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<document id="27D8F2D5C0E2E7B0E4BE37A2DE0C7E46" ID-DOI="10.11646/zootaxa.4040.3.1" ID-GBIF-Dataset="589c20cd-ac9b-4155-8b98-8c0742b5d849" ID-ISSN="1175-5326" ID-Zenodo-Dep="245740" ID-ZooBank="392A2F34-0B0C-4298-BBF5-76A82CED0C59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461335112013" checkinUser="plazi" docAuthor="Ruiz, Gustavo R. S. &amp; Maddison, Wayne P." docDate="2015" docId="03C087CBFF875446FF05FC20FD0DFE3C" docLanguage="en" docName="zt04040p279.pdf" docOrigin="Zootaxa 4040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Urupuyu Ruiz &amp; Maddison, 2015, gen. nov." docType="treatment" docVersion="8" lastPageNumber="254" masterDocId="FFF9FFB3FF855445FF92FF95FF8BFFEC" masterDocTitle="The new Andean jumping spider genus Urupuyu and its placement within a revised classification of the Amycoida (Araneae: Salticidae)" masterLastPageNumber="279" masterPageNumber="251" pageNumber="253" updateTime="1698619168940" updateUser="plazi">
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<mods:title id="F164D55BD0B73153A2B2F901A60A0E70">The new Andean jumping spider genus Urupuyu and its placement within a revised classification of the Amycoida (Araneae: Salticidae)</mods:title>
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<mods:namePart id="D5FF5A379CDEFC84885E63B4A6A3AED3">Ruiz, Gustavo R. S.</mods:namePart>
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<mods:namePart id="3CB003DED486285FAA350574B8CA3BA6">Maddison, Wayne P.</mods:namePart>
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<mods:date id="383886110FED9844D66129FB828B7801">2015</mods:date>
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<treatment id="03C087CBFF875446FF05FC20FD0DFE3C" ID-DOI="http://doi.org/10.5281/zenodo.6121668" ID-GBIF-Taxon="119644476" ID-Zenodo-Dep="6121668" LSID="urn:lsid:plazi:treatment:03C087CBFF875446FF05FC20FD0DFE3C" httpUri="http://treatment.plazi.org/id/03C087CBFF875446FF05FC20FD0DFE3C" lastPageId="3" lastPageNumber="254" pageId="2" pageNumber="253">
<subSubSection id="C3736556FF875447FF05FC20FEE4FC21" box="[151,367,948,973]" pageId="2" pageNumber="253" type="nomenclature">
<paragraph id="8BD636DDFF875447FF05FC20FEE4FC21" blockId="2.[151,367,948,973]" box="[151,367,948,973]" pageId="2" pageNumber="253">
<heading id="D09E81B1FF875447FF05FC20FEE4FC21" bold="true" box="[151,367,948,973]" fontSize="10" level="2" pageId="2" pageNumber="253" reason="2">
<emphasis id="B91DEACFFF875447FF05FC20FEE4FC21" bold="true" box="[151,367,948,973]" pageId="2" pageNumber="253">
<taxonomicName id="4C694D5EFF875447FF05FC20FF74FC20" ID-CoL="63VHS" ID-ENA="1766291" box="[151,255,949,972]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91DEACFFF875447FF05FC20FF74FC20" bold="true" box="[151,255,949,972]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57B4FF875447FE94FC21FEE4FC21" box="[262,367,948,973]" pageId="2" pageNumber="253" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3736556FF875447FF05FC69FDB2FBFE" box="[151,569,1020,1042]" pageId="2" pageNumber="253" type="materials_examined">
<paragraph id="8BD636DDFF875447FF05FC69FDB2FBFE" blockId="2.[151,569,1020,1042]" box="[151,569,1020,1042]" pageId="2" pageNumber="253">
