]" pageId="5" pageNumber="410">
|
||
<emphasis id="B908BC1CFFF8FFCEB9C0F971E037C38A" bold="true" box="[199,334,1697,1723]" pageId="5" pageNumber="410">Etymology.</emphasis>
|
||
The specific epithet refers to the fact that species of
|
||
<taxonomicName id="4C7C1B8DFFF8FFCEBA9AF971E29FC38A" authorityName="Latreille" authorityYear="1797" box="[925,998,1697,1723]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="410" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B908BC1CFFF8FFCEBA9AF971E29FC38A" box="[925,998,1697,1723]" italics="true" pageId="5" pageNumber="410">Stenus</emphasis>
|
||
</taxonomicName>
|
||
are predators.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |