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<document id="D51CEDAADB53865E15A208CEDBD020D0" ID-DOI="10.1080/00222930400008736" ID-ISSN="1464-5262" ID-Zenodo-Dep="5214684" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629257395829" checkinUser="felipe" docAuthor="Chen, I-Shiung &amp; Kottelat, Maurice" docDate="2005" docId="03C7915F37156D1282BB02DCFCF07388" docLanguage="en" docName="JNatHist.39.17.1407-1429.pdf" docOrigin="Journal of Natural History 39 (17)" docSource="http://dx.doi.org/10.1080/00222930400008736" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Rhinogobius sulcatus Chen &amp; Kottelat 2005, new species" docType="treatment" docVersion="4" lastPageNumber="1417" masterDocId="FFFEE927371D6D1E837A0657FFFB7740" masterDocTitle="Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam" masterLastPageNumber="1429" masterPageNumber="1407" pageNumber="1413" updateTime="1699065453420" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B4A636F97F1827F240CC3390703D0243">Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="2B01139D5260A7A41F2902FD9888EB3D" type="personal">
<mods:role id="B74833865F98A0DB04BE53A664ED1BBA">
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<mods:namePart id="6306906404C1593E1A5EDD95D6861E1D">Chen, I-Shiung</mods:namePart>
</mods:name>
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<mods:namePart id="AC25B7AD884806D6FF6CFB5B0D000539">Kottelat, Maurice</mods:namePart>
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<mods:typeOfResource id="46F2135BE7F7A588A2EBA1F06C26A6AD">text</mods:typeOfResource>
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<mods:title id="31125F585BAFCDF0EE131AB1304B76CB">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="AABD4C68EF85130FD0ED2B0E9DBDFE91">
<mods:date id="27F520FAA5F41FE216B1F9F165C8EBB6">2005</mods:date>
<mods:detail id="C1560C44784AD0E8978FF7D1E4C3E3B2" type="pubDate">
<mods:number id="7B24829A7AD2AED8B2E7EE051F25EEB4">2005-09-30</mods:number>
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<mods:detail id="30D997A01C3C2C28D04536DC06F280C1" type="volume">
<mods:number id="9058D448D7A2924E1CC019963C621ED1">39</mods:number>
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<mods:detail id="4F37A15D2FCE5A3D8E10ED23B010428F" type="issue">
<mods:number id="CC05F5A24387CC076F9D5EC55A1D03B7">17</mods:number>
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<mods:start id="E111D259A04F592A3AF9E995181940AF">1407</mods:start>
<mods:end id="B516FBB787B87B41FA9782E1BAFB3A98">1429</mods:end>
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<mods:url id="D09026506DB0A72F519FE761DFB65107">http://dx.doi.org/10.1080/00222930400008736</mods:url>
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<mods:classification id="F5CB85FACEE2CAA2F5ABC60553A9CE31">journal article</mods:classification>
<mods:identifier id="82520311EEE7A58D74FF05DCBDD882EF" type="DOI">10.1080/00222930400008736</mods:identifier>
<mods:identifier id="1DC2391696668166F40EAE996129A299" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="70DC5A868BA014C6E7F64A7E8C4FF8A1" type="Zenodo-Dep">5214684</mods:identifier>
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<treatment id="03C7915F37156D1282BB02DCFCF07388" LSID="urn:lsid:zoobank.org:act:F6339828-210C-4154-8F22-424270C7176A" httpUri="http://treatment.plazi.org/id/03C7915F37156D1282BB02DCFCF07388" lastPageId="12" lastPageNumber="1417" pageId="8" pageNumber="1413">
<subSubSection id="C37473C237156D1682BB02DCFC9073E3" box="[449,875,1163,1187]" pageId="8" pageNumber="1413" type="nomenclature">
<paragraph id="8BD1204937156D1682BB02DCFC9073E3" blockId="8.[138,875,1163,1258]" box="[449,875,1163,1187]" pageId="8" pageNumber="1413">
<heading id="D099972537156D1682BB02DCFC9073E3" box="[449,875,1163,1187]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="1413" reason="2">
<taxonomicName id="4C6E5BCA37156D1682BB02DCFD2273E3" ID-CoL="4S932" authority="Chen &amp; Kottelat, 2005" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[449,729,1163,1187]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="1413" phylum="Chordata" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B91AFC5B37156D1682BB02DCFD2273E3" bold="true" box="[449,729,1163,1187]" italics="true" pageId="8" pageNumber="1413">Rhinogobius sulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229412037156D16819802DCFC9073E3" box="[738,875,1163,1187]" pageId="8" pageNumber="1413" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37473C237156D12814A02E6FF127495" lastPageId="12" lastPageNumber="1417" pageId="8" pageNumber="1413" type="description">
<paragraph id="8BD1204937156D16814A02E6FCE873AA" blockId="8.[138,875,1163,1258]" pageId="8" pageNumber="1413">
(
<figureCitation id="13553CCC37156D16814202E6FD5E7389" box="[568,677,1201,1225]" captionStart="Figure 3" captionStartId="9.[125,185,1637,1656]" captionTargetBox="[241,1059,172,1596]" captionTargetId="figure-0@9.[232,1065,165,1601]" captionTargetPageId="9" captionText="Figure 3. Head lateral-line system of Rhinogobius sulcatus, paratype, CMK 14981, 25.4 mm SL. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214690" httpUri="https://zenodo.org/record/5214690/files/figure.png" pageId="8" pageNumber="1413">Figures 3</figureCitation>
,
<figureCitation id="13553CCC37156D1681CE02E6FD3E7389" box="[692,709,1201,1225]" captionStart="Figure 7" captionStartId="16.[138,198,1637,1656]" captionTargetBox="[187,1136,443,1594]" captionTargetId="figure-57@16.[182,1140,441,1601]" captionTargetPageId="16" captionText="Figure 7. Rhinogobius sulcatus, holotype, ZRC 49208, 27.0 mm SL, cephalic colour pattern of male. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214698" httpUri="https://zenodo.org/record/5214698/files/figure.png" pageId="8" pageNumber="1413">7</figureCitation>
,
<figureCitation id="13553CCC37156D1681A902E6FD0A7389" box="[723,753,1201,1225]" captionStart="Figure 11" captionStartId="19.[125,185,1348,1367]" captionTargetBox="[171,1128,819,1312]" captionTargetId="figure-93@19.[171,1128,819,1312]" captionTargetPageId="19" captionText="Figure 11. Rhinogobius sulcatus. (A) Male, holotype, ZRC 49208, 27.0 mm SL; (B) female, paratype, ZRC 49209, 26.8 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5214707" httpUri="https://zenodo.org/record/5214707/files/figure.png" pageId="8" pageNumber="1413">11</figureCitation>
)
<taxonomicName id="4C6E5BCA37156D1683F00285FD7A73AA" authority=": Kottelat 2001 a, p 61" authorityName="Kottelat" authorityPageNumber="61" authorityYear="2001" box="[138,641,1234,1258]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="1413" phylum="Chordata" rank="species" species="duospilus">
<emphasis id="B91AFC5B37156D1683F00285FE8273AA" box="[138,377,1234,1258]" italics="true" pageId="8" pageNumber="1413">Rhinogobius duospilus</emphasis>
:
<bibRefCitation id="EFFF5DB837156D1682F00285FDC773AA" author="Kottelat M" box="[394,572,1234,1258]" pageId="8" pageNumber="1413" refId="ref8906" refString="Kottelat M. 2001 a. Freshwater fishes of northern Vietnam Washington: World Bank. 140 p." type="book" year="2001">Kottelat 2001a</bibRefCitation>
, p 61
</taxonomicName>
, Figure 142.