<emphasis id="B91DEACFFF875447FF05FC69FEAEFBFE" bold="true" box="[151,293,1020,1042]" pageId="2" pageNumber="253">
<typeStatus id="54D2887FFF875447FF05FC69FF47FBFE" box="[151,204,1020,1042]" pageId="2" pageNumber="253">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C694D5EFF875447FEB9FC68FE68FBFE" box="[299,483,1021,1042]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="species" species="antisana" status="sp. nov.">
<emphasis id="B91DEACFFF875447FEB9FC68FE68FBFE" box="[299,483,1021,1042]" italics="true" pageId="2" pageNumber="253">Urupuyu antisana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57B4FF875447FE7BFC69FDB2FBFE" box="[489,569,1020,1042]" pageId="2" pageNumber="253" rank="species">
<emphasis id="B91DEACFFF875447FE7BFC69FDB2FBFE" bold="true" box="[489,569,1020,1042]" pageId="2" pageNumber="253">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3736556FF875447FF05FBABFB11FB90" pageId="2" pageNumber="253" type="etymology">
<paragraph id="8BD636DDFF875447FF05FBABFB11FB90" blockId="2.[151,1437,1086,2012]" pageId="2" pageNumber="253">
<emphasis id="B91DEACFFF875447FF05FBABFEAAFBBB" bold="true" box="[151,289,1086,1111]" pageId="2" pageNumber="253">Etymology:</emphasis>
The generic name is a euphonic combination of the Quechuan words “uru” (=spider) and “puyu” (=cloud), in allusion to the Andean cloud forests where these spiders live. Neuter in gender.
</paragraph>
</subSubSection>
<subSubSection id="C3736556FF875447FF55FB13FB6CF9C7" pageId="2" pageNumber="253" type="diagnosis">
<paragraph id="8BD636DDFF875447FF55FB13FB6CF9C7" blockId="2.[151,1437,1086,2012]" pageId="2" pageNumber="253">
<emphasis id="B91DEACFFF875447FF55FB13FEC8FB73" bold="true" box="[199,323,1158,1183]" pageId="2" pageNumber="253">Diagnosis:</emphasis>
<taxonomicName id="4C694D5EFF875447FEDEFB1DFE3BFB73" box="[332,432,1160,1183]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FEDEFB1DFE3BFB73" box="[332,432,1160,1183]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
is a member of the huriines, a group distinguished from other South American amycoid lineages (except bredines) by the combination of chelicerae having a single retromarginal tooth and several promarginal teeth, and male palp with well developed RvTA pointing distally (
<figureCitation id="13522A58FF875447FBBBFB5BFB08FB0B" box="[1065,1155,1230,1255]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 23</figureCitation>
,
<figureCitation id="13522A58FF875447FB03FB5AFB24FB0B" box="[1169,1199,1231,1255]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">28</figureCitation>
,
<figureCitation id="13522A58FF875447FB2CFB5AFB56FB0B" box="[1214,1245,1231,1255]" captionStart="FIGURES 32 35" captionStartId="9.[151,258,1490,1512]" captionTargetBox="[164,1436,788,1470]" captionTargetId="figure@9.[151,1436,788,1470]" captionTargetPageId="9" captionText="FIGURES 32 35. Urupuyu occidentale: 32 male left palp, ventral view; 33 same, palpal tibia, retrolateral view; 34 same, dorsal view; Urupuyu spp: 35 distributional records in Ecuador." httpUri="https://zenodo.org/record/245748/files/figure.png" pageId="2" pageNumber="253">33</figureCitation>
). Huriines differ from bredines in having a standard and more compact salticid body (flattened and elongate in bredines).
