534 lines
61 KiB
XML
534 lines
61 KiB
XML
<document id="65DC2C7015ACEEF7CAACBD6CB50CFEA1" ID-DOI="10.5281/zenodo.282919" ID-GBIF-Dataset="09d06f43-47e0-4f67-8ed9-54c16248c2c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="282919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460556013358" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2012" docId="03C087ACFF83FF9EAB84FA57FD0BC8B4" docLanguage="en" docName="zt03559p036.pdf" docOrigin="Zootaxa 3559" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cophixalus ateles Boulenger 1898" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FFF9FFD4FF82FF9AAB13FF9EFFF6C860" masterDocTitle="Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key" masterLastPageNumber="36" masterPageNumber="1" pageNumber="2" updateTime="1698319243426" updateUser="plazi">
|
||
<mods:mods id="EE7A73BD2AAC2F0C6C539793C4250E6F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6B62417A85C841DFEA73E7FB2E4B88B9">
|
||
<mods:title id="DBAE44766A5721EBDF915446ACCE3B66">Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4C677C88D5D1AD262897FC4BD0ED2224" type="personal">
|
||
<mods:role id="2B4D632EB50F14D76573F52546E4285D">
|
||
<mods:roleTerm id="261807F74AD58D9F1629083B0E4052E1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="917D021B6378DE576B5D1AA5D80AF80E">Kraus, Fred</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A4AE30C4C4E0DEA00690FDB282900371">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0F0A23E5453B704F737B0D6D19DB2034" type="host">
|
||
<mods:titleInfo id="8F3BAFA51620010F1958EE6ADB0F0BC1">
|
||
<mods:title id="3BFF9CBF1CFC2BDFBFFBF03C78504070">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="204AA05CA4F9A4528A0E164541EC9E3E">
|
||
<mods:date id="A591C64051CB2FA972D18CE5E8D1B5C7">2012</mods:date>
|
||
<mods:detail id="08564058FE0B122DF3315D1E45A5E00F" type="volume">
|
||
<mods:number id="BC5C1F031B0701796F15853FEEFE5BD4">3559</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="370DCD963B375E777138392570893F72" unit="page">
|
||
<mods:start id="0FF84F08ABB8345688F0F127A42C2C94">1</mods:start>
|
||
<mods:end id="C8761EC064873BBCD2AD1ABE056A593A">36</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D4E049DD2254EBEED123B27AD694AFD5">journal article</mods:classification>
|
||
<mods:identifier id="B2DACD9B301C75BE425BCADE968F7E73" type="DOI">10.5281/zenodo.282919</mods:identifier>
|
||
<mods:identifier id="F1A9878642045E6E091F081E953A6B4E" type="GBIF-Dataset">09d06f43-47e0-4f67-8ed9-54c16248c2c4</mods:identifier>
|
||
<mods:identifier id="128C0CB2C60252DF188D64F76BC92B2B" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="10958D2B3A57B5BA8CAE2906BD82C45B" type="Zenodo-Dep">282919</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C087ACFF83FF9EAB84FA57FD0BC8B4" ID-DOI="http://doi.org/10.5281/zenodo.6180266" ID-GBIF-Taxon="119555803" ID-Zenodo-Dep="6180266" LSID="urn:lsid:plazi:treatment:03C087ACFF83FF9EAB84FA57FD0BC8B4" httpUri="http://treatment.plazi.org/id/03C087ACFF83FF9EAB84FA57FD0BC8B4" lastPageId="4" lastPageNumber="4" pageId="1" pageNumber="2">
|
||
<subSubSection id="C3736531FF83FF9EAB84FA57FD0BC8B4" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="nomenclature">
|
||
<paragraph id="8BD636BAFF83FF9BAB84FA57FDB9CD82" blockId="1.[151,591,1480,1507]" box="[151,591,1480,1507]" pageId="1" pageNumber="2">
|
||
<heading id="D09E81D6FF83FF9BAB84FA57FDB9CD82" bold="true" box="[151,591,1480,1507]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
|
||
<taxonomicName id="4C694D39FF83FF9BAB84FA57FDB9CD82" ID-CoL="Y43H" authority="Boulenger, 1898" authorityName="Boulenger" authorityYear="1898" box="[151,591,1480,1507]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BAB84FA57FDB9CD82" bold="true" box="[151,591,1480,1507]" pageId="1" pageNumber="2">
|
||
<emphasis id="B91DEAA8FF83FF9BAB84FA57FE9BCD83" bold="true" box="[151,365,1481,1507]" italics="true" pageId="1" pageNumber="2">Cophixalus ateles</emphasis>
|
||
(Boulenger, 1898)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF83FF98AB84F991FDF0C95F" blockId="1.[151,1436,1551,2008]" lastBlockId="2.[151,1436,151,860]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
|
||
Boulenger (1898) described
|
||
<taxonomicName id="4C694D39FF83FF9BAACFF98EFD5ECE48" box="[476,680,1552,1576]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BAACFF98EFD5ECE48" box="[476,680,1552,1576]" italics="true" pageId="1" pageNumber="2">Cophixalus ateles</emphasis>
|
||
</taxonomicName>
|
||
on the basis of several specimens obtained from Moroka, a region in Central Province,
|
||
<collectingCountry id="F37E762AFF83FF9BAA9BF9ABFD9DCE2C" box="[392,619,1588,1612]" name="Papua New Guinea" pageId="1" pageNumber="2">Papua New Guinea</collectingCountry>
|
||
, that is approximately
|
||
<quantity id="4C919B5FFF83FF9BA867F9AAFC1FCE2C" box="[884,1001,1588,1612]" metricMagnitude="4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="2" unit="km" value="25.0" valueMax="30.0" valueMin="20.0">20–30 km</quantity>
|
||
E of Port Moresby in the foothills of the Owen Stanley Mts. These specimens reside in the British Museum of Natural History in London and the Museo Civico di Storia Naturale Giacomo Doria in Genoa. The most distinctive feature of this species was its very short first finger without a dilated disc, a feature that has proven diagnostic for several other
|
||
<taxonomicName id="4C694D39FF83FF9BAE09F93EFA6ACED8" box="[1306,1436,1696,1720]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEAA8FF83FF9BAE09F93EFA6ACED8" box="[1306,1436,1696,1720]" italics="true" pageId="1" pageNumber="2">Cophixalus</emphasis>
|
||
</taxonomicName>
|
||
species described since that time (
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA928F95AFD05CEBC" author="Zweifel" box="[571,755,1732,1756]" pageId="1" pageNumber="2" refString="Zweifel, R. G. (1956 a) Notes on microhylid frogs, genus Cophixalus, from New Guinea. American Museum Novitates, 1785, 1 - 8." type="journal article" year="1956" yearSuffix="a">Zweifel, 1956a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA816F95AFCB4CEBC" author="Zweifel" box="[773,834,1732,1756]" pageId="1" pageNumber="2" refString="Zweifel, R. G. (1980) Results of the Archbold Expeditions. No. 103. Frogs and lizards from the Huon Peninsula, Papua New Guinea. Bulletin of the American Museum of Natural History, 165, 390 - 434." type="journal article" year="1980">1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA840F95BFB76CEBC" author="Kraus" box="[851,1152,1732,1756]" pageId="1" pageNumber="2" refString="Kraus, F. & Allison, A. (2000) Two new species of Cophixalus from New Guinea. Journal of Herpetology, 34, 535 - 541." type="journal article" year="2000">Kraus and Allison, 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF84B4BFF83FF9BAF81F95AFB39CEBC" author="Allison" box="[1170,1231,1732,1756]" pageId="1" pageNumber="2" refString="Kraus, F. & Allison, A. (2009) New species of Cophixalus (Anura: Microhylidae) from Papua New Guinea. Zootaxa, 2128, 1 - 38." type="journal article" year="2009">2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF84B4BFF83FF9BAFF2F95AFA66CEBC" author="Gunther" box="[1249,1424,1732,1756]" pageId="1" pageNumber="2" refString="Gunther, R. (2006) Two new tiny Cophixalus species with reduced thumbs from the west of New Guinea. Herpetozoa, 19, 59 - 75." type="journal article" year="2006">Günther, 2006</bibRefCitation>
|
||
). However, since Boulenger’s description,
|
||
<taxonomicName id="4C694D39FF83FF9BA964F976FD2BCF60" box="[631,733,1768,1792]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BA964F976FD2BCF60" box="[631,733,1768,1792]" italics="true" pageId="1" pageNumber="2">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
has remained poorly known. The name had been applied to animals from the Central Highlands (
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA949F892FCE5CF44" author="Zweifel" box="[602,787,1804,1828]" pageId="1" pageNumber="2" refString="Zweifel, R. G. (1956 b) Results of the Archbold Expeditions. No. 72. Microhylid frogs from New Guinea, with descriptions of new species. American Museum Novitates, 1766, 1 - 49." type="journal article" year="1956" yearSuffix="b">Zweifel, 1956b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA837F892FC31CF44" author="Tyler" box="[804,967,1804,1828]" pageId="1" pageNumber="2" refString="Tyler, M. J. (1963 a) An account of collections of frogs from central New Guinea. Records of the Australian Museum, 26, 113 - 130." type="journal article" year="1963" yearSuffix="a">Tyler, 1963 a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA8CBF892FC11CF44" author="Tyler" box="[984,999,1804,1828]" pageId="1" pageNumber="2" refString="Tyler, M. J. (1963 b) A taxonomic study of amphibians and reptiles of the Central Highlands of New Guinea, with notes on their ecology and biology. Transactions of the Royal Society of South Australia, 36, 11 - 29." type="journal article" year="1963" yearSuffix="b">b</bibRefCitation>
|
||
) that are now referred to
|
||
<taxonomicName id="4C694D39FF83FF9BAE39F893FEBFCF27" authority="Zweifel, 1956" authorityName="Zweifel" authorityYear="1956" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BAE39F893FA6ACF44" box="[1322,1436,1804,1828]" italics="true" pageId="1" pageNumber="2">C. shellyi</emphasis>
|
||
<bibRefCitation id="EFF84B4BFF83FF9BAB84F8B1FEBFCF27" author="Zweifel" box="[151,329,1839,1863]" pageId="1" pageNumber="2" refString="Zweifel, R. G. (1956 a) Notes on microhylid frogs, genus Cophixalus, from New Guinea. American Museum Novitates, 1785, 1 - 8." type="journal article" year="1956" yearSuffix="a">Zweifel, 1956a</bibRefCitation>
|
||
</taxonomicName>
|
||
. From those mid-Twentieth-Century publications it appeared that
|
||
<taxonomicName id="4C694D39FF83FF9BAF5FF8AEFB44CF28" box="[1100,1202,1840,1864]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BAF5FF8AEFB44CF28" box="[1100,1202,1840,1864]" italics="true" pageId="1" pageNumber="2">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF83FF9BAFFEF8AEFAADCF28" box="[1261,1371,1840,1864]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BAFFEF8AEFAADCF28" box="[1261,1371,1840,1864]" italics="true" pageId="1" pageNumber="2">C. shellyi</emphasis>
|
||
</taxonomicName>
|
||
were similar species: “It is not certain that the apparently diagnostic characters of
|
||
<taxonomicName id="4C694D39FF83FF9BAF11F8CBFB98CF0C" box="[1026,1134,1876,1900]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BAF11F8CBFB98CF0C" box="[1026,1134,1876,1900]" italics="true" pageId="1" pageNumber="2">C. shellyi</emphasis>
|
||
</taxonomicName>
|
||
will continue to appear so when more specimens of
|
||
<taxonomicName id="4C694D39FF83FF9BAADEF8E6FDEECFF0" box="[461,536,1912,1936]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BAADEF8E6FDEECFF0" box="[461,536,1912,1936]" italics="true" pageId="1" pageNumber="2">shellyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF83FF9BA94AF8E6FD6ACFF0" box="[601,668,1912,1936]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BA94AF8E6FD6ACFF0" box="[601,668,1912,1936]" italics="true" pageId="1" pageNumber="2">ateles</emphasis>
|
||
</taxonomicName>
|
||
become available...” (
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA8BAF8E6FB97CFF0" author="Zweifel" box="[937,1121,1912,1936]" pageId="1" pageNumber="2" refString="Zweifel, R. G. (1956 a) Notes on microhylid frogs, genus Cophixalus, from New Guinea. American Museum Novitates, 1785, 1 - 8." type="journal article" year="1956" yearSuffix="a">Zweifel, 1956a</bibRefCitation>
|
||
), and “It does appear that
|
||
<taxonomicName id="4C694D39FF83FF9BAB84F802FF14CFD4" box="[151,226,1948,1972]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BAB84F802FF14CFD4" box="[151,226,1948,1972]" italics="true" pageId="1" pageNumber="2">shellyi</emphasis>
|
||
</taxonomicName>
|
||
is even closer to
|
||
<taxonomicName id="4C694D39FF83FF9BAAA7F802FE01CFD4" box="[436,503,1948,1972]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF83FF9BAAA7F802FE01CFD4" box="[436,503,1948,1972]" italics="true" pageId="1" pageNumber="2">ateles</emphasis>
|
||
</taxonomicName>
|
||
than originally suspected, but until additional topotypic material of the latter is obtained, the specific status of
|
||
<taxonomicName id="4C694D39FF83FF9BA932F85EFD9ACFB8" box="[545,620,1984,2008]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="shellyi">
|
||
<emphasis id="B91DEAA8FF83FF9BA932F85EFD9ACFB8" box="[545,620,1984,2008]" italics="true" pageId="1" pageNumber="2">shellyi</emphasis>
|
||
</taxonomicName>
|
||
must remain suspect...” (
|
||
<bibRefCitation id="EFF84B4BFF83FF9BA8BEF85EFBBACFB8" author="Tyler" box="[941,1100,1984,2008]" pageId="1" pageNumber="2" refString="Tyler, M. J. (1963 b) A taxonomic study of amphibians and reptiles of the Central Highlands of New Guinea, with notes on their ecology and biology. Transactions of the Royal Society of South Australia, 36, 11 - 29." type="journal article" year="1963" yearSuffix="b">Tyler, 1963b</bibRefCitation>
|
||
). This assumption of close relationship was based primarily on the shared character of a reduced first finger, but both
|
||
<bibRefCitation id="EFF84B4BFF80FF98AFA2FF09FA9CC8CF" author="Zweifel" box="[1201,1386,151,175]" pageId="2" pageNumber="3" refString="Zweifel, R. G. (1956 a) Notes on microhylid frogs, genus Cophixalus, from New Guinea. American Museum Novitates, 1785, 1 - 8." type="journal article" year="1956" yearSuffix="a">Zweifel (1956a)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFF84B4BFF80FF98AB84FF22FECCC8B4" author="Tyler" box="[151,314,188,212]" pageId="2" pageNumber="3" refString="Tyler, M. J. (1963 b) A taxonomic study of amphibians and reptiles of the Central Highlands of New Guinea, with notes on their ecology and biology. Transactions of the Royal Society of South Australia, 36, 11 - 29." type="journal article" year="1963" yearSuffix="b">Tyler (1963b)</bibRefCitation>
|
||
noted that Boulenger’s (1898) illustration of the hand of
|
||
<taxonomicName id="4C694D39FF80FF98AF16FF23FB86C8B4" box="[1029,1136,188,212]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98AF16FF23FB86C8B4" box="[1029,1136,188,212]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
did not agree well with available specimens. Those discrepancies, the faded color patterns and soft state of preservation of the
|
||
<typeStatus id="54D28818FF80FF98AE78FF7EFA6AC898" box="[1387,1436,224,248]" pageId="2" pageNumber="3">type</typeStatus>
|
||
series (
|
||
<bibRefCitation id="EFF84B4BFF80FF98ABFAFE9AFE8FC97C" author="Tyler" box="[233,377,260,284]" pageId="2" pageNumber="3" refString="Tyler, M. J. (1963 b) A taxonomic study of amphibians and reptiles of the Central Highlands of New Guinea, with notes on their ecology and biology. Transactions of the Royal Society of South Australia, 36, 11 - 29." type="journal article" year="1963" yearSuffix="b">Tyler,1963b</bibRefCitation>
|
||
), and the lack of additional topotypic material of
|
||
<taxonomicName id="4C694D39FF80FF98A8AAFE9BFBE9C97C" box="[953,1055,260,284]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98A8AAFE9BFBE9C97C" box="[953,1055,260,284]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
have made exact identity of that species uncertain to the present.
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF80FF98ABD4FED3FDE6CAE4" blockId="2.[151,1436,151,860]" pageId="2" pageNumber="3">
|
||
More recently,
|
||
<bibRefCitation id="EFF84B4BFF80FF98AA64FED3FD68C904" author="Kraus" box="[375,670,332,356]" pageId="2" pageNumber="3" refString="Kraus, F. & Allison, A. (2006) Three new species of Cophixalus (Anura: Microhylidae) from southeastern New Guinea. Herpetologica, 62, 202 - 220." type="journal article" year="2006">Kraus and Allison (2006)</bibRefCitation>
|
||
named three new species of closely related
|
||
<taxonomicName id="4C694D39FF80FF98AF8DFED2FAD6C904" box="[1182,1312,332,356]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEAA8FF80FF98AF8DFED2FAD6C904" box="[1182,1312,332,356]" italics="true" pageId="2" pageNumber="3">Cophixalus</emphasis>
|
||
</taxonomicName>
|
||
that occur in the Papuan Peninsula of New
|
||
<collectingCountry id="F37E762AFF80FF98A905FEF1FD9BC9E7" box="[534,621,367,391]" name="Guinea-Bissau" pageId="2" pageNumber="3">Guinea</collectingCountry>
|
||
. Of these,
|
||
<taxonomicName id="4C694D39FF80FF98A9F8FEEEFC99C9E8" box="[747,879,368,392]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98A9F8FEEEFC99C9E8" box="[747,879,368,392]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
was diagnosed by its small size (SV =
|
||
<quantity id="4C919B5FFF80FF98AE2CFEF1FF3EC9CC" metricMagnitude="-2" metricUnit="m" metricValue="1.305" metricValueMax="1.41" metricValueMin="1.2" pageId="2" pageNumber="3" unit="mm" value="13.05" valueMax="14.1" valueMin="12.0">12–14.1 mm</quantity>
|
||
), tuberculate dorsum, reduced first finger with a poorly developed disc lacking a circum-marginal groove, variegated or striped dorsal pattern, and a variety of morphometric details distinguishing it from its close relatives
|
||
<taxonomicName id="4C694D39FF80FF98AA13FE43FE66C994" box="[256,400,476,500]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="B91DEAA8FF80FF98AA13FE43FE66C994" box="[256,400,476,500]" italics="true" pageId="2" pageNumber="3">C. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF80FF98AADBFE43FDC8C994" box="[456,574,476,500]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B91DEAA8FF80FF98AADBFE43FDC8C994" box="[456,574,476,500]" italics="true" pageId="2" pageNumber="3">C. timidus</emphasis>
|
||
</taxonomicName>
|
||
, also named in that same paper. Directly relevant to the present discussion is that
|
||
<bibRefCitation id="EFF84B4BFF80FF98ABFCFD9EFDD3CA77" author="Kraus" box="[239,549,511,535]" pageId="2" pageNumber="3" refString="Kraus, F. & Allison, A. (2006) Three new species of Cophixalus (Anura: Microhylidae) from southeastern New Guinea. Herpetologica, 62, 202 - 220." type="journal article" year="2006">Kraus and Allison (2006)</bibRefCitation>
|
||
distinguished
|
||
<taxonomicName id="4C694D39FF80FF98A9CBFD9EFC96CA78" box="[728,864,512,536]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98A9CBFD9EFC96CA78" box="[728,864,512,536]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C694D39FF80FF98A8A1FD9EFBEACA78" box="[946,1052,512,536]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98A8A1FD9EFBEACA78" box="[946,1052,512,536]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
by the tuberculate dorsum and variegated or striped dorsal pattern of the former. This determination was based on photographs of two
|
||
<typeStatus id="54D28818FF80FF98AB84FDD9FECBCA3F" box="[151,317,583,607]" pageId="2" pageNumber="3" type="paralectotype">paralectotypes</typeStatus>
|
||
in the British Museum, which were very faded, as noted earlier (
|
||
<bibRefCitation id="EFF84B4BFF80FF98AF2DFDD9FB20CA3F" author="Tyler" box="[1086,1238,583,607]" pageId="2" pageNumber="3" refString="Tyler, M. J. (1963 b) A taxonomic study of amphibians and reptiles of the Central Highlands of New Guinea, with notes on their ecology and biology. Transactions of the Royal Society of South Australia, 36, 11 - 29." type="journal article" year="1963" yearSuffix="b">Tyler, 1963b</bibRefCitation>
|
||
), and showed no evidence of dorsal tuberculation.