</paragraph>
<paragraph id="8BD1204937156D1683F00370FEA3727F" blockId="8.[138,344,1319,1343]" box="[138,344,1319,1343]" pageId="8" pageNumber="1413">
<emphasis id="B91AFC5B37156D1683F00370FEA3727F" box="[138,344,1319,1343]" italics="true" pageId="8" pageNumber="1413">Material examined</emphasis>
</paragraph>
<paragraph id="8BD1204937156D1683F00300FF0D72AF" blockId="8.[138,1185,1367,1519]" pageId="8" pageNumber="1413">
<materialsCitation id="3B062A1437156D1683F00300FC3D72EF" accessionNumber="ZRC 49208" collectingDate="1998-10-05" collectorName="M. Kottelat" country="Vietnam" location="Cam Pha District" pageId="8" pageNumber="1413" specimenCount="1" stateProvince="Quang Ninh" typeStatus="holotype">
<typeStatus id="54D59EEB37156D1683F00300FF01722F" box="[138,250,1367,1391]" pageId="8" pageNumber="1413" type="holotype">Holotype</typeStatus>
: ZRC 49208, 27.0 mm SL,
<collectingCountry id="F37960D937156D1681030300FD1B722F" box="[633,736,1367,1391]" name="Vietnam" pageId="8" pageNumber="1413">Vietnam</collectingCountry>
:
<collectingRegion id="49AAEEAB37156D16818D0300FBEA722F" box="[759,1041,1367,1391]" country="Vietnam" name="Quang Ninh" pageId="8" pageNumber="1413">Quang Ninh Province</collectingRegion>
,
<location id="8EB1769237156D1687520300FF1E72CF" LSID="urn:lsid:plazi:treatment:03C7915F37156D1282BB02DCFCF07388:8EB1769237156D1687520300FF1E72CF" country="Vietnam" name="Cam Pha District" pageId="8" pageNumber="1413" stateProvince="Quang Ninh">Cam Pha District</location>
, small coastal stream, at about km 10 on road from Mong Duong to Tien Yen, 21
<emphasis id="B91AFC5B37156D1683D203C0FF4972EF" bold="true" box="[168,178,1431,1455]" italics="true" pageId="8" pageNumber="1413">°</emphasis>
059380N, 107
<emphasis id="B91AFC5B37156D16822C03C0FE9B72EF" bold="true" box="[342,352,1431,1455]" italics="true" pageId="8" pageNumber="1413">°</emphasis>
219020E, coll.
<collectorName id="269B459F37156D16816B03C0FD5B72EF" box="[529,672,1431,1455]" pageId="8" pageNumber="1413">M. Kottelat</collectorName>
et al.,
<date id="FFD0068937156D16807A03C0FC3D72EF" box="[768,966,1431,1455]" pageId="8" pageNumber="1413" value="1998-10-05">
<collectingDate id="EF94FF6137156D16807A03C0FC3D72EF" box="[768,966,1431,1455]" pageId="8" pageNumber="1413" value="1998-10-05">5 October 1998</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B062A1437156D1680AD03C0FE64728F" accessionNumber="CMK 14981" country="Vietnam" pageId="8" pageNumber="1413" specimenCount="1" stateProvince="Quang Ninh" typeStatus="paratype">
<typeStatus id="54D59EEB37156D1680AD03C0FBB072EF" box="[983,1099,1431,1455]" pageId="8" pageNumber="1413" type="paratype">Paratypes</typeStatus>
: CMK 14981,
<specimenCount id="9D68EBC037156D16839E03E0FE64728F" box="[228,415,1463,1487]" pageId="8" pageNumber="1413" type="generic">eight specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B062A1437156D1682D103E0FD02728F" accessionNumber="ZRC 49209" box="[427,761,1463,1487]" country="Vietnam" pageId="8" pageNumber="1413" specimenCount="1" stateProvince="Quang Ninh" typeStatus="paratype">
ZRC 49209,
<specimenCount id="9D68EBC037156D16813D03E0FD02728F" box="[583,761,1463,1487]" pageId="8" pageNumber="1413" type="generic">four specimens</specimenCount>
</materialsCitation>
,
<quantity id="4C968DAC37156D16807C03E0FC56728F" box="[774,941,1463,1487]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" metricValueMax="2.76" metricValueMin="2.06" pageId="8" pageNumber="1413" unit="mm" value="24.1" valueMax="27.6" valueMin="20.6">20.6–27.6 mm</quantity>
SL, other data same as above.