<taxonomicName id="4C694D5EFF875447FAAAFB61FA17FAE7" box="[1336,1436,1268,1291]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FAAAFB61FA17FAE7" box="[1336,1436,1268,1291]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
differs from remaining huriine genera in having a complex RTA divided into two lobes, a more dorsal one forming a hook towards the cymbium (
<figureCitation id="13522A58FF875447FE7AFAAEFDB5FABF" box="[488,574,1339,1364]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 24</figureCitation>
,
<figureCitation id="13522A58FF875447FDDBFAAEFDECFABF" box="[585,615,1339,1363]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">29</figureCitation>
,
<figureCitation id="13522A58FF875447FDE0FAAEFD1AFABF" box="[626,657,1339,1363]" captionStart="FIGURES 32 35" captionStartId="9.[151,258,1490,1512]" captionTargetBox="[164,1436,788,1470]" captionTargetId="figure@9.[151,1436,788,1470]" captionTargetPageId="9" captionText="FIGURES 32 35. Urupuyu occidentale: 32 male left palp, ventral view; 33 same, palpal tibia, retrolateral view; 34 same, dorsal view; Urupuyu spp: 35 distributional records in Ecuador." httpUri="https://zenodo.org/record/245748/files/figure.png" pageId="2" pageNumber="253">34</figureCitation>
), and a more ventral one, which can bear an acute projection (
<figureCitation id="13522A58FF875447FAD3FAAEFA1CFABF" box="[1345,1431,1339,1364]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 23</figureCitation>
,
<figureCitation id="13522A58FF875447FF05FACAFF3DFA9B" box="[151,182,1375,1399]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">28</figureCitation>
,
<figureCitation id="13522A58FF875447FF53FACAFF6BFA9B" box="[193,224,1375,1399]" captionStart="FIGURES 32 35" captionStartId="9.[151,258,1490,1512]" captionTargetBox="[164,1436,788,1470]" captionTargetId="figure@9.[151,1436,788,1470]" captionTargetPageId="9" captionText="FIGURES 32 35. Urupuyu occidentale: 32 male left palp, ventral view; 33 same, palpal tibia, retrolateral view; 34 same, dorsal view; Urupuyu spp: 35 distributional records in Ecuador." httpUri="https://zenodo.org/record/245748/files/figure.png" pageId="2" pageNumber="253">33</figureCitation>
); having a membranous projection along the embolus at the middle of its length (
<figureCitation id="13522A58FF875447FBFFFACBFB69FA9B" box="[1133,1250,1374,1399]" captionStart="FIGURES 9 13" captionStartId="5.[151,258,1681,1703]" captionTargetBox="[151,1436,191,1659]" captionTargetId="figure@5.[151,1436,191,1660]" captionTargetPageId="5" captionText="FIGURES 9 13. Urupuyu antisana: 9 male left palp, ventral view (white arrow shows projection on embolus; black arrow in white triangle shows tegular cleft); 10 detail of same; 11 chelicerae, frontal view; 12 right chelicera, promarginal teeth (arrows); 13 left chelicera, promarginal teeth (arrows)." httpUri="https://zenodo.org/record/245743/files/figure.png" pageId="2" pageNumber="253">Figs 910</figureCitation>
); and being dark and more or less glabrous (
<figureCitation id="13522A58FF875447FE55FA16FDA6FA70" box="[455,557,1411,1436]" captionStart="FIGURES 1 5" captionStartId="3.[151,258,1949,1971]" captionTargetBox="[151,1436,502,1927]" captionTargetId="figure@3.[151,1436,502,1928]" captionTargetPageId="3" captionText="FIGURES 1 5. Urupuyu spp. U. antisana: 1 male from Papallacta; 2 female from Papallacta; 3 female from Sendero Jumandy. U. edwardsi: 4 male; 5 same, sitting on fern leaf. Figs 1 - 5 © 2015 W. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/245741/files/figure.png" pageId="2" pageNumber="253">Figs 15</figureCitation>
). Besides that,
<taxonomicName id="4C694D5EFF875447FD4BFA11FCB6FA77" box="[729,829,1412,1435]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FD4BFA11FCB6FA77" box="[729,829,1412,1435]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