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF80FF98ABD4FD11FDB4CB3C" blockId="2.[151,1436,151,860]" pageId="2" pageNumber="3">
|
||
Since the description of
|
||
<taxonomicName id="4C694D39FF80FF98AAC8FD0EFD37CAC8" box="[475,705,656,680]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98AAC8FD0EFD37CAC8" box="[475,705,656,680]" italics="true" pageId="2" pageNumber="3">Cophixalus sisyphus</emphasis>
|
||
</taxonomicName>
|
||
two additional specimens of that species (BPBM 27492, 27494) have become available from near Moroka, Central Province,
|
||
<collectingCountry id="F37E762AFF80FF98A85EFD2AFC73CAAC" box="[845,901,692,716]" name="Papua New Guinea" pageId="2" pageNumber="3">PNG</collectingCountry>
|
||
, the
|
||
<typeStatus id="54D28818FF80FF98A8AFFD2BFC1ACAAD" box="[956,1004,693,717]" pageId="2" pageNumber="3">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C694D39FF80FF98AF63FD2BFB25CAAC" box="[1136,1235,692,716]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98AF63FD2BFB25CAAC" box="[1136,1235,692,716]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
. This unexpected discovery, and recent opportunities to directly compare
|
||
<typeStatus id="54D28818FF80FF98A81FFD46FC8ECA90" box="[780,888,728,752]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C694D39FF80FF98A88CFD46FBE9CA90" box="[927,1055,728,752]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98A88CFD46FBE9CA90" box="[927,1055,728,752]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 19304–05, 19314) to the
|
||
<typeStatus id="54D28818FF80FF98AB84FD62FF08CB74" box="[151,254,764,788]" pageId="2" pageNumber="3" type="lectotype">lectotype</typeStatus>
|
||
(MSNG 29116) and seven
|
||
<typeStatus id="54D28818FF80FF98A920FD62FD23CB74" box="[563,725,764,788]" pageId="2" pageNumber="3" type="paralectotype">paralectotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C694D39FF80FF98A9E8FD63FCABCB74" box="[763,861,764,788]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98A9E8FD63FCABCB74" box="[763,861,764,788]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
(MSNG 50182 [
|
||
<specimenCount id="9D6FFD33FF80FF98AF08FD62FB51CB74" box="[1051,1191,764,788]" pageId="2" pageNumber="3" type="generic">5 specimens</specimenCount>
|
||
under same number], BMNH 1947.2.12.6–7) allowed me to determine whether the presumptive diagnostic features of
|
||
<taxonomicName id="4C694D39FF80FF98AFCEFCBEFAABCB58" box="[1245,1373,800,824]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98AFCEFCBEFAABCB58" box="[1245,1373,800,824]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
were valid vis-à-vis
|
||
<taxonomicName id="4C694D39FF80FF98AA2EFCDBFE69CB3C" box="[317,415,836,860]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98AA2EFCDBFE69CB3C" box="[317,415,836,860]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
. They are not.
|
||
</paragraph>
|
||
<caption id="DF166632FF80FF98AB84F8BEFE52CFD1" httpUri="https://zenodo.org/record/282920/files/figure.png" pageId="2" pageNumber="3" targetBox="[170,1429,926,1788]" targetPageId="2">
|
||
<paragraph id="8BD636BAFF80FF98AB84F8BEFE52CFD1" blockId="2.[151,1436,1824,1969]" pageId="2" pageNumber="3">
|
||
<emphasis id="B91DEAA8FF80FF98AB84F8BEFEE2CF55" bold="true" box="[151,276,1824,1847]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
|
||
Bivariate plot of principal component 1 (eigenvalue = 0.125) versus principal component 2 (eigenvalue = 0.025) for two paralectotypes of
|
||
<taxonomicName id="4C694D39FF80FF98AAB2F8A1FDAECF35" box="[417,600,1855,1877]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98AAB2F8A1FDAECF35" box="[417,600,1855,1877]" italics="true" pageId="2" pageNumber="3">Cophixalus ateles</emphasis>
|
||
</taxonomicName>
|
||
(filled diamonds), 24 types of
|
||
<taxonomicName id="4C694D39FF80FF98A888F8DEFBE6CF35" box="[923,1040,1855,1877]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98A888F8DEFBE6CF35" box="[923,1040,1855,1877]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
(triangles), one recent specimen of
|
||
<taxonomicName id="4C694D39FF80FF98AE97F8DEFF1BCF14" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98AE97F8DEFF1BCF14" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
from the type locality for
|
||
<taxonomicName id="4C694D39FF80FF98A910F8C0FDA8CF14" box="[515,606,1886,1908]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98A910F8C0FDA8CF14" box="[515,606,1886,1908]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
(open diamond), 14 types of
|
||
<taxonomicName id="4C694D39FF80FF98A887F8C0FC09CF14" box="[916,1023,1886,1908]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B91DEAA8FF80FF98A887F8C0FC09CF14" box="[916,1023,1886,1908]" italics="true" pageId="2" pageNumber="3">C. timidus</emphasis>
|
||
</taxonomicName>
|
||
(squares), and 49 types of
|
||
<taxonomicName id="4C694D39FF80FF98AE08F8C0FA6ACF14" box="[1307,1436,1886,1908]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="B91DEAA8FF80FF98AE08F8C0FA6ACF14" box="[1307,1436,1886,1908]" italics="true" pageId="2" pageNumber="3">C. variabilis</emphasis>
|
||
</taxonomicName>
|
||
(circles) showing no morphometric discrimination between
|
||
<taxonomicName id="4C694D39FF80FF98A9EAF8E3FCA5CFF2" box="[761,851,1916,1938]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF80FF98A9EAF8E3FCA5CFF2" box="[761,851,1916,1938]" italics="true" pageId="2" pageNumber="3">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF80FF98A896F8E3FC0FCFF2" box="[901,1017,1916,1938]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF80FF98A896F8E3FC0FCFF2" box="[901,1017,1916,1938]" italics="true" pageId="2" pageNumber="3">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
. Component loadings presented in Table 1. All specimens are male.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD636BAFF81FF99ABD4FF09FF1DCA8F" blockId="3.[151,1436,151,751]" pageId="3" pageNumber="4">
|
||
When compared side by side, there is no morphological feature to distinguish the
|
||
<typeStatus id="54D28818FF81FF99AF6DFF06FB4FC8D0" box="[1150,1209,152,176]" pageId="3" pageNumber="4">types</typeStatus>
|
||
of the two species. Further, except for the smooth skin and some rather shrunken measurements of some of the
|
||
<typeStatus id="54D28818FF81FF99AF85FF23FB27C8B5" box="[1174,1233,189,213]" pageId="3" pageNumber="4">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C694D39FF81FF99AFE6FF23FAA0C8B4" box="[1269,1366,188,212]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99AFE6FF23FAA0C8B4" box="[1269,1366,188,212]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