</paragraph>
<paragraph id="8BD1204937156D1683F0007AFF087105" blockId="8.[138,243,1581,1605]" box="[138,243,1581,1605]" pageId="8" pageNumber="1413">
<emphasis id="B91AFC5B37156D1683F0007AFF087105" box="[138,243,1581,1605]" italics="true" pageId="8" pageNumber="1413">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD1204937156D1483F0000AFCD97623" blockId="8.[138,1185,1629,1685]" lastBlockId="10.[138,1185,137,355]" lastPageId="10" lastPageNumber="1415" pageId="8" pageNumber="1413">
<taxonomicName id="4C6E5BCA37156D1683F0000AFE977135" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[138,364,1629,1653]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="1413" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37156D1683F0000AFE977135" box="[138,364,1629,1653]" italics="true" pageId="8" pageNumber="1413">Rhinogobius sulcatus</emphasis>
</taxonomicName>
is distinguished from all congeners by the unique combination of the following features: cheek and opercle with three (rarely four) oblique brownish black bands and one to two stripes, respectively; branchiostegal membrane greyish with 14–17 round light spots (red or orange in life) in male; first dorsal fin with a median brownish black blotch in front of third spinous ray in male; pectoral fin whitish, base with two conspicuous black spots in adults; caudal fin base with a median black spot; body with six to seven conspicuous deep brown blotches on lateral trunk; second dorsal fin rays modally eight; anal fin rays modally seven; pectoral fin rays modally 15; longitudinal scale rows 26–29; predorsal median series 8–11; and vertebrae modally 27.
</paragraph>
<caption id="DF1170C137146D1783070032FB6E7138" ID-DOI="http://doi.org/10.5281/zenodo.5214690" ID-Zenodo-Dep="5214690" box="[125,1173,1637,1656]" httpUri="https://zenodo.org/record/5214690/files/figure.png" pageId="9" pageNumber="1414" startId="9.[125,185,1637,1656]" targetBox="[241,1059,172,1596]" targetPageId="9">
<paragraph id="8BD1204937146D1783070032FB6E7138" blockId="9.[125,1173,1637,1656]" box="[125,1173,1637,1656]" pageId="9" pageNumber="1414">
Figure 3. Head lateral-line system of
<taxonomicName id="4C6E5BCA37146D1782A00032FD767138" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[474,653,1637,1656]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="1414" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37146D1782A00032FD767138" box="[474,653,1637,1656]" italics="true" pageId="9" pageNumber="1414">Rhinogobius sulcatus</emphasis>
</taxonomicName>
, paratype, CMK 14981, 25.4 mm SL. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BD1204937176D1483F007CBFEF876F4" blockId="10.[138,259,412,436]" box="[138,259,412,436]" pageId="10" pageNumber="1415">
<emphasis id="B91AFC5B37176D1483F007CBFEF876F4" box="[138,259,412,436]" italics="true" pageId="10" pageNumber="1415">Description</emphasis>
</paragraph>
<paragraph id="8BD1204937176D1483F0079AFF4275E7" blockId="10.[138,1185,461,679]" pageId="10" pageNumber="1415">Body cylindrical anteriorly, compressed posteriorly. Body proportions in Table I. Head moderately large, slightly depressed anteriorly. Eye large, lips thick. Mouth oblique, rear edge reaching beyond vertical through anterior margin of pupil in both sexes. Both jaws with three to four rows of conical teeth, outer rows enlarged. Tongue margin rounded. Anterior nostril a short tube, posterior one a round hole. Gill opening extending to vertical through margin of preopercle. Isthmus broad. 10211+16217527228 vertebrae (mode 27).</paragraph>
<paragraph id="8BD1204937176D1483F00487FD2174CA" blockId="10.[138,1185,720,906]" pageId="10" pageNumber="1415">
<emphasis id="B91AFC5B37176D1483F00487FF3975A8" box="[138,194,720,744]" italics="true" pageId="10" pageNumber="1415">Fins.</emphasis>
D1 V–VI (mode VI); D2 I/7–9 (mode 8); A I/6–7 (mode 7); P 15–17 (mode 15); V I/ 5+I/5 (frequency distribution in Table II). D1 rays about equal, III, IV longest, rear tip extending to base of first branched ray of D
<quantity id="4C968DAC37176D1481020546FD587469" box="[632,675,785,809]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="1415" unit="in" value="2.0">2 in</quantity>
male, not reaching origin of D
<quantity id="4C968DAC37176D1487780546FBD67469" box="[1026,1069,785,809]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="1415" unit="in" value="2.0">2 in</quantity>
female. A origin below second branched ray of D2. P large, rear margin extending to vertical of anus in male, not reaching this point in female. V disc rounded, spinous rays with pointed membranous lobe. C elliptical, rear edge rounded.
</paragraph>
<paragraph id="8BD1204937176D1483F005E4FD0973CD" blockId="10.[138,1185,947,1165]" pageId="10" pageNumber="1415">
<emphasis id="B91AFC5B37176D1483F005E4FF2E748B" box="[138,213,947,971]" italics="true" pageId="10" pageNumber="1415">Scales.</emphasis>
Body with large ctenoid scales, anterior part of predorsal area naked; scales on posterior predorsal region and belly cycloid; scales in longitudinal series 26–29 (mode 28); transverse series nine (mode nine); predorsal median series 8–11 (mode 10); series between first dorsal and upper pectoral fin origin six (frequency distribution in Table III). Head including opercle, preopercle and prepelvic areas naked. Predorsal squamation with slightly trifurcate anterior edge, anterior extension of median series reaching midline vertical of gap between anterior and posterior oculoscapular canals.
</paragraph>
<paragraph id="8BD1204937176D1483F002E1FF217230" blockId="10.[138,1185,1206,1490]" pageId="10" pageNumber="1415">
<emphasis id="B91AFC5B37176D1483F002E1FE5D738E" box="[138,422,1206,1230]" italics="true" pageId="10" pageNumber="1415">Head lateral-line system.</emphasis>
Canals: nasal extension of anterior oculoscapular canal with terminal pores
<emphasis id="B91AFC5B37176D1482420280FEBC73AF" box="[312,327,1239,1263]" italics="true" pageId="10" pageNumber="1415">S</emphasis>
in front of posterior nostrils. Anterior interorbital section of oculoscapular canal separated, with paired pores
<emphasis id="B91AFC5B37176D14816202A0FDDE724F" box="[536,549,1271,1295]" italics="true" pageId="10" pageNumber="1415">l</emphasis>
. A single pore
<emphasis id="B91AFC5B37176D1481AB02A0FD1B724F" box="[721,736,1271,1295]" italics="true" pageId="10" pageNumber="1415">K</emphasis>
in posterior interorbital region. Pore
<emphasis id="B91AFC5B37176D1487F402A0FB5A724F" box="[1166,1185,1271,1295]" italics="true" pageId="10" pageNumber="1415">v</emphasis>
present at posterior edge of eye. Gap between anterior and posterior oculoscapular canals about equal to length of posterior oculoscapular canal. Preopercular canal with three pores
<emphasis id="B91AFC5B37176D1483F0030FFF6D7230" box="[138,150,1368,1392]" italics="true" pageId="10" pageNumber="1415">c</emphasis>
,
<emphasis id="B91AFC5B37176D1483DC030FFF487230" box="[166,179,1368,1392]" italics="true" pageId="10" pageNumber="1415">d</emphasis>
, E.
</paragraph>
<paragraph id="8BD1204937176D1483DF032EFE187292" blockId="10.[138,1185,1206,1490]" pageId="10" pageNumber="1415">
Sensory papillae: row
<emphasis id="B91AFC5B37176D1482D0032DFE4372D1" box="[426,440,1402,1425]" italics="true" pageId="10" pageNumber="1415">a</emphasis>
reaching midline of orbit. Length of row
<emphasis id="B91AFC5B37176D1480D0032EFC4D72D1" box="[938,950,1401,1425]" italics="true" pageId="10" pageNumber="1415">b</emphasis>
about half of orbit. Row
<emphasis id="B91AFC5B37176D1483BD03CDFF2A72F1" box="[199,209,1434,1457]" italics="true" pageId="10" pageNumber="1415">c</emphasis>
,
<emphasis id="B91AFC5B37176D14839B03CDFF1572F2" box="[225,238,1434,1458]" italics="true" pageId="10" pageNumber="1415">d</emphasis>
long, not reaching vertical of pore
<emphasis id="B91AFC5B37176D1481F203CDFD6E72F2" box="[648,661,1434,1458]" italics="true" pageId="10" pageNumber="1415">a</emphasis>
. A single
<emphasis id="B91AFC5B37176D14807103CDFCD972F1" box="[779,802,1434,1457]" italics="true" pageId="10" pageNumber="1415">cp</emphasis>
papilla. Row
<emphasis id="B91AFC5B37176D1480BE03CDFC3072F2" box="[964,971,1434,1458]" italics="true" pageId="10" pageNumber="1415">f</emphasis>
paired. Opercular rows
<emphasis id="B91AFC5B37176D1483B303ECFF267292" box="[201,221,1467,1490]" italics="true" pageId="10" pageNumber="1415">ot</emphasis>
and
<emphasis id="B91AFC5B37176D14826003EDFED67292" box="[282,301,1466,1490]" italics="true" pageId="10" pageNumber="1415">oi</emphasis>
well separated.
</paragraph>
<paragraph id="8BD1204937176D1283F003ACFF2777A1" blockId="10.[138,1185,1531,1684]" lastBlockId="12.[138,1185,137,452]" lastPageId="12" lastPageNumber="1417" pageId="10" pageNumber="1415">
<emphasis id="B91AFC5B37176D1483F003ACFE937153" box="[138,360,1531,1555]" italics="true" pageId="10" pageNumber="1415">Coloration in alcohol.</emphasis>
Head and body light yellowish brown. Body with six to seven conspicuous deep brown blotches on flank and five deep brown blotches in dorsal region. Marking of scale pocket indistinct. Dorsal side of snout with a pair of brown lines united at tip of snout. A longitudinal brown line behind orbit. Nape with some irregular brown marks. Cheek and opercle always with three (rarely four) oblique brownish black stripes and one to two stripes, respectively. A brown blotch below eye in female. Branchiostegal membrane greyish with 14–17 round light spots (red or orange in life) in male, uniformly light yellowish in female.
</paragraph>
<caption id="DF1170C137166D158307002CFB6E71CE" ID-DOI="http://doi.org/10.5281/zenodo.5214692" ID-Zenodo-Dep="5214692" box="[125,1173,1659,1678]" httpUri="https://zenodo.org/record/5214692/files/figure.png" pageId="11" pageNumber="1416" startId="11.[125,185,1659,1678]" targetBox="[208,1091,192,1622]" targetPageId="11">
<paragraph id="8BD1204937166D158307002CFB6E71CE" blockId="11.[125,1173,1659,1678]" box="[125,1173,1659,1678]" pageId="11" pageNumber="1416">
Figure 4. Head lateral-line system of
<taxonomicName id="4C6E5BCA37166D1582AE002CFD6C71CE" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[468,663,1659,1678]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="1416" phylum="Chordata" rank="species" species="variolatus">
<emphasis id="B91AFC5B37166D1582AE002CFD6C71CE" box="[468,663,1659,1678]" italics="true" pageId="11" pageNumber="1416">Rhinogobius variolatus</emphasis>
</taxonomicName>
, paratype, CMK 14982, 25.9 mm SL. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BD1204937116D1283DF06BDFC177684" blockId="12.[138,1185,137,452]" pageId="12" pageNumber="1417">First dorsal fin with light margin and a median brownish black blotch in front of third spinous ray in male, none in female. Second dorsal fin grey with two basal rows of blackish brown spots in male, whitish with three rows of longitudinal brown spots in female. Anal fin greyish, with light margin. Pectoral fin whitish, base with two conspicuous black spots in adults of both sexes. Caudal fin greyish in male, pale white in female, with three to six rows of vertical greyish brown spots and base with a median black spot. Pelvic fin greyish in male, pale white in female.</paragraph>
<paragraph id="8BD1204937116D1283F0045CFEF07563" blockId="12.[138,267,523,547]" box="[138,267,523,547]" pageId="12" pageNumber="1417">
<emphasis id="B91AFC5B37116D1283F0045CFEF07563" box="[138,267,523,547]" italics="true" pageId="12" pageNumber="1417">Distribution</emphasis>
</paragraph>
<paragraph id="8BD1204937116D1283F0046CFE5B75B5" blockId="12.[138,1185,571,757]" pageId="12" pageNumber="1417">
<taxonomicName id="4C6E5BCA37116D1283F0046CFE987513" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[138,355,571,595]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D1283F0046CFE987513" box="[138,355,571,595]" italics="true" pageId="12" pageNumber="1417">Rhinogobius sulcatus</emphasis>
</taxonomicName>
<materialsCitation id="3B062A1437116D128210046CFCD87534" country="Vietnam" location="Cam Pha" pageId="12" pageNumber="1417" specimenCount="1">
is definitively known only from the
<typeStatus id="54D59EEB37116D12807D046CFCC37513" box="[775,824,571,595]" pageId="12" pageNumber="1417">type</typeStatus>
locality, a small coastal stream north of
<location id="8EB1769237116D128394040BFEA57534" LSID="urn:lsid:plazi:treatment:03C7915F37156D1282BB02DCFCF07388:8EB1769237116D128394040BFEA57534" box="[238,350,604,628]" country="Vietnam" name="Cam Pha" pageId="12" pageNumber="1417">Cam Pha</location>
, on the north-eastern coast of
<collectingCountry id="F37960D937116D1281C6040BFCD87534" box="[700,803,604,628]" name="Vietnam" pageId="12" pageNumber="1417">Vietnam</collectingCountry>
</materialsCitation>
. It was collected together with
<taxonomicName id="4C6E5BCA37116D1287FC040BFF0E75D4" authorityName="Chen &amp; Kottelat" authorityYear="2005" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="variolatus">
<emphasis id="B91AFC5B37116D1287FC040BFF0E75D4" italics="true" pageId="12" pageNumber="1417">R. variolatus</emphasis>
</taxonomicName>
. The specimen figured in
<bibRefCitation id="EFFF5DB837116D128148042BFD1475D4" author="Kottelat M" box="[562,751,636,660]" pageId="12" pageNumber="1417" refId="ref8906" refString="Kottelat M. 2001 a. Freshwater fishes of northern Vietnam Washington: World Bank. 140 p." type="book" year="2001">Kottelat (2001a</bibRefCitation>
, Figure 142) probably represents the same species, but this cannot be confirmed as the material is no longer available. This material was from a small hill stream in the Song Lo drainage (a tributary of Song Hong), in north-western
<collectingCountry id="F37960D937116D12824F048AFE6775B5" box="[309,412,733,757]" name="Vietnam" pageId="12" pageNumber="1417">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BD1204937116D1283F00579FF007406" blockId="12.[138,251,814,838]" box="[138,251,814,838]" pageId="12" pageNumber="1417">
<emphasis id="B91AFC5B37116D1283F00579FF007406" box="[138,251,814,838]" italics="true" pageId="12" pageNumber="1417">Etymology</emphasis>
</paragraph>
<paragraph id="8BD1204937116D1283F00508FBB67437" blockId="12.[138,1101,863,887]" box="[138,1101,863,887]" pageId="12" pageNumber="1417">
From the latin
<taxonomicName id="4C6E5BCA37116D1282470508FE6A7437" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[317,401,863,887]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D1282470508FE6A7437" box="[317,401,863,887]" italics="true" pageId="12" pageNumber="1417">sulcatus</emphasis>
</taxonomicName>
(scarred). An adjective (feminine:
<emphasis id="B91AFC5B37116D1280560508FC8C7437" box="[812,887,863,887]" italics="true" pageId="12" pageNumber="1417">sulcata</emphasis>
; neuter
<emphasis id="B91AFC5B37116D1280A10508FBC07437" box="[987,1083,863,887]" italics="true" pageId="12" pageNumber="1417">sulcatum</emphasis>
).