seems to be the only huriine genus with a cleft on the tegulum (
<figureCitation id="13522A58FF875447FE91FA33FEC5FA53" box="[259,334,1446,1471]" captionStart="FIGURES 9 13" captionStartId="5.[151,258,1681,1703]" captionTargetBox="[151,1436,191,1659]" captionTargetId="figure@5.[151,1436,191,1660]" captionTargetPageId="5" captionText="FIGURES 9 13. Urupuyu antisana: 9 male left palp, ventral view (white arrow shows projection on embolus; black arrow in white triangle shows tegular cleft); 10 detail of same; 11 chelicerae, frontal view; 12 right chelicera, promarginal teeth (arrows); 13 left chelicera, promarginal teeth (arrows)." httpUri="https://zenodo.org/record/245743/files/figure.png" pageId="2" pageNumber="253">Figs 9</figureCitation>
, arrow within triangle, 22). The epigyne in
<taxonomicName id="4C694D5EFF875447FCD6FA3DFC23FA53" box="[836,936,1448,1471]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FCD6FA3DFC23FA53" box="[836,936,1448,1471]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C694D5EFF875447FB06FA32FB75FA52" box="[1172,1278,1447,1470]" class="Arachnida" family="Salticidae" genus="Scoturius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FB06FA32FB75FA52" box="[1172,1278,1447,1470]" italics="true" pageId="2" pageNumber="253">Scoturius</emphasis>
</taxonomicName>
in having the initial portion of the copulation ducts dilated as a chamber (marked with arrow in
<figureCitation id="13522A58FF875447FBF5FA5EFB4AFA08" box="[1127,1217,1483,1508]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 26</figureCitation>
,
<figureCitation id="13522A58FF875447FB43FA5EFB7BFA0F" box="[1233,1264,1483,1507]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">31</figureCitation>
; the female of
<taxonomicName id="4C694D5EFF875447FF05FA7BFF75F9EB" box="[151,254,1518,1543]" class="Arachnida" family="Salticidae" genus="Atelurius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FF05FA7BFF75F9EB" box="[151,254,1518,1543]" italics="true" pageId="2" pageNumber="253">Atelurius</emphasis>
</taxonomicName>
is still unknown), but differs in having more separate copulation openings and longer copulation ducts (
<figureCitation id="13522A58FF875447FF0DF986FEABF9C0" box="[159,288,1555,1580]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 2526</figureCitation>
,
<figureCitation id="13522A58FF875447FEB9F986FEFDF9C7" box="[299,374,1555,1579]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">3031</figureCitation>
). The molecular data suggest
<taxonomicName id="4C694D5EFF875447FD51F981FCACF9C7" box="[707,807,1556,1579]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FD51F981FCACF9C7" box="[707,807,1556,1579]" italics="true" pageId="2" pageNumber="253">Urupuyu</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694D5EFF875447FCCFF981FC4CF9C7" box="[861,967,1556,1579]" class="Arachnida" family="Salticidae" genus="Scoturius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF875447FCCFF981FC4CF9C7" box="[861,967,1556,1579]" italics="true" pageId="2" pageNumber="253">Scoturius</emphasis>
</taxonomicName>
are distinct (
<figureCitation id="13522A58FF875447FBCAF986FB50F9C7" box="[1112,1243,1555,1580]" captionStart="FIGURES 37 38" captionStartId="16.[151,258,1890,1912]" captionTargetBox="[167,1421,206,1858]" captionTargetId="figure@16.[151,1436,191,1868]" captionTargetPageId="16" captionText="FIGURES 37 38. Maximum likelikhood phylogenies from Chimera and Non-chimera matrices, combined 5 gene analyses. Numbers show likelihood bootstrap % for Amycoids only; in parenthesis are bootstrap percentages for analysis without Nilakantha and Gypogyna (shown for deep amycoid branches only). Rectangles at nodes indicate bootstrap support in parsimony analysis (filled rectangle at least 75 %; open 50 % to 74 %). Sites partitioned as per PartitionFinder analysis." httpUri="https://zenodo.org/record/245750/files/figure.png" pageId="2" pageNumber="253">Figs 37, 38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3736556FF875446FF55F9A3FE35FE41" lastPageId="3" lastPageNumber="254" pageId="2" pageNumber="253" type="description">