, there is no obvious difference in shape of body proportions to distinguish them from the
|
||
<typeStatus id="54D28818FF81FF99AF27FF7EFB69C898" box="[1076,1183,224,248]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C694D39FF81FF99AFD7FF7EFAB5C898" box="[1220,1347,224,248]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AFD7FF7EFAB5C898" box="[1220,1347,224,248]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
. This is confirmed by principal components analysis of these specimens vs.
|
||
<typeStatus id="54D28818FF81FF99A8ACFE9BFC19C97D" box="[959,1007,261,285]" pageId="3" pageNumber="4">type</typeStatus>
|
||
specimens of the closely related
|
||
<taxonomicName id="4C694D39FF81FF99AE91FE9BFF00C920" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AE91FE9BFF00C920" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D39FF81FF99AA14FEB6FE8BC920" box="[263,381,296,320]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B91DEAA8FF81FF99AA14FEB6FE8BC920" box="[263,381,296,320]" italics="true" pageId="3" pageNumber="4">C. timidus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C694D39FF81FF99AAD1FEB6FDA7C920" box="[450,593,296,320]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="B91DEAA8FF81FF99AAD1FEB6FDA7C920" box="[450,593,296,320]" italics="true" pageId="3" pageNumber="4">C. variabilis</emphasis>
|
||
</taxonomicName>
|
||
. In both bivariate (
|
||
<figureCitation id="13522A3FFF81FF99A826FEB9FC8FC95F" box="[821,889,295,319]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1847]" captionTargetBox="[170,1429,926,1788]" captionTargetId="figure@2.[151,1436,916,1800]" captionTargetPageId="2" captionText="FIGURE 1. Bivariate plot of principal component 1 (eigenvalue = 0.125) versus principal component 2 (eigenvalue = 0.025) for two paralectotypes of Cophixalus ateles (filled diamonds), 24 types of C. sisyphus (triangles), one recent specimen of C. sisyphus from the type locality for C. ateles (open diamond), 14 types of C. timidus (squares), and 49 types of C. variabilis (circles) showing no morphometric discrimination between C. ateles and C. sisyphus. Component loadings presented in Table 1. All specimens are male." httpUri="https://zenodo.org/record/282920/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
) and 3-dimensional (not shown) plots of wellpreserved specimens, the two specimens of
|
||
<taxonomicName id="4C694D39FF81FF99A996FED3FD11C904" box="[645,743,332,356]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99A996FED3FD11C904" box="[645,743,332,356]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
cluster with the
|
||
<typeStatus id="54D28818FF81FF99A8B7FED3FC22C905" box="[932,980,333,357]" pageId="3" pageNumber="4">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C694D39FF81FF99AF50FED3FB35C904" box="[1091,1219,332,356]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AF50FED3FB35C904" box="[1091,1219,332,356]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
to the exclusion of the similar
|
||
<taxonomicName id="4C694D39FF81FF99AA07FEEEFE73C9E8" box="[276,389,368,392]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B91DEAA8FF81FF99AA07FEEEFE73C9E8" box="[276,389,368,392]" italics="true" pageId="3" pageNumber="4">C. timidus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF81FF99AAA8FEEEFDB0C9E8" box="[443,582,368,392]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="B91DEAA8FF81FF99AAA8FEEEFDB0C9E8" box="[443,582,368,392]" italics="true" pageId="3" pageNumber="4">C. variabilis</emphasis>
|
||
</taxonomicName>
|
||
. The specimens of
|
||
<taxonomicName id="4C694D39FF81FF99A80DFEEEFC89C9E8" box="[798,895,368,392]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99A80DFEEEFC89C9E8" box="[798,895,368,392]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
are, indeed, at one extreme of the distribution of
|
||
<taxonomicName id="4C694D39FF81FF99AB84FE0BFEE1C9CC" box="[151,279,404,428]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AB84FE0BFEE1C9CC" box="[151,279,404,428]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
along PC1. This axis largely reflects specimen size, judging by the uniformly positive loadings of the constituent variables (
|
||
<tableCitation id="C6EB0301FF81FF99AA86FE29FE1AC9AF" box="[405,492,439,463]" captionStart="TABLE 1" captionStartId="3.[151,239,799,823]" captionTargetBox="[165,1351,889,1570]" captionTargetPageId="3" captionText="TABLE 1. Loadings for log-transformed characters used in principal components analysis of covariance matrix for adult males of Cophixalus ateles, C. sisyphus, C. timidus, and C. variabilis. Included specimens are listed in the Appendix." httpUri="http://table.plazi.org/id/DF166632FF81FF99AB84FC81FA8CCB37" pageId="3" pageNumber="4" tableUuid="DF166632FF81FF99AB84FC81FA8CCB37">Table 1</tableCitation>
|
||
). The position of the
|
||
<taxonomicName id="4C694D39FF81FF99A9E6FE26FCADC9B0" box="[757,859,440,464]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99A9E6FE26FCADC9B0" box="[757,859,440,464]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
specimens at the low end of PC1 seems to be a reflection of their smaller measurements for a number of features relative to their overall body size. This apparently reflects measurement error—especially with respect to disc width—due to the soft and somewhat shrunken state of preservation of these specimens. When disc widths for these two specimens are artificially increased to be in line with those obtained from the recent
|
||
<taxonomicName id="4C694D39FF81FF99A955FDD6FD3CCA00" box="[582,714,584,608]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99A955FDD6FD3CCA00" box="[582,714,584,608]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D28818FF81FF99A9C6FDD6FCE6CA00" box="[725,784,584,608]" pageId="3" pageNumber="4">types</typeStatus>
|
||
of those body sizes, both
|
||
<taxonomicName id="4C694D39FF81FF99AF5CFDD6FB43CA00" box="[1103,1205,584,608]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99AF5CFDD6FB43CA00" box="[1103,1205,584,608]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D28818FF81FF99AFD2FDD6FB0ACA00" box="[1217,1276,584,608]" pageId="3" pageNumber="4">types</typeStatus>
|
||
more closely approach the distribution of the
|
||