</paragraph>
<paragraph id="8BD1204937116D1283F005EAFF127495" blockId="12.[138,233,957,981]" box="[138,233,957,981]" pageId="12" pageNumber="1417">
<emphasis id="B91AFC5B37116D1283F005EAFF127495" box="[138,233,957,981]" italics="true" pageId="12" pageNumber="1417">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37473C237116D1283F005B9FCF07388" pageId="12" pageNumber="1417" type="reference_group">
<paragraph id="8BD1204937116D1283F005B9FCF07388" blockId="12.[138,1185,1006,1224]" pageId="12" pageNumber="1417">
<taxonomicName id="4C6E5BCA37116D1283F005B9FE967346" ID-CoL="4S932" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[138,365,1006,1030]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D1283F005B9FE967346" box="[138,365,1006,1030]" italics="true" pageId="12" pageNumber="1417">Rhinogobius sulcatus</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C6E5BCA37116D12812905B8FC6D7346" ID-CoL="4S8YZ" authority="(Herre, 1935 a)" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[595,918,1006,1030]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="duospilus">
<emphasis id="B91AFC5B37116D12812905B8FD207346" box="[595,731,1006,1030]" italics="true" pageId="12" pageNumber="1417">R. duospilus</emphasis>
(
<bibRefCitation id="EFFF5DB837116D12819705B9FC757346" author="Herre AWCT" box="[749,910,1006,1030]" pageId="12" pageNumber="1417" pagination="285 - 293" refId="ref8783" refString="Herre AWCT. 1935 a. Note on the fishes in the Zoological Museum of Standford University, new and rare Hongkong fishes obtained in 1934. Hong Kong Nature 6: 285 - 293." type="journal article" year="1935">Herre, 1935a</bibRefCitation>
)
</taxonomicName>
(a species endemic to
<collectingCountry id="F37960D937116D1283F00259FEE47366" box="[138,287,1038,1062]" name="Hong Kong" pageId="12" pageNumber="1417">Hong Kong</collectingCountry>
) than to any other species in southern
<collectingCountry id="F37960D937116D1280500259FC8E7366" box="[810,885,1038,1062]" name="China" pageId="12" pageNumber="1417">China</collectingCountry>
. However, they can be distinguished by the following combinations of features: (1) body scales: body with fewer lateral scales in
<taxonomicName id="4C6E5BCA37116D1282470207FE487327" ID-CoL="4S932" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[317,435,1103,1127]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D1282470207FE487327" box="[317,435,1103,1127]" italics="true" pageId="12" pageNumber="1417">R. sulcatus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C6E5BCA37116D12816B0207FD6E7327" ID-CoL="4S8YZ" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[529,661,1103,1127]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="duospilus">
<emphasis id="B91AFC5B37116D12816B0207FD6E7327" box="[529,661,1103,1127]" italics="true" pageId="12" pageNumber="1417">R. duospilus</emphasis>
</taxonomicName>
(always 27–28 versus 30–31); (2) pectoral fin rays: fewer pectoral fin rays in
<taxonomicName id="4C6E5BCA37116D12829F0227FDA073C7" ID-CoL="4S932" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[485,603,1135,1159]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D12829F0227FDA073C7" box="[485,603,1135,1159]" italics="true" pageId="12" pageNumber="1417">R. sulcatus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C6E5BCA37116D1281C20227FCC773C7" ID-CoL="4S8YZ" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[696,828,1135,1159]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="duospilus">
<emphasis id="B91AFC5B37116D1281C20227FCC773C7" box="[696,828,1135,1159]" italics="true" pageId="12" pageNumber="1417">R. duospilus</emphasis>
</taxonomicName>
(usually 15 versus 16–17); and (3) coloration pattern in male: opercle with one to two brownish black stripes in
<taxonomicName id="4C6E5BCA37116D12875102C7FB5A73E8" ID-CoL="4S932" authorityName="Chen &amp; Kottelat" authorityYear="2005" box="[1067,1185,1168,1192]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="B91AFC5B37116D12875102C7FB5A73E8" box="[1067,1185,1168,1192]" italics="true" pageId="12" pageNumber="1417">R. sulcatus</emphasis>
</taxonomicName>
versus none on upper region of opercle in
<taxonomicName id="4C6E5BCA37116D12810702E6FCF87388" ID-CoL="4S8YZ" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[637,771,1200,1224]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="duospilus">
<emphasis id="B91AFC5B37116D12810702E6FCF87388" box="[637,771,1200,1224]" italics="true" pageId="12" pageNumber="1417">R. duospilus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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