<paragraph id="8BD636DDFF875446FF55F9A3FA0DFEAC" blockId="2.[151,1437,1086,2012]" lastBlockId="3.[151,1436,151,464]" lastPageId="3" lastPageNumber="254" pageId="2" pageNumber="253">
<emphasis id="B91DEACFFF875447FF55F9A3FED0F9A3" bold="true" box="[199,347,1590,1615]" pageId="2" pageNumber="253">Description:</emphasis>
Small (2.53.0 mm), dark salticids (
<figureCitation id="13522A58FF875447FCB1F9A3FC06F9A3" box="[803,909,1590,1615]" captionStart="FIGURES 1 5" captionStartId="3.[151,258,1949,1971]" captionTargetBox="[151,1436,502,1927]" captionTargetId="figure@3.[151,1436,502,1928]" captionTargetPageId="3" captionText="FIGURES 1 5. Urupuyu spp. U. antisana: 1 male from Papallacta; 2 female from Papallacta; 3 female from Sendero Jumandy. U. edwardsi: 4 male; 5 same, sitting on fern leaf. Figs 1 - 5 © 2015 W. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/245741/files/figure.png" pageId="2" pageNumber="253">Figs 15</figureCitation>
) from vegetation of high elevation forests. Carapace covered with small depressions, in the middle of which there is a pore (
<figureCitation id="13522A58FF875447FBD8F9CEFB1FF99F" box="[1098,1172,1627,1652]" captionStart="FIGURES 1 5" captionStartId="3.[151,258,1949,1971]" captionTargetBox="[151,1436,502,1927]" captionTargetId="figure@3.[151,1436,502,1928]" captionTargetPageId="3" captionText="FIGURES 1 5. Urupuyu spp. U. antisana: 1 male from Papallacta; 2 female from Papallacta; 3 female from Sendero Jumandy. U. edwardsi: 4 male; 5 same, sitting on fern leaf. Figs 1 - 5 © 2015 W. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/245741/files/figure.png" pageId="2" pageNumber="253">Figs 1</figureCitation>
,
<figureCitation id="13522A58FF875447FB36F9CEFB5AF998" box="[1188,1233,1627,1652]" captionStart="FIGURES 6 8" captionStartId="4.[151,258,1942,1964]" captionTargetBox="[151,1436,644,1920]" captionTargetId="figure@4.[151,1436,644,1922]" captionTargetPageId="4" captionText="FIGURES 6 8. Urupuyu antisana: 6 male carapace; 7 same, showing depressions (larger, arrowed) and punctuations (smaller); 8 same, detail showing perforation in depression." httpUri="https://zenodo.org/record/245742/files/figure.png" pageId="2" pageNumber="253">68</figureCitation>
; see note below). Between these depressions with pores there are minute punctuations. The clypeus is low in both sexes. No scales on carapace. Chelicera small and vertical in both sexes, with 34 promarginal teeth (
<figureCitation id="13522A58FF875447FBE6F936FB7DF957" box="[1140,1270,1699,1724]" captionStart="FIGURES 9 13" captionStartId="5.[151,258,1681,1703]" captionTargetBox="[151,1436,191,1659]" captionTargetId="figure@5.[151,1436,191,1660]" captionTargetPageId="5" captionText="FIGURES 9 13. Urupuyu antisana: 9 male left palp, ventral view (white arrow shows projection on embolus; black arrow in white triangle shows tegular cleft); 10 detail of same; 11 chelicerae, frontal view; 12 right chelicera, promarginal teeth (arrows); 13 left chelicera, promarginal teeth (arrows)." httpUri="https://zenodo.org/record/245743/files/figure.png" pageId="2" pageNumber="253">Figs 1113</figureCitation>
) and a single, simple retromarginal tooth. Male palp: unmodified femur and patella. Tibia with a wide RTA forming a hook dorsally (