<taxonomicName id="4C694D39FF81FF99AAEDFDF3FD8BCAE4" box="[510,637,620,644]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AAEDFDF3FD8BCAE4" box="[510,637,620,644]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D28818FF81FF99A997FDF3FD34CAE5" box="[644,706,621,645]" pageId="3" pageNumber="4">types</typeStatus>
|
||
, with the
|
||
<taxonomicName id="4C694D39FF81FF99A823FDF3FC65CAE4" box="[816,915,620,644]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99A823FDF3FC65CAE4" box="[816,915,620,644]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
specimen lower on the graph now lying in the midst of adjacent
|
||
<taxonomicName id="4C694D39FF81FF99AA71FD0EFE14CAC8" box="[354,482,656,680]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AA71FD0EFE14CAC8" box="[354,482,656,680]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
specimens (graph not shown). Thus the little difference that appears between the
|
||
<taxonomicName id="4C694D39FF81FF99AE91FD0EFF2ECAAC" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99AE91FD0EFF2ECAAC" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D28818FF81FF99ABF7FD2BFEE9CAAD" box="[228,287,693,717]" pageId="3" pageNumber="4">types</typeStatus>
|
||
and
|
||
<taxonomicName id="4C694D39FF81FF99AA4EFD2BFE14CAAC" box="[349,482,692,716]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AA4EFD2BFE14CAAC" box="[349,482,692,716]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D28818FF81FF99AAFFFD2BFDD1CAAD" box="[492,551,693,717]" pageId="3" pageNumber="4">types</typeStatus>
|
||
in multivariate analysis seems to merely reflect differences in preservation quality.
|
||
</paragraph>
|
||
<caption id="DF166632FF81FF99AB84FC81FA8CCB37" ID-Table-UUID="DF166632FF81FF99AB84FC81FA8CCB37" httpUri="http://table.plazi.org/id/DF166632FF81FF99AB84FC81FA8CCB37" pageId="3" pageNumber="4" targetBox="[165,1351,889,1570]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="8BD636BAFF81FF99AB84FC81FA8CCB37" blockId="3.[151,1435,799,855]" pageId="3" pageNumber="4">
|
||
<emphasis id="B91DEAA8FF81FF99AB84FC81FEFFCB57" bold="true" box="[151,265,799,823]" pageId="3" pageNumber="4">TABLE 1.</emphasis>
|
||
Loadings for log-transformed characters used in principal components analysis of covariance matrix for adult males of
|
||
<taxonomicName id="4C694D39FF81FF99ABEBFCDEFE40CB37" box="[248,438,832,855]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99ABEBFCDEFE40CB37" box="[248,438,832,855]" italics="true" pageId="3" pageNumber="4">Cophixalus ateles</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D39FF81FF99AAD7FCDEFDCBCB37" box="[452,573,832,855]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AAD7FCDEFDCBCB37" box="[452,573,832,855]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D39FF81FF99A958FCDEFD4ECB37" box="[587,696,832,855]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B91DEAA8FF81FF99A958FCDEFD4ECB37" box="[587,696,832,855]" italics="true" pageId="3" pageNumber="4">C. timidus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C694D39FF81FF99A9E0FCDEFC8ECB37" box="[755,888,832,855]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="variabilis">
|
||
<emphasis id="B91DEAA8FF81FF99A9E0FCDEFC8ECB37" box="[755,888,832,855]" italics="true" pageId="3" pageNumber="4">C. variabilis</emphasis>
|
||
</taxonomicName>
|
||
. Included specimens are listed in the Appendix.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD636BAFF81FF99A819FCE7FAC9CE42" pageId="3" pageNumber="4">
|
||
<table id="F969C41AFF810065ABB6FCE7FAB1CE42" box="[165,1351,889,1570]" gridcols="4" gridrows="16" pageId="3" pageNumber="4">
|
||
<tr id="355934F8FF810065ABB6FCE7FAB1CBEE" box="[165,1351,889,910]" gridrow="0" pageId="3" pageNumber="4" rowspan-0="1">
|
||
<th id="76885D84FF810065A9E5FCE7FCBFCBEE" box="[758,841,889,910]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">PC1</th>
|
||
<th id="76885D84FF810065A8EAFCE7FBBACBEE" box="[1017,1100,889,910]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">PC2</th>
|
||
<th id="76885D84FF810065AFE6FCE7FAB1CBEE" box="[1269,1351,889,910]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">PC3</th>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FC3AFAB1CBDA" box="[165,1351,932,954]" gridrow="1" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FC3AFE3BCBDA" box="[165,461,932,954]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Eigenvalue</th>
|
||
<td id="76885D84FF810065A9E5FC3AFCBFCBDA" box="[758,841,932,954]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">0.12466</td>
|
||
<td id="76885D84FF810065A8EAFC3AFBBACBDA" box="[1017,1100,932,954]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">0.02471</td>
|
||
<td id="76885D84FF810065AFE6FC3AFAB1CBDA" box="[1269,1351,932,954]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">0.01154</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FC4EFAB1CB86" box="[165,1351,976,998]" gridrow="2" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FC4EFE3BCB86" box="[165,461,976,998]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Cumulative percent variation</th>
|
||
<td id="76885D84FF810065A9E5FC4EFCBFCB86" box="[758,841,976,998]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">0.678</td>
|
||
<td id="76885D84FF810065A8EAFC4EFBBACB86" box="[1017,1100,976,998]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">0.812</td>
|
||
<td id="76885D84FF810065AFE6FC4EFAB1CB86" box="[1269,1351,976,998]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">0.875</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FC62FAB1CC72" box="[165,1351,1020,1042]" gridrow="3" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76885D84FF810065ABB6FC62FE3BCC72" box="[165,461,1020,1042]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Variable</th>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FBB6FAB1CC5E" box="[165,1351,1064,1086]" gridrow="4" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FBB6FE3BCC5E" box="[165,461,1064,1086]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">logSV</th>
|
||
<td id="76885D84FF810065A9E5FBB6FCBFCC5E" box="[758,841,1064,1086]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">0.269</td>
|
||
<td id="76885D84FF810065A8EAFBB6FBBACC5E" box="[1017,1100,1064,1086]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">0.087</td>
|
||
<td id="76885D84FF810065AFE6FBB6FAB1CC5E" box="[1269,1351,1064,1086]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">0.202</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FBCAFAB1CC0A" box="[165,1351,1108,1130]" gridrow="5" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FBCAFE3BCC0A" box="[165,461,1108,1130]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">logTL</th>