<figureCitation id="13522A58FF875447FE92F97EFED2F8EF" box="[256,345,1771,1796]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 24</figureCitation>
,
<figureCitation id="13522A58FF875447FEF7F97EFE08F8EF" box="[357,387,1771,1795]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">29</figureCitation>
,
<figureCitation id="13522A58FF875447FE1DF97EFE27F8EF" box="[399,428,1771,1795]" captionStart="FIGURES 32 35" captionStartId="9.[151,258,1490,1512]" captionTargetBox="[164,1436,788,1470]" captionTargetId="figure@9.[151,1436,788,1470]" captionTargetPageId="9" captionText="FIGURES 32 35. Urupuyu occidentale: 32 male left palp, ventral view; 33 same, palpal tibia, retrolateral view; 34 same, dorsal view; Urupuyu spp: 35 distributional records in Ecuador." httpUri="https://zenodo.org/record/245748/files/figure.png" pageId="2" pageNumber="253">34</figureCitation>
) (also with an acute retrolateral projection in some species) and an acute RvTA (
<figureCitation id="13522A58FF875447FAD2F97EFA1CF8EF" box="[1344,1431,1771,1796]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 23</figureCitation>
,
<figureCitation id="13522A58FF875447FF05F89AFF3DF8CB" box="[151,182,1807,1831]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">28</figureCitation>
,
<figureCitation id="13522A58FF875447FF56F89AFF68F8CB" box="[196,227,1807,1831]" captionStart="FIGURES 32 35" captionStartId="9.[151,258,1490,1512]" captionTargetBox="[164,1436,788,1470]" captionTargetId="figure@9.[151,1436,788,1470]" captionTargetPageId="9" captionText="FIGURES 32 35. Urupuyu occidentale: 32 male left palp, ventral view; 33 same, palpal tibia, retrolateral view; 34 same, dorsal view; Urupuyu spp: 35 distributional records in Ecuador." httpUri="https://zenodo.org/record/245748/files/figure.png" pageId="2" pageNumber="253">33</figureCitation>
). Cymbium drop-shaped. Tegulum roundish, bearing a cleft (
<figureCitation id="13522A58FF875447FC37F89BFC77F8CB" box="[933,1020,1806,1831]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Fig. 22</figureCitation>
). Embolus fixed to the tegulum and curling around it, with a median blunt projection lying in a pit of the prolateral side of tegulum (
<figureCitation id="13522A58FF875447FB4EF8A6FAD9F8A7" box="[1244,1362,1843,1868]" captionStart="FIGURES 9 13" captionStartId="5.[151,258,1681,1703]" captionTargetBox="[151,1436,191,1659]" captionTargetId="figure@5.[151,1436,191,1660]" captionTargetPageId="5" captionText="FIGURES 9 13. Urupuyu antisana: 9 male left palp, ventral view (white arrow shows projection on embolus; black arrow in white triangle shows tegular cleft); 10 detail of same; 11 chelicerae, frontal view; 12 right chelicera, promarginal teeth (arrows); 13 left chelicera, promarginal teeth (arrows)." httpUri="https://zenodo.org/record/245743/files/figure.png" pageId="2" pageNumber="253">Figs 910</figureCitation>
). Legs can be black or yellow (
<figureCitation id="13522A58FF875447FE21F8C3FD97F883" box="[435,540,1878,1903]" captionStart="FIGURES 1 5" captionStartId="3.[151,258,1949,1971]" captionTargetBox="[151,1436,502,1927]" captionTargetId="figure@3.[151,1436,502,1928]" captionTargetPageId="3" captionText="FIGURES 1 5. Urupuyu spp. U. antisana: 1 male from Papallacta; 2 female from Papallacta; 3 female from Sendero Jumandy. U. edwardsi: 4 male; 5 same, sitting on fern leaf. Figs 1 - 5 © 2015 W. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/245741/files/figure.png" pageId="2" pageNumber="253">Figs 15</figureCitation>
). Leg formula: males 1423, females 4123. Male second tarsus with modified scale-like setae (