|
||
<td id="76885D84FF810065A9E5FBCAFCBFCC0A" box="[758,841,1108,1130]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">0.361</td>
|
||
<td id="76885D84FF810065A8EAFBCAFBBACC0A" box="[1017,1100,1108,1130]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">0.011</td>
|
||
<td id="76885D84FF810065AFE6FBCAFAB1CC0A" box="[1269,1351,1108,1130]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">0.292</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FB1EFAB1CCF6" box="[165,1351,1152,1174]" gridrow="6" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FB1EFE3BCCF6" box="[165,461,1152,1174]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">logEN</th>
|
||
<td id="76885D84FF810065A9E5FB1EFCBFCCF6" box="[758,841,1152,1174]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">0.248</td>
|
||
<td id="76885D84FF810065A8EAFB1EFBBACCF6" box="[1017,1100,1152,1174]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">0.097</td>
|
||
<td id="76885D84FF810065AFE6FB1EFAB1CCF6" box="[1269,1351,1152,1174]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">0.081</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FB32FAB1CCA2" box="[165,1351,1196,1218]" gridrow="7" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FB32FE3BCCA2" box="[165,461,1196,1218]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">logIN</th>
|
||
<td id="76885D84FF810065A9E5FB32FCBFCCA2" box="[758,841,1196,1218]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">0.125</td>
|
||
<td id="76885D84FF810065A8EAFB32FBBACCA2" box="[1017,1100,1196,1218]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">0.008</td>
|
||
<td id="76885D84FF810065AFE6FB32FAB1CCA2" box="[1269,1351,1196,1218]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">-0.031</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FB46FAB1CC8E" box="[165,1351,1240,1262]" gridrow="8" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FB46FE3BCC8E" box="[165,461,1240,1262]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">logSN</th>
|
||
<td id="76885D84FF810065A9E5FB46FCBFCC8E" box="[758,841,1240,1262]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">0.228</td>
|
||
<td id="76885D84FF810065A8EAFB46FBBACC8E" box="[1017,1100,1240,1262]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">0.079</td>
|
||
<td id="76885D84FF810065AFE6FB46FAB1CC8E" box="[1269,1351,1240,1262]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">0.042</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FA9AFAB1CD7A" box="[165,1351,1284,1306]" gridrow="9" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FA9AFE3BCD7A" box="[165,461,1284,1306]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">logTY</th>
|
||
<td id="76885D84FF810065A9E5FA9AFCBFCD7A" box="[758,841,1284,1306]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">0.267</td>
|
||
<td id="76885D84FF810065A8EAFA9AFBBACD7A" box="[1017,1100,1284,1306]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">-0.951</td>
|
||
<td id="76885D84FF810065AFE6FA9AFAB1CD7A" box="[1269,1351,1284,1306]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">-0.115</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FAAEFAB1CD26" box="[165,1351,1328,1350]" gridrow="10" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FAAEFE3BCD26" box="[165,461,1328,1350]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">logEY</th>
|
||
<td id="76885D84FF810065A9E5FAAEFCBFCD26" box="[758,841,1328,1350]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">0.208</td>
|
||
<td id="76885D84FF810065A8EAFAAEFBBACD26" box="[1017,1100,1328,1350]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">0.064</td>
|
||
<td id="76885D84FF810065AFE6FAAEFAB1CD26" box="[1269,1351,1328,1350]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">0.258</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FAC2FAB1CD12" box="[165,1351,1372,1394]" gridrow="11" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FAC2FE3BCD12" box="[165,461,1372,1394]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">logHW</th>
|
||
<td id="76885D84FF810065A9E5FAC2FCBFCD12" box="[758,841,1372,1394]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">0.240</td>
|
||
<td id="76885D84FF810065A8EAFAC2FBBACD12" box="[1017,1100,1372,1394]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">0.017</td>
|
||
<td id="76885D84FF810065AFE6FAC2FAB1CD12" box="[1269,1351,1372,1394]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">0.149</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FA16FAB1CDFE" box="[165,1351,1416,1438]" gridrow="12" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FA16FE3BCDFE" box="[165,461,1416,1438]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">logHL</th>
|
||
<td id="76885D84FF810065A9E5FA16FCBFCDFE" box="[758,841,1416,1438]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">0.240</td>
|
||
<td id="76885D84FF810065A8EAFA16FBBACDFE" box="[1017,1100,1416,1438]" gridcol="2" gridrow="12" pageId="3" pageNumber="4">0.014</td>
|
||
<td id="76885D84FF810065AFE6FA16FAB1CDFE" box="[1269,1351,1416,1438]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">0.194</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FA2AFAB1CDAA" box="[165,1351,1460,1482]" gridrow="13" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FA2AFE3BCDAA" box="[165,461,1460,1482]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">logHandL</th>
|
||
<td id="76885D84FF810065A9E5FA2AFCBFCDAA" box="[758,841,1460,1482]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">0.397</td>
|
||
<td id="76885D84FF810065A8EAFA2AFBBACDAA" box="[1017,1100,1460,1482]" gridcol="2" gridrow="13" pageId="3" pageNumber="4">0.109</td>
|
||
<td id="76885D84FF810065AFE6FA2AFAB1CDAA" box="[1269,1351,1460,1482]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">0.227</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6FA7EFAB1CD96" box="[165,1351,1504,1526]" gridrow="14" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6FA7EFE3BCD96" box="[165,461,1504,1526]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">log3rdF</th>
|
||
<td id="76885D84FF810065A9E5FA7EFCBFCD96" box="[758,841,1504,1526]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">0.469</td>
|
||
<td id="76885D84FF810065A8EAFA7EFBBACD96" box="[1017,1100,1504,1526]" gridcol="2" gridrow="14" pageId="3" pageNumber="4">0.172</td>