<figureCitation id="13522A58FF875447FEC8F8EEFE69F878" box="[346,482,1915,1940]" captionStart="FIGURES 14 17" captionStartId="6.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@6.[151,1436,191,1155]" captionTargetPageId="6" captionText="FIGURES 14 17. Urupuyu antisana: 14 epiandric region (arrows show fusules); 15 male left tarsus II; 16 detail of same, showing modified white setae; 17 same." httpUri="https://zenodo.org/record/245744/files/figure.png" pageId="2" pageNumber="253">Figs 1517</figureCitation>
). Tarsal claws small. Claw tufts poorly developed. Males with dorsal abdominal scutum and epiandrous fusules (
<figureCitation id="13522A58FF875447FD91F80BFDDCF85B" box="[515,599,1950,1975]" captionStart="FIGURES 14 17" captionStartId="6.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@6.[151,1436,191,1155]" captionTargetPageId="6" captionText="FIGURES 14 17. Urupuyu antisana: 14 epiandric region (arrows show fusules); 15 male left tarsus II; 16 detail of same, showing modified white setae; 17 same." httpUri="https://zenodo.org/record/245744/files/figure.png" pageId="2" pageNumber="253">Fig. 14</figureCitation>
). Epigyne: posterior border slightly indented, with no conspicuous pocket (
<figureCitation id="13522A58FF875447FF0DF856FF7CF830" box="[159,247,1987,2012]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="2" pageNumber="253">Figs 25</figureCitation>
,
<figureCitation id="13522A58FF875447FE96F856FEAFF837" box="[260,292,1987,2011]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="2" pageNumber="253">30</figureCitation>
). U-shaped copulation openings connected to each other, forming an atrium. Copulation ducts begin wide and extend backwards, fold at the center and extend anteriorly. Beneath the copulation openings they fold backwards again and connect to small spermathecae, from which fertilization ducts emerge (
<figureCitation id="13522A58FF865446FB6CFF29FAD7FF39" box="[1278,1372,188,213]" captionStart="FIGURES 22 26" captionStartId="8.[151,258,1096,1118]" captionTargetBox="[162,1430,387,1075]" captionTargetId="figure@8.[151,1436,371,1075]" captionTargetPageId="8" captionText="FIGURES 22 26. Urupuyu antisana: 22 male left palp, ventral view; 23 same, palpal tibia, retrolateral view; 24 same, dorsal view; 25 epigyne, ventral view; 26 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245746/files/figure.png" pageId="3" pageNumber="254">Figs 26</figureCitation>
,
<figureCitation id="13522A58FF865446FAE2FF29FA04FF38" box="[1392,1423,188,212]" captionStart="FIGURES 27 31" captionStartId="8.[151,258,1950,1972]" captionTargetBox="[165,1429,1210,1920]" captionTargetId="figure@8.[151,1436,1196,1929]" captionTargetPageId="8" captionText="FIGURES 27 31. Urupuyu edwardsi: 27 male left palp, ventral view; 28 same, palpal tibia, retrolateral view; 29 same, dorsal view; 30 epigyne, ventral view; 31 same, cleared, dorsal view (arrow indicates initial portion of copulation duct dilated as a chamber)." httpUri="https://zenodo.org/record/245747/files/figure.png" pageId="3" pageNumber="254">31</figureCitation>
). Spinnerets (
<figureCitation id="13522A58FF865446FE8EFF4AFE15FF14" box="[284,414,223,248]" captionStart="FIGURES 18 21" captionStartId="7.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@7.[151,1436,191,1155]" captionTargetPageId="7" captionText="FIGURES 18 21. Urupuyu antisana: 18 male spinnerets; 19 right anterior lateral spinneret; 20 posterior median spinnerets; 21 left posterior lateral spinneret." httpUri="https://zenodo.org/record/245745/files/figure.png" pageId="3" pageNumber="254">Figs 1821</figureCitation>
; examined only in
<taxonomicName id="4C694D5EFF865446FDE1FF74FD7EFF14" box="[627,757,225,248]" class="Arachnida" family="Salticidae" genus="Urupuyu" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="antisana">
<emphasis id="B91DEACFFF865446FDE1FF74FD7EFF14" box="[627,757,225,248]" italics="true" pageId="3" pageNumber="254">U. antisana</emphasis>
</taxonomicName>
, male): the ALS has one major ampullate spigot (MAP) and one nubbin (n) surrounded by about nine piriform (pi) spigots (
<figureCitation id="13522A58FF865446FCE6FE91FC41FEF0" box="[884,970,260,285]" captionStart="FIGURES 18 21" captionStartId="7.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@7.[151,1436,191,1155]" captionTargetPageId="7" captionText="FIGURES 18 21. Urupuyu antisana: 18 male spinnerets; 19 right anterior lateral spinneret; 20 posterior median spinnerets; 21 left posterior lateral spinneret." httpUri="https://zenodo.org/record/245745/files/figure.png" pageId="3" pageNumber="254">Fig. 19</figureCitation>
); the PMS has a single minor ampullate spigot (mAP) and four aciniform (ac) spigots (
<figureCitation id="13522A58FF865446FD0DFEB2FD7FFEAC" box="[671,756,295,320]" captionStart="FIGURES 18 21" captionStartId="7.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@7.[151,1436,191,1155]" captionTargetPageId="7" captionText="FIGURES 18 21. Urupuyu antisana: 18 male spinnerets; 19 right anterior lateral spinneret; 20 posterior median spinnerets; 21 left posterior lateral spinneret." httpUri="https://zenodo.org/record/245745/files/figure.png" pageId="3" pageNumber="254">Fig. 20</figureCitation>
); the PLS has about five or six aciniform spigots (
<figureCitation id="13522A58FF865446FAB7FEB2FAF3FEAC" box="[1317,1400,295,320]" captionStart="FIGURES 18 21" captionStartId="7.[151,258,1176,1198]" captionTargetBox="[151,1436,191,1155]" captionTargetId="figure@7.[151,1436,191,1155]" captionTargetPageId="7" captionText="FIGURES 18 21. Urupuyu antisana: 18 male spinnerets; 19 right anterior lateral spinneret; 20 posterior median spinnerets; 21 left posterior lateral spinneret." httpUri="https://zenodo.org/record/245745/files/figure.png" pageId="3" pageNumber="254">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BD636DDFF865446FF55FED9FE35FE41" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="254">
<emphasis id="B91DEACFFF865446FF55FED9FE8DFE89" bold="true" box="[199,262,332,357]" pageId="3" pageNumber="254">Note.</emphasis>
Such depressions with pores on the carapace have also been found in non-salticids
<taxonomicName id="4C694D5EFF865446FB5DFED8FEFDFE64" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF865446FB5DFED8FAB2FE88" box="[1231,1337,333,356]" italics="true" pageId="3" pageNumber="254">Teutamus</emphasis>
Thorell, 1890 (Liocranidae)
</taxonomicName>
,
<taxonomicName id="4C694D5EFF865446FE17FEFAFCDCFE64" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[389,855,367,392]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEACFFF865446FE17FEFAFE71FE64" box="[389,506,367,392]" italics="true" pageId="3" pageNumber="254">Orthobula</emphasis>
Simon, 1897 (Phrurolithidae)
</taxonomicName>
and in trachelid spiders (
<bibRefCitation id="EFF84B2CFF865446FB1BFEE5FE39FE41" author="Ramirez" pageId="3" pageNumber="254" refString="Ramirez, M. L. (2014) The morphology and phylogeny of dionychan spiders (Araneae, Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">
Ramírez 2014: fig. 6D, character
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phylogeny
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3736556FF865446FF55FE22FD0DFE3C" box="[199,646,439,464]" pageId="3" pageNumber="254" type="distribution">
<paragraph id="8BD636DDFF865446FF55FE22FD0DFE3C" blockId="3.[151,1436,151,464]" box="[199,646,439,464]" pageId="3" pageNumber="254">
<emphasis id="B91DEACFFF865446FF55FE22FEEBFE3C" bold="true" box="[199,352,439,464]" pageId="3" pageNumber="254">Distribution.</emphasis>
Cloud forests of
<collectingCountry id="F37E764DFF865446FDB1FE22FD09FE3C" box="[547,642,439,464]" name="Ecuador" pageId="3" pageNumber="254">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>