|
||
<td id="76885D84FF810065AFE6FA7EFAB1CD96" box="[1269,1351,1504,1526]" gridcol="3" gridrow="14" pageId="3" pageNumber="4">-0.455</td>
|
||
</tr>
|
||
<tr id="355934F8FF810065ABB6F992FAB1CE42" box="[165,1351,1548,1570]" gridrow="15" pageId="3" pageNumber="4">
|
||
<th id="76885D84FF810065ABB6F992FE3BCE42" box="[165,461,1548,1570]" gridcol="0" gridrow="15" pageId="3" pageNumber="4">log4thT</th>
|
||
<td id="76885D84FF810065A9E5F992FCBFCE42" box="[758,841,1548,1570]" gridcol="1" gridrow="15" pageId="3" pageNumber="4">0.246</td>
|
||
<td id="76885D84FF810065A8EAF992FBBACE42" box="[1017,1100,1548,1570]" gridcol="2" gridrow="15" pageId="3" pageNumber="4">0.160</td>
|
||
<td id="76885D84FF810065AFE6F992FAB1CE42" box="[1269,1351,1548,1570]" gridcol="3" gridrow="15" pageId="3" pageNumber="4">-0.683</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF81FF9EABD4F9FAFD0BC8B4" blockId="3.[151,1436,1636,2020]" lastBlockId="4.[151,1436,151,212]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Color pattern for several of the
|
||
<typeStatus id="54D28818FF81FF99A95BF9FBFD8ECE1D" box="[584,632,1637,1661]" pageId="3" pageNumber="4">type</typeStatus>
|
||
specimens is faded, but others show the dorsal and ventral pattern elements typical of
|
||
<taxonomicName id="4C694D39FF81FF99AA6AF917FE0CCEC0" box="[377,506,1672,1696]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AA6AF917FE0CCEC0" box="[377,506,1672,1696]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
, including dark dorsal blotches and supratympanic line (two specimens), a broad mid-dorsal pale stripe margined in black (one specimen), dark punctations on the chin and chest (six specimens), and paired pale pectoral spots (two specimens). Furthermore, in basing their diagnosis of
|
||
<taxonomicName id="4C694D39FF81FF99AF82F94FFAE4CE88" box="[1169,1298,1744,1768]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AF82F94FFAE4CE88" box="[1169,1298,1744,1768]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
vis-à-vis
|
||
<taxonomicName id="4C694D39FF81FF99AE91F94FFF2ECF6C" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99AE91F94FFF2ECF6C" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
on the smooth skin and lack of color pattern in the latter,
|
||
<bibRefCitation id="EFF84B4BFF81FF99A87AF96BFB7FCF6C" author="Kraus" box="[873,1161,1780,1804]" pageId="3" pageNumber="4" refString="Kraus, F. & Allison, A. (2006) Three new species of Cophixalus (Anura: Microhylidae) from southeastern New Guinea. Herpetologica, 62, 202 - 220." type="journal article" year="2006">Kraus and Allison (2006)</bibRefCitation>
|
||
overlooked the fact that Boulenger (1898) mentioned that
|
||
<taxonomicName id="4C694D39FF81FF99A901F887FD82CF50" box="[530,628,1816,1840]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99A901F887FD82CF50" box="[530,628,1816,1840]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
had small warts and some pattern elements, although the latter were not specified in detail beyond noting the occasional presence of black dorsolateral lines and a light vertebral stripe. The frequent hourglass pattern seen on the dorsum and the dark flecking and blotching often seen in
|
||
<taxonomicName id="4C694D39FF81FF99AFDAF8FFFE83CFFC" authority="Kraus and Allison, 2006" authorityName="Kraus and Allison" authorityYear="2006" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AFDAF8FFFABFCF18" box="[1225,1353,1888,1912]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF84B4BFF81FF99AE44F8FEFE9BCFFC" author="Kraus" pageId="3" pageNumber="4" refString="Kraus, F. & Allison, A. (2006) Three new species of Cophixalus (Anura: Microhylidae) from southeastern New Guinea. Herpetologica, 62, 202 - 220." type="journal article" year="2006">Kraus and Allison, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
were not mentioned by Boulenger, although the black dorsolateral lines he did mention presumably represent the lateral margins of the hourglass markings described for
|
||
<taxonomicName id="4C694D39FF81FF99AF52F837FB35CFA0" box="[1089,1219,1960,1984]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99AF52F837FB35CFA0" box="[1089,1219,1960,1984]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
. Given the lack of structural differences between
|
||
<taxonomicName id="4C694D39FF81FF99AAE4F853FDAACF84" box="[503,604,1996,2020]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF81FF99AAE4F853FDAACF84" box="[503,604,1996,2020]" italics="true" pageId="3" pageNumber="4">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D39FF81FF99A98BF853FCEDCF84" box="[664,795,1996,2020]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF81FF99A98BF853FCEDCF84" box="[664,795,1996,2020]" italics="true" pageId="3" pageNumber="4">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
and the fact that the few color-pattern details given by Boulenger for
|
||
<taxonomicName id="4C694D39FF86FF9EAA29FF06FE6DC8D0" box="[314,411,152,176]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF86FF9EAA29FF06FE6DC8D0" box="[314,411,152,176]" italics="true" pageId="4" pageNumber="5">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
are consistent with some of the patterns seen in
|
||
<taxonomicName id="4C694D39FF86FF9EA8ABFF06FBCEC8D0" box="[952,1080,152,176]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF86FF9EA8ABFF06FBCEC8D0" box="[952,1080,152,176]" italics="true" pageId="4" pageNumber="5">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
, I synonymize
|
||
<taxonomicName id="4C694D39FF86FF9EAFF7FF06FA95C8D0" box="[1252,1379,152,176]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="sisyphus">
|
||
<emphasis id="B91DEAA8FF86FF9EAFF7FF06FA95C8D0" box="[1252,1379,152,176]" italics="true" pageId="4" pageNumber="5">C. sisyphus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C694D39FF86FF9EAB84FF23FF0EC8B4" box="[151,248,188,212]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF86FF9EAB84FF23FF0EC8B4" box="[151,248,188,212]" italics="true" pageId="4" pageNumber="5">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
. The taxonomic history of
|
||
<taxonomicName id="4C694D39FF86FF9EA939FF23FD7DC8B4" box="[554,651,188,212]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ateles">
|
||
<emphasis id="B91DEAA8FF86FF9EA939FF23FD7DC8B4" box="[554,651,188,212]" italics="true" pageId="4" pageNumber="5">C. ateles</emphasis>
|
||
</taxonomicName>
|
||
becomes:
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |