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<document id="091380DCBD0D53AE0D635D88BDD994BC" ID-DOI="10.5281/zenodo.214120" ID-GBIF-Dataset="b78e4624-fed5-4ee5-a98d-1590b9adf040" ID-ISSN="1175-5326" ID-Zenodo-Dep="214120" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460514526012" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Itou, Isamu" docDate="2012" docId="03B45B33FFC3FFF46BFE5EEAC06AA640" docLanguage="en" docName="zt03440p062.pdf" docOrigin="Zootaxa 3440" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onycocaris hayamaensis Komai &amp; Itou, 2012, n. sp." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="FF8D234BFFC2FFFF6B695F58C41EA048" masterDocTitle="A new species of the pontoniine shrimp genus Onycocaris Nobili, 1904 (Crustacea: Decapoda: Caridea: Palaemonidae) from Sagami Bay, central Japan" masterLastPageNumber="62" masterPageNumber="50" pageNumber="51" updateTime="1698313762796" updateUser="plazi">
<mods:mods id="AD6F3966D1F2BD0F8A14BC2B7F951674" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A19B311B68B48E943C273C4487F0951F">
<mods:title id="6F881761743245861972910767A4A052">A new species of the pontoniine shrimp genus Onycocaris Nobili, 1904 (Crustacea: Decapoda: Caridea: Palaemonidae) from Sagami Bay, central Japan</mods:title>
</mods:titleInfo>
<mods:name id="5AA27CE1992B224734D3E2D196A5C45A" type="personal">
<mods:role id="D0F56C9F9EBFA0B3D89E033A5106453D">
<mods:roleTerm id="FE536D42767F6D9869340513A2A02302">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB023D1A303A6B2C38EBE206CE39F53E">Komai, Tomoyuki</mods:namePart>
</mods:name>
<mods:name id="68D89BB4C8E4F0A3A3C29BCA4C442A2A" type="personal">
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<mods:namePart id="C11E4C421230AC13464B2A133F5DAD60">Itou, Isamu</mods:namePart>
</mods:name>
<mods:typeOfResource id="552C3487E6A56E8F94BA143E098FC1E0">text</mods:typeOfResource>
<mods:relatedItem id="6D1B8915674EB53906F699CA8A17B44D" type="host">
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<mods:title id="3927B2824DF5D8BE9EBBA7C7F2564D8A">Zootaxa</mods:title>
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<mods:date id="0C8C13DC5A2CDFA9F6350B7B7AC886C7">2012</mods:date>
<mods:detail id="51B4EE9A692B8055446C5FAEAC74F391" type="volume">
<mods:number id="F8683F0A01CA5E92081661E981A6CE96">3440</mods:number>
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<mods:classification id="BFFFD45C98BC6AFE3262E465CA0C90AE">journal article</mods:classification>
<mods:identifier id="4DA852D0A5459329469C5F8CD2CDA0EF" type="DOI">10.5281/zenodo.214120</mods:identifier>
<mods:identifier id="7650A79ADBAE0ED23C0A2A755BB5B00F" type="GBIF-Dataset">b78e4624-fed5-4ee5-a98d-1590b9adf040</mods:identifier>
<mods:identifier id="D814894150FBD33C9BFEB88092BACAA1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1FBE6B20AA0B47987E4C5E3616981789" type="Zenodo-Dep">214120</mods:identifier>
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<treatment id="03B45B33FFC3FFF46BFE5EEAC06AA640" ID-DOI="http://doi.org/10.5281/zenodo.6166556" ID-GBIF-Taxon="119523858" ID-Zenodo-Dep="6166556" LSID="urn:lsid:plazi:treatment:03B45B33FFC3FFF46BFE5EEAC06AA640" httpUri="http://treatment.plazi.org/id/03B45B33FFC3FFF46BFE5EEAC06AA640" lastPageId="11" lastPageNumber="61" pageId="1" pageNumber="51">
<subSubSection id="C307B9AEFFC3FFFE6BFE5EEAC515A1A5" pageId="1" pageNumber="51" type="nomenclature">
<paragraph id="8BA2EA25FFC3FFFE6BFE5EEAC612A184" blockId="1.[151,524,434,493]" box="[151,524,434,460]" pageId="1" pageNumber="51">
<heading id="D0EA5D49FFC3FFFE6BFE5EEAC612A184" bold="true" box="[151,524,434,460]" fontSize="11" level="1" pageId="1" pageNumber="51" reason="1">
<emphasis id="B9693637FFC3FFFE6BFE5EEAC612A184" bold="true" box="[151,524,434,460]" pageId="1" pageNumber="51">
<taxonomicName id="4C1D91A6FFC3FFFE6BFE5EEAC5DDA184" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[151,451,434,460]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="51" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC3FFFE6BFE5EEAC5DDA184" bold="true" box="[151,451,434,460]" italics="true" pageId="1" pageNumber="51">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC3FFFE6AA35EEAC612A184" box="[458,524,434,460]" pageId="1" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2EA25FFC3FFFE6BFE5E8DC515A1A5" blockId="1.[151,524,434,493]" box="[151,267,469,493]" pageId="1" pageNumber="51">
(
<figureCitation id="1326F6A0FFC3FFFE6BF65E8DC51CA1A5" box="[159,258,469,493]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="1.[151,250,1448,1471]" captionStartId-1="2.[151,250,1778,1801]" captionStartId-2="3.[151,250,1857,1880]" captionStartId-3="5.[151,250,1905,1928]" captionStartId-4="7.[151,250,1880,1903]" captionStartId-5="8.[151,250,1901,1924]" captionStartId-6="10.[151,250,1941,1964]" captionTargetBox-0="[151,1435,765,1424]" captionTargetBox-1="[184,1406,158,1777]" captionTargetBox-2="[226,1348,185,1831]" captionTargetBox-3="[181,1406,180,1904]" captionTargetBox-4="[183,1388,182,1879]" captionTargetBox-5="[211,1383,217,1854]" captionTargetBox-6="[171,1414,317,1922]" captionTargetId-0="figure@1.[151,1436,765,1424]" captionTargetId-1="figure@2.[170,1434,123,1803]" captionTargetId-2="figure@3.[201,1387,147,1845]" captionTargetId-3="figure@5.[151,1436,173,1912]" captionTargetId-4="figure@7.[172,1416,173,1890]" captionTargetId-5="figure@8.[176,1410,193,1880]" captionTargetId-6="figure@10.[171,1415,317,1922]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionText-0="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." captionText-1="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." captionText-2="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." captionText-3="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." captionText-4="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." captionText-5="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." captionText-6="FIGURE 7. Underwater photographs. A, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm) (indicated by red arrow), and host sponge Haliclona sp.; B, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm); C, Onycocaris hayamaensis n. sp., male, spouse of holotype, not collected." httpUri-0="https://zenodo.org/record/214121/files/figure.png" httpUri-1="https://zenodo.org/record/214122/files/figure.png" httpUri-2="https://zenodo.org/record/214123/files/figure.png" httpUri-3="https://zenodo.org/record/214124/files/figure.png" httpUri-4="https://zenodo.org/record/214125/files/figure.png" httpUri-5="https://zenodo.org/record/214126/files/figure.png" httpUri-6="https://zenodo.org/record/214127/files/figure.png" pageId="1" pageNumber="51">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFC3FFF66BFE5D45C5C2A242" lastPageId="9" lastPageNumber="59" pageId="1" pageNumber="51" type="materials_examined">
<paragraph id="8BA2EA25FFC3FFFE6BFE5D45C07BA211" blockId="1.[151,1437,541,709]" pageId="1" pageNumber="51">
<emphasis id="B9693637FFC3FFFE6BFE5D45C567A27E" bold="true" box="[151,377,541,566]" pageId="1" pageNumber="51">Material examined</emphasis>
.
<typeStatus id="54A65487FFC3FFFE6AEF5D45C5EDA27D" box="[390,499,541,565]" pageId="1" pageNumber="51" type="holotype">Holotype</typeStatus>
: ovigerous female (cl
<quantity id="4CE547C0FFC3FFFE69815D45C75FA27D" box="[744,833,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="51" unit="mm" value="3.4">3.4 mm</quantity>
), Sagami Bay, Hayama, Kanagawa Prefecture,
<quantity id="4CE547C0FFC3FFFE6E265D45C189A27D" box="[1359,1431,541,565]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="51" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
,
<date id="FFA3CCE5FFC3FFFE6BFE5D19C552A211" box="[151,332,577,601]" pageId="1" pageNumber="51" value="2011-08-14">14 August 2011</date>
,
<collectionCode id="ED0C72E0FFC3FFFE6A3E5D19C5AAA211" box="[343,436,577,601]" pageId="1" pageNumber="51">SCUBA</collectionCode>
diving, coll. Satoko Komai and Isamu Itou, CBM-ZC 11215.
</paragraph>
<paragraph id="8BA2EA25FFC3FFFE6BFE5D3EC61BA28D" blockId="1.[151,1437,541,709]" pageId="1" pageNumber="51">
<typeStatus id="54A65487FFC3FFFE6BFE5D3EC516A236" box="[151,264,614,638]" pageId="1" pageNumber="51" type="paratype">Paratypes</typeStatus>
: 3 ovigerous females (cl
<quantity id="4CE547C0FFC3FFFE69775D3DC6B7A235" box="[542,681,613,637]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.8" metricValueMin="2.9" pageId="1" pageNumber="51" unit="mm" value="3.3499999999999996" valueMax="3.8" valueMin="2.9">2.9–3.8 mm</quantity>
),
<specimenCount id="9D1B21ACFFC3FFFE69D55D3DC70CA235" box="[700,786,613,637]" pageId="1" pageNumber="51" type="male">3 males</specimenCount>
(cl
<quantity id="4CE547C0FFC3FFFE68545D3DC7D6A235" box="[829,968,613,637]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="1" pageNumber="51" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
), same data as
<typeStatus id="54A65487FFC3FFFE6F075D3DC0CDA235" box="[1134,1235,613,637]" pageId="1" pageNumber="51" type="holotype">holotype</typeStatus>
, CBM-ZC 11216.
<emphasis id="B9693637FFC3FFFE6BAE5DD1C54FA2EA" bold="true" box="[199,337,649,674]" pageId="1" pageNumber="51">Description</emphasis>
.
<emphasis id="B9693637FFC3FFFE6A0B5DD1C5DFA2E9" box="[354,449,649,673]" italics="true" pageId="1" pageNumber="51">Females</emphasis>
. Body (
<figureCitation id="1326F6A0FFC3FFFE69495DD1C674A2E9" box="[544,618,649,673]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="1" pageNumber="51">Figs 1</figureCitation>
,
<figureCitation id="1326F6A0FFC3FFFE69125DD1C697A2E9" box="[635,649,649,673]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="1" pageNumber="51">2</figureCitation>
A–C) stoutly built, plump, subcylindrical; relatively large for genus, attaining
<quantity id="4CE547C0FFC3FFFE6A695DF5C548A28D" box="[256,342,685,709]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="51" unit="mm" value="3.5">3.5 mm</quantity>
in cl in female.
</paragraph>
<caption id="DF62BAADFFC3FFFE6BFE5AF0C4C2A595" httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="1" pageNumber="51" targetBox="[151,1435,765,1424]" targetPageId="1">
<paragraph id="8BA2EA25FFC3FFFE6BFE5AF0C4C2A595" blockId="1.[151,1436,1448,1501]" pageId="1" pageNumber="51">
<emphasis id="B9693637FFC3FFFE6BFE5AF0C50AA5F6" bold="true" box="[151,276,1448,1471]" pageId="1" pageNumber="51">FIGURE 1.</emphasis>
<taxonomicName id="4C1D91A6FFC3FFFE6A725AF1C604A5F6" box="[283,538,1448,1470]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="51" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC3FFFE6A725AF1C604A5F6" box="[283,538,1448,1470]" italics="true" pageId="1" pageNumber="51">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC3FFFE694B5AF1C642A5F6" box="[546,604,1449,1470]" pageId="1" pageNumber="51" rank="species">
<emphasis id="B9693637FFC3FFFE694B5AF1C642A5F6" bold="true" box="[546,604,1449,1470]" pageId="1" pageNumber="51">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFC3FFFE6BAE597CC7D9A683" blockId="1.[151,1436,1571,2027]" pageId="1" pageNumber="51">
Rostrum (
<figureCitation id="1326F6A0FFC3FFFE6A5F597BC569A673" box="[310,375,1571,1595]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="1" pageNumber="51">Fig. 3</figureCitation>
A) short, broadly triangular, reaching midlength of eyestalks; dorsal surface with short, distinct carina extending to level of posterior margin of orbit, without teeth; lateral margin merging into orbital margin. Carapace smooth, glabrous; dorsal surface rounded, sloping down toward rostrum; orbital margin concave; antennal tooth absent; inferior orbital angle broadly subtriangular, subacutely pointed, extending as far as rostrum; pterygostomial angle rounded, slightly exceeding beyond antennal tooth.
</paragraph>
<paragraph id="8BA2EA25FFC3FFFE6BAE598FC0FAA7A3" blockId="1.[151,1436,1571,2027]" pageId="1" pageNumber="51">
Abdomen (
<figureCitation id="1326F6A0FFC3FFFE6A21598FC590A6A7" box="[328,398,1751,1775]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="1" pageNumber="51">Fig. 2</figureCitation>
C) with all pleura rounded; first to third pleura strongly expanded, enclosing large brood chamber. Sixth somite (
<figureCitation id="1326F6A0FFC3FFFE6ACA59A3C5FBA75B" box="[419,485,1787,1811]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="1" pageNumber="51">Fig. 3</figureCitation>
B) depressed dorsoventrally, widened posteriorly, subequal in length to fifth somite, 2.0 times wider than long; posteroventral angle triangular, subacutely pointed; posterolateral process triangular, sharply pointed; posterodorsal margin straight, unarmed. Telson (
<figureCitation id="1326F6A0FFC3FFFE681E581BC7A7A713" box="[887,953,1859,1883]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="1" pageNumber="51">Fig. 3</figureCitation>
C) about half length of carapace, 1.8 times as long as anterior width; lateral margins feebly convex, posteriorly convergent, with 2 pairs of small marginal dorsal spines (anterior pair located at 0.4 of telson length, posterior pair at 0.7), these spines subequal in length, about 0.05 of telson length; posterior margin convex, without median point, armed with 3 pairs of spines (lateral spines 0.06 of telson length, intermediate spines 0.10 and submedian spines 0.09 of telson length).
</paragraph>
<caption id="DF62BAADFFC0FFFD6BFE59AAC755A70E" httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="2" pageNumber="52" targetBox="[184,1406,158,1777]" targetPageId="2">
<paragraph id="8BA2EA25FFC0FFFD6BFE59AAC755A70E" blockId="2.[151,1436,1778,1862]" pageId="2" pageNumber="52">
<emphasis id="B9693637FFC0FFFD6BFE59AAC50AA740" bold="true" box="[151,276,1778,1801]" pageId="2" pageNumber="52">FIGURE 2.</emphasis>
<taxonomicName id="4C1D91A6FFC0FFFD6A7559ABC605A741" box="[284,539,1779,1801]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC0FFFD6A7559ABC605A741" box="[284,539,1779,1801]" italics="true" pageId="2" pageNumber="52">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC0FFFD694A59ABC640A740" box="[547,606,1779,1800]" pageId="2" pageNumber="52" rank="species">
<emphasis id="B9693637FFC0FFFD694A59ABC640A740" bold="true" box="[547,606,1779,1800]" pageId="2" pageNumber="52">n. sp.</emphasis>
</taxonomicNameLabel>
A–C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFC0FFFD6BAE5828C717A7C0" blockId="2.[151,1437,1904,2028]" box="[199,777,1904,1928]" pageId="2" pageNumber="52">Epistome with small rounded submedian swellings.</paragraph>
<paragraph id="8BA2EA25FFC0FFFD6BAE58CAC624A7A4" blockId="2.[151,1437,1904,2028]" pageId="2" pageNumber="52">
Eye (
<figureCitation id="1326F6A0FFC0FFFD6A6A58C9C558A7E1" box="[259,326,1937,1961]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="2" pageNumber="52">Fig. 3</figureCitation>
A) subquadrate in dorsal view, corneas hemispherical, lightly pigmented, without ocellar spot, its width about 0.1 of cl. Eyestalks swollen, narrowly separated; anterior surface flat, slightly sloping posteriorly, anteromesial angle weakly delimited.
</paragraph>
<caption id="DF62BAADFFC1FFFC6BFE5819C171A7B9" httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="3" pageNumber="53" targetBox="[226,1348,185,1831]" targetPageId="3">
<paragraph id="8BA2EA25FFC1FFFC6BFE5819C171A7B9" blockId="3.[151,1436,1857,2033]" pageId="3" pageNumber="53">
<emphasis id="B9693637FFC1FFFC6BFE5819C50BA71F" bold="true" box="[151,277,1857,1880]" pageId="3" pageNumber="53">FIGURE 3.</emphasis>
<taxonomicName id="4C1D91A6FFC1FFFC6A76581BC63EA710" box="[287,544,1858,1880]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="53" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC1FFFC6A76581BC63EA710" box="[287,544,1858,1880]" italics="true" pageId="3" pageNumber="53">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC1FFFC6940581AC67BA71F" box="[553,613,1858,1879]" pageId="3" pageNumber="53" rank="species">
<emphasis id="B9693637FFC1FFFC6940581AC67BA71F" bold="true" box="[553,613,1858,1879]" pageId="3" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A–E, H, K–N; 0.25 mm for I, J; 0.1 mm for F, G.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFC6FFFB6BAE5FCFC09DA1CF" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Antennular peduncle (
<figureCitation id="1326F6A0FFC6FFFB6AA95FCFC61FA0E7" box="[448,513,151,175]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
A) short, about 0.3 times as long as carapace. First segment about twice as long as wide; lateral and mesial margins sub-parallel; distolateral tooth reaching midlength of second segment; ventromesial margin without tooth; stylocerite reaching half segment length, acute, with several short plumose setae distally; statocyst poorly developed, with obsolescent statolith. Second and third segments subequal in length, combined length about half of first segment. Outer flagellum biramous, proximal 5 or 6 articles fused, bearing 5 or 6 groups of long aesthetascs; short free ramus consisting of single article, longer ramus short, consisting of 8–10 articles; inner flagellum slender, longer than outer flagellum, consisting of 10–12 articles.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5ECCC5AEA25F" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Antenna (
<figureCitation id="1326F6A0FFC6FFFB6A5E5ECCC565A1E4" box="[311,379,404,428]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
A, D) with stout basicerite unarmed. Carpocerite subcylindrical, 4.0 times longer than wide, slightly overreaching distal margin of antennal scale. Antennal scale well-developed, about 0.3 times as long as carapace, 1.7 times longer than wide; lateral margin straight, with strong distal tooth, far exceeding distal margin of broadly rounded lamella.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5D7CC626A274" blockId="4.[151,1437,151,1220]" box="[199,568,548,572]" pageId="4" pageNumber="54">Labrum without special features.</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5D1FC5B7A284" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Mandible (
<figureCitation id="1326F6A0FFC6FFFB6A285D1FC59CA217" box="[321,386,583,607]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
E) without palp. Molar process moderately slender, subcylindrical, distally obliquely truncate; distal surface with thick marginal row of setae on outer side and marginal row of spinules on inner side (
<figureCitation id="1326F6A0FFC6FFFB6E5C5D34C169A2CC" box="[1333,1399,620,644]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
G). Incisor process moderately broad, slightly curved, tapering to acute distal tooth, mesial edge armed with 4 minute subdistal teeth (
<figureCitation id="1326F6A0FFC6FFFB6A2E5DECC597A284" box="[327,393,692,716]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
F).
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5D8FC7EBA37F" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Maxillule (
<figureCitation id="1326F6A0FFC6FFFB6A2F5D8FC594A2A7" box="[326,394,727,751]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
H) of normal form; palp simple, stout, with shallow excavation terminally (
<figureCitation id="1326F6A0FFC6FFFB6F9D5D8FC127A2A7" box="[1268,1337,727,751]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
J). Basal endite moderately broad, rounded distal margin bearing row of several setulose setae and 2 spines (
<figureCitation id="1326F6A0FFC6FFFB6F9C5DA4C127A35C" box="[1269,1337,764,788]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">
Fig.
<date id="FFA3CCE5FFC6FFFB6E435DA4C127A35C" box="[1322,1337,764,788]" pageId="4" pageNumber="54">3</date>
</figureCitation>
I). Coxal endite slender, tapering distally, bearing tuft of stiff setulose setae terminally.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5C1CC609A337" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Maxilla (
<figureCitation id="1326F6A0FFC6FFFB6A5B5C1CC567A314" box="[306,377,836,860]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
K) with moderately slender tapering, non-setose palp. Endites damaged during dissection. Scaphognathite moderately broad.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5CD4C648A447" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
First maxilliped (
<figureCitation id="1326F6A0FFC6FFFB6AE55CD4C5D1A3EC" box="[396,463,908,932]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
L) with well-developed, non-setose palp, extending well beyond caridean lobe. Basal and coxal endites fused, distal margin broadly rounded, mesial margin straight, with sparse long setulose setae, coxal portion slightly convex. Exopod with slender flagellum with 4 plumose terminal setae; caridean lobe well developed, rounded. Epipod broken off.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE5B44C703A48C" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="54">
Second maxilliped (
<figureCitation id="1326F6A0FFC6FFFB6AC35B44C5F5A47C" box="[426,491,1052,1076]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="4" pageNumber="54">Fig. 3</figureCitation>
M) of normal form, consisting of 6 segments. Dactylus with mesial margin slightly concave, with numerous coarsely setulose spines; outer face with large patch of short setae. Propodus with anteromesial margin nearly straight, with several stiff finely setulose setae. Carpus and merus without special features. Ischiobasis widened proximally, slightly excavate mesially. Coxa with mesial margin rounded. Exopod long, with several plumose terminal setae. Epipod absent.
</paragraph>
<paragraph id="8BA2EA25FFC6FFFB6BAE59B1C765A791" blockId="4.[151,1437,1769,2009]" pageId="4" pageNumber="54">
Third maxilliped (
<figureCitation id="1326F6A0FFC6FFFB6AF059B1C5C1A749" box="[409,479,1769,1793]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="4" pageNumber="54">Fig. 4</figureCitation>
A, B) with endopod slender, extending to distal margin of antennal scale. Ultimate segment subequal to carpal length, slightly tapering distally. Penultimate segment (= carpus) 0.6 times as long as antepenultimate segment, subcylindrical, 2.0 times longer than wide. Antepenultimate segment 2.3 times as long as proximal width, tapering distally; lateral margin nearly straight, mesial margin slightly concave, with row of setae. Exopod well developed, distinctly overreaching distal end of antepenultimate segment, with some terminal plumose setae. Coxa large, with large, broadly rounded lateral plate; mesial margin convex, non-setose. Rudimentary arthrobranch present above base of third maxilliped.
</paragraph>
<caption id="DF62BAADFFC7FFFA6BFE5829C02CA7AC" httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="5" pageNumber="55" targetBox="[181,1406,180,1904]" targetPageId="5">
<paragraph id="8BA2EA25FFC7FFFA6BFE5829C02CA7AC" blockId="5.[151,1436,1905,2020]" pageId="5" pageNumber="55">
<emphasis id="B9693637FFC7FFFA6BFE5829C50CA7CF" bold="true" box="[151,274,1905,1928]" pageId="5" pageNumber="55">FIGURE 4.</emphasis>
<taxonomicName id="4C1D91A6FFC7FFFA6A71582BC608A7C0" box="[280,534,1906,1928]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="55" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC7FFFA6A71582BC608A7C0" box="[280,534,1906,1928]" italics="true" pageId="5" pageNumber="55">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC7FFFA6972582AC64AA7CF" box="[539,596,1906,1927]" pageId="5" pageNumber="55" rank="species">
<emphasis id="B9693637FFC7FFFA6972582AC64AA7CF" bold="true" box="[539,596,1906,1927]" pageId="5" pageNumber="55">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A–C, E, G, H; 0.25 mm for D; 0.1 mm for F.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFC4FFF96BAE5FCFC721A30A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
First pereopod (
<figureCitation id="1326F6A0FFC4FFF96A145FCFC5DDA0E7" box="[381,451,151,175]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig. 4</figureCitation>
C) slender, overreaching distal margin of antennal scale by distal 0.2 of merus. Chela (
<figureCitation id="1326F6A0FFC4FFF96BF65FE2C4FCA09A" box="[159,226,186,210]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig. 4</figureCitation>
D) subcylindrical, slightly tapering distally, 0.6 times as long as carpus, 5.5 times as long as greatest width, with sparse grooming setae proximoventrally; dactylus about 0.3 of palm length; fingers distally with curved setae forming basket-like structure. Carpus 8.0–9.0 times as long as distal width, tapering slightly proximally. Merus slightly curved, subequal in length to carpus, 6.0–7.0 times as long as greatest width. Ischium 0.7 of carpal length. Second pereopods slightly unequal, similar. Major second pereopod (
<figureCitation id="1326F6A0FFC4FFF968DD5E1DC7E7A115" box="[948,1017,325,349]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
A, B) stout. Chela moderately large, about 0.8 times as long as carapace, 2.3 times longer than greatest width, ventral margin gently sinuous; palm compressed, smooth, about 1.4 times as long as greatest width, slightly projecting posterodorsally. Dactylus about 0.7 of palm length, 3.2 times as long as proximal width, scaphoid with strongly hooked acute tip; lateral surface with scattered tufts of short setae; mesial surface with 3 rows of tufts of short to long setae; cutting edge chitinous with proximal one-third slightly concave, with 1 or 2 small triangular teeth, distal two-thirds chitinous, serrate with row of sharp (acute?) teeth increasing in size distally (
<figureCitation id="1326F6A0FFC4FFF968475D4DC76BA265" box="[814,885,533,557]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
C). Fixed finger tapering, deeply bifid distally, longitudinally grooved (
<figureCitation id="1326F6A0FFC4FFF96ACE5D6FC5F4A207" box="[423,490,567,591]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
D); lateral surface with scattered tufts of short setae; mesial surface with irregular 2 rows of tufts of short setae along mesial cutting edge; lateral cutting edge with 1 or 2 triangular teeth on proximal two-fifths, distal three-fifths forming chitinous flange with slightly convex, microscopically serrate margin (teeth increasing in size distally) (
<figureCitation id="1326F6A0FFC4FFF96AA65DC7C60DA2FF" box="[463,531,671,695]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
C); mesial edge mostly entire, but with some minute acute denticles on strongly curved, acute distal part (
<figureCitation id="1326F6A0FFC4FFF96AAE5D9AC610A292" box="[455,526,706,730]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
C). Carpus cup-shaped, about 0.3 of chela length, smooth; distal part deeply excavate to accommodate posterior extension of chela. Merus 1.3–1.4 times longer than carpus, centrally swollen, 2.0–2.2 times as long as wide, distoventrally excavate; ventral margin smooth. Ischium about half of meral length, slightly tapered proximally, unarmed. Basis and coxa robust.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5C15C034A3C0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
Minor second pereopod (
<figureCitation id="1326F6A0FFC4FFF96A885C15C63CA32D" box="[481,546,845,869]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
E–G) with palm about 1.5 times longer than greatest width; proximal one-third of cutting edge of fixed finger irregularly denticulate to bearing up to 5 tiny teeth.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5CCAC0EBA4D5" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
Third pereopod (
<figureCitation id="1326F6A0FFC4FFF96AE25CCAC5D8A3E2" box="[395,454,914,938]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig.4</figureCitation>
E) moderately robust, overreaching distal margin of antennal scale by half length of propodus. Dactylus (
<figureCitation id="1326F6A0FFC4FFF96AE25CEDC5CDA385" box="[395,467,949,973]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig. 4</figureCitation>
F) strongly compressed laterally, 2.0–2.1 times longer than wide, extensor margin convex; unguis curved, without ventral denticulations; flexor margin nearly straight, armed with large, faintly bicuspidate, distal accessory tooth followed by row of 5 acute, obliquely erect denticles decreasing in size proximally in distal 0.6. Propodus about 4.3 times as long as greatest width, slightly tapering distally, slightly compressed laterally, sparsely setose, with single long flexor distal spine and 4 widely spaced, small flexor spines. Carpus 0.7 times as long as propodus, slightly widened distally. Merus about 3.0 times longer than greatest width, unarmed. Ischium 0.8 times as long as merus, unarmed. Basis and coxa without distinctive features.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BFE5BF0C66BA488" blockId="6.[151,1437,151,2013]" box="[151,629,1192,1216]" pageId="6" pageNumber="56">
Fourth pereopod (
<figureCitation id="1326F6A0FFC4FFF96A095BF0C5BCA488" box="[352,418,1192,1216]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig. 4</figureCitation>
G) similar to third.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5B92C182A54D" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
Fifth pereopod (
<figureCitation id="1326F6A0FFC4FFF96A125B92C5A0A4AA" box="[379,446,1226,1250]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="6" pageNumber="56">Fig. 4</figureCitation>
H) generally similar to third. Propodus about 5.3 times longer than wide, with cluster of submarginal grooming setae on distal flexor part, distal flexor margin with single spine, flexor margin otherwise unarmed.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5A48C641A560" blockId="6.[151,1437,151,2013]" box="[199,607,1296,1320]" pageId="6" pageNumber="56">Gill formula summarized in Table 1.</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5A6AC6B5A59D" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
Pleopods without distinctive features. Uropod (
<figureCitation id="1326F6A0FFC4FFF9698B5A6AC73BA502" box="[738,805,1330,1354]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
N) with protopodite unarmed, posterolaterally rounded. Endopod longer than telson, about 2.3 times longer than wide; lateral margin gently convex; posterior margin rounded. Exopod slightly shorter than endopod, slightly widened posteriorly; lateral margin slightly convex, sparsely setose, terminating in small acute tooth; spinule present just mesial to base of posterolateral tooth; posterior margin broadly rounded; no diaeresis.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5AB9C6C5A5B0" blockId="6.[151,1437,151,2013]" box="[199,731,1504,1529]" pageId="6" pageNumber="56">
Eggs numerous (
<figureCitation id="1326F6A0FFC4FFF96AED5AB8C5DBA5B0" box="[388,453,1504,1528]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="6" pageNumber="56">Fig. 1</figureCitation>
), 1.0–1.2 x 1.0–
<quantity id="4CE547C0FFC4FFF969165AB8C6C6A5B0" box="[639,728,1504,1528]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="56" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE595BC590A628" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
<emphasis id="B9693637FFC4FFF96BAE595BC515A653" box="[199,267,1539,1563]" italics="true" pageId="6" pageNumber="56">Males</emphasis>
. Body less stout than in females (
<figureCitation id="1326F6A0FFC4FFF969F0595AC6C1A652" box="[665,735,1538,1562]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
D, E). Carapace (
<figureCitation id="1326F6A0FFC4FFF968C1595AC7F3A652" box="[936,1005,1538,1562]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
D) with dorsal margin in lateral view nearly straight; pterygostomial angle more narrowly pointed. Abdominal pleura not expanded; sixth somite less compressed (
<figureCitation id="1326F6A0FFC4FFF96A435910C570A628" box="[298,366,1608,1632]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
E).
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE5932C7E2A6ED" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="56">
Cornea (
<figureCitation id="1326F6A0FFC4FFF96A4E5932C574A6CA" box="[295,362,1642,1666]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
D) relatively larger, its diameter about 0.2 of carapace length. Antennal scale about 0.4 times as long as carapace; carpocerite slightly falling short of distal margin of lamella.
</paragraph>
<paragraph id="8BA2EA25FFC4FFF96BAE59E8C782A680" blockId="6.[151,1437,151,2013]" box="[199,924,1712,1736]" pageId="6" pageNumber="56">First, third to fifth pereopods generally similar to those of males.</paragraph>
<paragraph id="8BA2EA25FFC4FFF66BAE598AC7C4A15F" blockId="6.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,2013]" lastPageId="9" lastPageNumber="59" pageId="6" pageNumber="56">
Second pereopods markedly unequal and dissimilar. Major second pereopod (
<figureCitation id="1326F6A0FFC4FFF96F41598AC074A6A2" box="[1064,1130,1746,1770]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="6" pageNumber="56">Fig. 6</figureCitation>
A, B) massive. Chela about 1.8 times longer than carapace, about 2.0 times longer than greatest width, ventral margin sinuous; palm strongly compressed, about 1.3 times as long as greatest width, slightly projecting posterodorsally; mesial surface with shallow excavation proximally; ventral margin faintly tuberculate. Dactylus about 0.6 of palm length, terminally in-curved, tip acute; lateral surface with row of tufts of short setae along cutting edge located slightly laterally, with 2 prominent triangular teeth (proximal tooth subacute, distal tooth blunt, much lower than proximal tooth). Fixed finger feebly deflexed, terminating in hooked tip; lateral surface with row of tufts of short setae along cutting edge; mesial surface with scattered tufts of short setae; cutting edge with 3 prominent teeth (proximal and distal teeth triangular, middle tooth subrectangular). Carpus short, cup-shaped, smooth; distodorsal part deeply excavate to accommodate posterior extension of chela. Merus 1.6 times longer than carpus, centrally slightly swollen, 2.2–2.3 times longer than wide, distoventrally excavate; ventral margin with few minute tubercles. Ischium about 0.6 of meral length, slightly tapered proximally, unarmed. Basis and coxa robust.
</paragraph>
<caption id="DF62BAADFFC5FFF86BFE5800C5B7A7A1" httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="7" pageNumber="57" targetBox="[183,1388,182,1879]" targetPageId="7">
<paragraph id="8BA2EA25FFC5FFF86BFE5800C5B7A7A1" blockId="7.[151,1437,1880,2025]" pageId="7" pageNumber="57">
<emphasis id="B9693637FFC5FFF86BFE5800C50DA726" bold="true" box="[151,275,1880,1903]" pageId="7" pageNumber="57">FIGURE 5.</emphasis>
<taxonomicName id="4C1D91A6FFC5FFF86A735801C607A726" box="[282,537,1880,1902]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC5FFF86A735801C607A726" box="[282,537,1880,1902]" italics="true" pageId="7" pageNumber="57">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC5FFF869495801C644A726" box="[544,602,1881,1902]" pageId="7" pageNumber="57" rank="species">
<emphasis id="B9693637FFC5FFF869495801C644A726" bold="true" box="[544,602,1881,1902]" pageId="7" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D–F; 0.25 mm for C.
</paragraph>
</caption>
<caption id="DF62BAADFFCAFFF76BFE5835C650A7A8" httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="8" pageNumber="58" targetBox="[211,1383,217,1854]" targetPageId="8">
<paragraph id="8BA2EA25FFCAFFF76BFE5835C650A7A8" blockId="8.[151,1436,1901,2016]" pageId="8" pageNumber="58">
<emphasis id="B9693637FFCAFFF76BFE5835C50AA7CB" bold="true" box="[151,276,1901,1924]" pageId="8" pageNumber="58">FIGURE 6.</emphasis>
<taxonomicName id="4C1D91A6FFCAFFF76A755837C602A7CC" box="[284,540,1902,1924]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="58" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFCAFFF76A755837C602A7CC" box="[284,540,1902,1924]" italics="true" pageId="8" pageNumber="58">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFCAFFF7694D5836C67EA7CB" box="[548,608,1902,1923]" pageId="8" pageNumber="58" rank="species">
<emphasis id="B9693637FFCAFFF7694D5836C67EA7CB" bold="true" box="[548,608,1902,1923]" pageId="8" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A–D; 0.5 mm for E; 0.25 mm for I.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFCBFFF66BAE5E7AC7E2A172" blockId="9.[151,1436,151,2013]" box="[199,1020,290,314]" pageId="9" pageNumber="59">
Minor second pereopod (
<figureCitation id="1326F6A0FFCBFFF66A895E7AC63DA172" box="[480,547,290,314]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="9" pageNumber="59">Fig. 6</figureCitation>
C, D) similar to female second pereopods.
</paragraph>
<paragraph id="8BA2EA25FFCBFFF66BAE5E1DC7E2A1EA" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
First pleopod (
<figureCitation id="1326F6A0FFCBFFF66A195E1DC5ABA115" box="[368,437,325,349]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="9" pageNumber="59">Fig. 6</figureCitation>
E) with small, slender endopod about 0.4 times as long as exopod, tapering distally to rounded tip, sparsely setose. Endopod of second pleopod with appendix masculina greatly reduced to socket for single long slender setulose seta located at base of appendix interna (
<figureCitation id="1326F6A0FFCBFFF668F35ED2C7C3A1EA" box="[922,989,394,418]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="9" pageNumber="59">Fig. 6</figureCitation>
F).
</paragraph>
<paragraph id="8BA2EA25FFCBFFF66BAE5EF5C5C2A242" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
<emphasis id="B9693637FFCBFFF66BAE5EF5C58DA18E" bold="true" box="[199,403,429,454]" pageId="9" pageNumber="59">Coloration in life</emphasis>
. Body and appendages translucent. Carapace with scattered red chromatophores. Abdomen with numerous scattered white chromatophores and less numerous red chromatophores. Ovary green. Eggs pale greenish gray. See
<figureCitation id="1326F6A0FFCBFFF66A035EAAC5A8A242" box="[362,438,498,522]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="9" pageNumber="59">Figs. 1</figureCitation>
,
<figureCitation id="1326F6A0FFCBFFF66AAD5EAAC5CDA242" box="[452,467,498,522]" captionStart="FIGURE 7" captionStartId="10.[151,250,1941,1964]" captionTargetBox="[171,1414,317,1922]" captionTargetId="figure@10.[171,1415,317,1922]" captionTargetPageId="10" captionText="FIGURE 7. Underwater photographs. A, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm) (indicated by red arrow), and host sponge Haliclona sp.; B, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm); C, Onycocaris hayamaensis n. sp., male, spouse of holotype, not collected." httpUri="https://zenodo.org/record/214127/files/figure.png" pageId="9" pageNumber="59">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFCBFFF66BAE5D4DC11AA2FF" pageId="9" pageNumber="59" type="description">
<paragraph id="8BA2EA25FFCBFFF66BAE5D4DC11AA2FF" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
<emphasis id="B9693637FFCBFFF66BAE5D4DC52BA266" bold="true" box="[199,309,533,558]" pageId="9" pageNumber="59">Variation</emphasis>
.
<collectingCountry id="F30AAAB5FFCBFFF66A235D4EC577A266" box="[330,361,534,558]" name="American Samoa" pageId="9" pageNumber="59">As</collectingCountry>
is apparent from the above description, the size and structure of the chelae of the second pereopods exhibit strong sexual dimorphism in this new species. In males, the second pereopods are greatly unequal in the size and dissimilar in the structure and armature of the fingers. In contrast, in females, the second pereopods are slightly unequal and similar, and generally similar to the minor second pereopod of males. In addition, the body is plump in females, with strongly expanded pleura forming broad brood chamber.
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFCBFFF66BAE5D9AC660A30B" pageId="9" pageNumber="59" type="distribution">
<paragraph id="8BA2EA25FFCBFFF66BAE5D9AC0B2A292" blockId="9.[151,1436,151,2013]" box="[199,1196,706,731]" pageId="9" pageNumber="59">
<emphasis id="B9693637FFCBFFF66BAE5D9AC547A293" bold="true" box="[199,345,706,731]" pageId="9" pageNumber="59">Distribution</emphasis>
. Presently known only from Sagami Bay, central
<collectingCountry id="F30AAAB5FFCBFFF668EE5D9BC7D4A292" box="[903,970,707,730]" name="Japan" pageId="9" pageNumber="59">Japan</collectingCountry>
, at depths of
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5–
<specimenCount id="9D1B21ACFFCBFFF66F125D9AC0B6A292" box="[1147,1192,706,730]" pageId="9" pageNumber="59" type="generic">6 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8BA2EA25FFCBFFF66BAE5DBDC660A30B" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
<emphasis id="B9693637FFCBFFF66BAE5DBDC53AA2B5" bold="true" box="[199,292,741,765]" pageId="9" pageNumber="59">Ecology</emphasis>
. Specimens were all found in association with
<taxonomicName id="4C1D91A6FFCBFFF668225DBDC7A5A2B5" box="[843,955,741,765]" class="Demospongiae" family="Chalinidae" genus="Haliclona" kingdom="Animalia" order="Haplosclerida" pageId="9" pageNumber="59" phylum="Porifera" rank="genus">
<emphasis id="B9693637FFCBFFF668225DBDC7A5A2B5" box="[843,955,741,765]" italics="true" pageId="9" pageNumber="59">Haliclona</emphasis>
</taxonomicName>
sp., encrusted on bottom surfaces of free rocks (about
<quantity id="4CE547C0FFCBFFF66A415C50C589A368" box="[296,407,776,800]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="59" unit="cm" value="65.0" valueMax="70.0" valueMin="60.0">60–70 cm</quantity>
in longer axis) (
<figureCitation id="1326F6A0FFCBFFF669245C50C691A368" box="[589,655,776,800]" captionStart="FIGURE 7" captionStartId="10.[151,250,1941,1964]" captionTargetBox="[171,1414,317,1922]" captionTargetId="figure@10.[171,1415,317,1922]" captionTargetPageId="10" captionText="FIGURE 7. Underwater photographs. A, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm) (indicated by red arrow), and host sponge Haliclona sp.; B, Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm); C, Onycocaris hayamaensis n. sp., male, spouse of holotype, not collected." httpUri="https://zenodo.org/record/214127/files/figure.png" pageId="9" pageNumber="59">Fig. 7</figureCitation>
A). The animals lived in small cavities of the sponge in a heterosexual pair. Females were all ovigerous in August.
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFCBFFF66BAE5C15C001A680" pageId="9" pageNumber="59" type="discussion">
<paragraph id="8BA2EA25FFCBFFF66BAE5C15C066A488" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
<emphasis id="B9693637FFCBFFF66BAE5C15C52AA32D" bold="true" box="[199,308,845,869]" pageId="9" pageNumber="59">Remarks</emphasis>
.
<taxonomicName id="4C1D91A6FFCBFFF66A2D5C16C640A32D" box="[324,606,845,869]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFCBFFF66A2D5C16C640A32D" box="[324,606,845,869]" italics="true" pageId="9" pageNumber="59">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFCBFFF6690E5C15C6B9A32D" box="[615,679,845,869]" pageId="9" pageNumber="59" rank="species">
<emphasis id="B9693637FFCBFFF6690E5C15C6B9A32D" bold="true" box="[615,679,845,869]" pageId="9" pageNumber="59">n. sp.</emphasis>
</taxonomicNameLabel>
is referable to the
<taxonomicName id="4C1D91A6FFCBFFF668EB5C16C068A32D" box="[898,1142,845,869]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFCBFFF668EB5C16C068A32D" box="[898,1142,845,869]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF668EB5C16C787A32D" box="[898,921,846,869]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. quadratophthalma
</emphasis>
</taxonomicName>
species group because of the following characters: (1) rostrum short, unarmed; (2) chelae of second pereopods lacking mesial row of prominent teeth on fingers: and (3) meri and ischia of second pereopods unarmed. This informal species group contains the following four species:
<taxonomicName id="4C1D91A6FFCBFFF669455CEEC703A385" authority="Bruce, 2011" authorityName="Bruce" authorityYear="2011" box="[556,797,949,973]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF669455CEEC690A385" box="[556,654,949,973]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF669455CEEC65DA385" box="[556,579,950,973]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
Bruce, 2011
</taxonomicName>
from the Northern Territory,
<collectingCountry id="F30AAAB5FFCBFFF66F0D5CEDC0D5A385" box="[1124,1227,949,973]" name="Australia" pageId="9" pageNumber="59">Australia</collectingCountry>
(
<bibRefCitation id="EF8C97D4FFCBFFF66FB05CEDC170A385" author="Bruce" box="[1241,1390,949,973]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce 2011b</bibRefCitation>
);
<taxonomicName id="4C1D91A6FFCBFFF66EE85CEEC56DA3B8" authority="Bruce, 2011" authorityName="Bruce" authorityYear="2011" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFCBFFF66EE85CEEC4FEA3B8" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66EE85CEEC186A385" box="[1409,1432,950,973]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. fujinoi
</emphasis>
Bruce, 2011
</taxonomicName>
, from the Ryukyu Islands,
<collectingCountry id="F30AAAB5FFCBFFF669CB5C81C6FCA3B8" box="[674,738,985,1008]" name="Japan" pageId="9" pageNumber="59">Japan</collectingCountry>
(
<bibRefCitation id="EF8C97D4FFCBFFF669985C80C799A3B8" author="Bruce" box="[753,903,984,1008]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce 2011b</bibRefCitation>
);
<taxonomicName id="4C1D91A6FFCBFFF668F35C80C093A3B8" box="[922,1165,984,1008]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFCBFFF668F35C80C093A3B8" box="[922,1165,984,1008]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF668F35C80C7AFA3A7" box="[922,945,984,1007]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. quadratophthalma
</emphasis>
</taxonomicName>
from Western
<collectingCountry id="F30AAAB5FFCBFFF66E5C5C80C182A3B8" box="[1333,1436,984,1008]" name="Australia" pageId="9" pageNumber="59">Australia</collectingCountry>
and
<collectingCountry id="F30AAAB5FFCBFFF66BAF5CA3C554A45B" box="[198,330,1019,1043]" name="Hong Kong" pageId="9" pageNumber="59">Hong Kong</collectingCountry>
(
<bibRefCitation id="EF8C97D4FFCBFFF66A305CA2C5FEA45A" author="Balss" box="[345,480,1018,1042]" pageId="9" pageNumber="59" refString="Balss, H. (1921) Results of Dr. E. Mjobergs Swedish Scientific Expeditions to Australia 1910 - 13. XXIX: Stomatopoda, Macrura, Paguridea und Galatheidea. Kungliga Svenska Vetenskapsakademiens Handlinger, 61, 1 - 24." type="journal article" year="1921">Balss, 1921</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFCBFFF66A855CA3C66FA45A" author="Bruce" box="[492,625,1018,1042]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">Bruce 1992</bibRefCitation>
,
<bibRefCitation id="EF8C97D4FFCBFFF669155CA2C6D8A45A" author="Bruce" box="[636,710,1018,1042]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">2011b</bibRefCitation>
); and
<taxonomicName id="4C1D91A6FFCBFFF668605CA3C011A45A" authority="Bruce, 1978" authorityName="Bruce" authorityYear="1978" box="[777,1039,1018,1043]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF668605CA3C761A45B" box="[777,895,1019,1043]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF668605CA3C73EA45A" box="[777,800,1019,1042]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
<bibRefCitation id="EF8C97D4FFCBFFF668EF5CA3C011A45A" author="Bruce" box="[902,1039,1018,1042]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce, 1978</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F30AAAB5FFCBFFF66F3A5CA2C0C4A45A" box="[1107,1242,1018,1042]" name="Madagascar" pageId="9" pageNumber="59">Madagascar</collectingCountry>
(
<bibRefCitation id="EF8C97D4FFCBFFF66F805CA2C16FA45A" author="Bruce" box="[1257,1393,1018,1042]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978</bibRefCitation>
). In previous works (e.g.,
<bibRefCitation id="EF8C97D4FFCBFFF66AE35B45C691A47D" author="Fujino" box="[394,655,1053,1077]" pageId="9" pageNumber="59" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFCBFFF669F25B45C727A47D" author="Hayashi" box="[667,825,1053,1077]" pageId="9" pageNumber="59" refString="Hayashi, K. (2003) Shrimps and lobsters from Japan (127). Family Palaemonidae, subfamily Pontoniinae - Genus Onycocaris. Aquabiology, 144, 21 - 26." type="journal article" year="2003">Hayashi 2003</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFCBFFF6682C5B45C7D5A47D" author="Marin" box="[837,971,1053,1077]" pageId="9" pageNumber="59" refString="Marin, I. (2005) Pontoniine shrimps (Crustacea, Decapoda, Pontoniinae) from Viet Nam. Onycocaris temiri sp. n., a new sponge-associated shrimp from Nha Trang Bay. Arthropoda Selecta, 13, 113 - 122." type="journal article" year="2005">Marin 2005</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFCBFFF668BE5B46C079A47D" author="Bruce" box="[983,1127,1053,1077]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 a) Notes on some Indo-Pacific Pontoniinae, XLVIII. Onycocaris nieli sp. nov., a new pontoniine shrimp from Heron Island, Queensland. Crustaceana, 84, 319 - 330." type="journal article" year="2011" yearSuffix="a">Bruce 2011a</bibRefCitation>
,
<bibRefCitation id="EF8C97D4FFCBFFF66F1A5B45C0A2A47D" author="Bruce" box="[1139,1212,1053,1077]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">2011b</bibRefCitation>
), characters derived from chelae of the second pereopods have been routinely used in differentiating species of
<taxonomicName id="4C1D91A6FFCBFFF66FD25B18C122A41F" box="[1211,1340,1088,1111]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B9693637FFCBFFF66FD25B18C122A41F" box="[1211,1340,1088,1111]" italics="true" pageId="9" pageNumber="59">Onycocaris</emphasis>
</taxonomicName>
without careful consideration on the sexual dimorphism and heteromorph between right and left. The recent report on
<taxonomicName id="4C1D91A6FFCBFFF66EE85B3BC5ADA4D5" authority="Bruce, 1980" authorityName="Bruce" authorityYear="1980" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B9693637FFCBFFF66EE85B3BC507A4D5" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66EE85B3BC186A432" box="[1409,1432,1123,1146]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. longirostris
</emphasis>
<bibRefCitation id="EF8C97D4FFCBFFF66A4C5BDEC5ADA4D5" author="Bruce" box="[293,435,1157,1181]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1980) On some pontoniine shrimps from Noumea, New Caledonia. Cahiers de l'Indo-Pacifique, 2, 1 - 9, figs 1 - 14." type="journal article" year="1980">Bruce, 1980</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EF8C97D4FFCBFFF66A815BDEC69CA4D5" author="Bruce" box="[488,642,1157,1181]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2012) Onycocaris longirostris Bruce, 1980 (Crustacea: Decapoda: Pontoniinae), new to the Australian fauna. Zootaxa, 3299, 61 - 65." type="journal article" year="2012">Bruce (2012)</bibRefCitation>
and the discovery of the present new species led us to assess the characters of the second pereopods more carefully with special attention to the variation.
</paragraph>
<paragraph id="8BA2EA25FFCBFFF66BAE5B93C001A680" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="59">
Among the four species,
<taxonomicName id="4C1D91A6FFCBFFF66A8B5B93C658A4AB" box="[482,582,1227,1251]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66A8B5B93C658A4AB" box="[482,582,1227,1251]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66A8B5B93C5E7A4AA" box="[482,505,1227,1250]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
appears closest to
<taxonomicName id="4C1D91A6FFCBFFF668755B93C7D1A4AB" box="[796,975,1227,1251]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFCBFFF668755B93C7D1A4AB" box="[796,975,1227,1251]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF668755B93C72DA4AA" box="[796,819,1227,1250]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFCBFFF668BE5B93C008A4AB" box="[983,1046,1227,1251]" pageId="9" pageNumber="59" rank="species">
<emphasis id="B9693637FFCBFFF668BE5B93C008A4AB" bold="true" box="[983,1046,1227,1251]" pageId="9" pageNumber="59">n. sp.</emphasis>
</taxonomicNameLabel>
in the fixed fingers of both chelae of the second pereopods with deeply bifid tips and well-developed lateral flanges and the proportionally short fingers of the first pereopod (less than 0.3 times as long as the palm) (
<bibRefCitation id="EF8C97D4FFCBFFF668C05A48C021A560" author="Bruce" box="[937,1087,1296,1320]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce 2011b</bibRefCitation>
).
<collectingCountry id="F30AAAB5FFCBFFF66F385A48C06EA560" box="[1105,1136,1296,1320]" name="American Samoa" pageId="9" pageNumber="59">As</collectingCountry>
only the ovigerous female
<typeStatus id="54A65487FFCBFFF66BFE5A6AC4E7A502" box="[151,249,1330,1354]" pageId="9" pageNumber="59" type="holotype">holotype</typeStatus>
is known for
<taxonomicName id="4C1D91A6FFCBFFF66AFD5A6BC5E9A503" box="[404,503,1331,1355]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66AFD5A6BC5E9A503" box="[404,503,1331,1355]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66AFD5A6BC5B5A502" box="[404,427,1331,1354]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
, the following comparison is limited to females. Fortunately, the second pereopods of the
<typeStatus id="54A65487FFCBFFF66BB75A0DC55EA525" box="[222,320,1365,1389]" pageId="9" pageNumber="59" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1D91A6FFCBFFF66A0D5A0EC5D6A525" box="[356,456,1365,1389]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66A0D5A0EC5D6A525" box="[356,456,1365,1389]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66A0D5A0EC565A525" box="[356,379,1366,1389]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
are both preserved (
<bibRefCitation id="EF8C97D4FFCBFFF669C25A0DC75EA525" author="Bruce" box="[683,832,1365,1389]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce 2011b</bibRefCitation>
). The new species can be distinguished from
<taxonomicName id="4C1D91A6FFCBFFF66E505A0EC182A525" box="[1337,1436,1365,1389]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66E505A0EC182A525" box="[1337,1436,1365,1389]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66E505A0EC14EA525" box="[1337,1360,1366,1389]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
by the different armature of the cutting edges of the chela of the second pereopods. In
<taxonomicName id="4C1D91A6FFCBFFF66F0A5A20C109A5D8" box="[1123,1303,1400,1424]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF66F0A5A20C109A5D8" box="[1123,1303,1400,1424]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66F0A5A20C064A5C7" box="[1123,1146,1400,1423]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
, the cutting edges of dactyli and lateral flanges of the fixed finger are serrated with fine teeth increasing in size distally (
<figureCitation id="1326F6A0FFCBFFF66E075AC2C4BBA59D" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="9" pageNumber="59">Fig. 5</figureCitation>
C). In contrast, in
<taxonomicName id="4C1D91A6FFCBFFF66A125AE6C5FFA59D" box="[379,481,1469,1493]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66A125AE6C5FFA59D" box="[379,481,1469,1493]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66A125AE6C58CA59D" box="[379,402,1470,1493]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
, the cutting edges of the fingers are minutely serrate with denticles of similar size (
<bibRefCitation id="EF8C97D4FFCBFFF66BF75AB8C5ACA5B0" author="Bruce" box="[158,434,1504,1528]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">
Bruce 2011b:
<figureCitation id="1326F6A0FFCBFFF66A545AB8C59EA5B0" box="[317,384,1504,1528]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="9" pageNumber="59">Fig. 4</figureCitation>
C, F
</bibRefCitation>
). Furthermore, the dorsal margin of the carapace is more strongly sloping anteriorly in
<taxonomicName id="4C1D91A6FFCBFFF66EE85AB8C536A653" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF66EE85AB8C536A653" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66EE85AB8C186A5BF" box="[1409,1432,1504,1527]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D91A6FFCBFFF66AF9595BC5E6A653" box="[400,504,1539,1563]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66AF9595BC5E6A653" box="[400,504,1539,1563]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66AF9595BC5B9A652" box="[400,423,1539,1562]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFCBFFF66965595AC64CA652" box="[524,594,1538,1562]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="9" pageNumber="59">Fig. 2</figureCitation>
A versus
<bibRefCitation id="EF8C97D4FFCBFFF669AF595BC740A652" author="Bruce" box="[710,862,1538,1562]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce 2011b</bibRefCitation>
:
<figureCitation id="1326F6A0FFCBFFF66804595AC7B1A652" box="[877,943,1538,1562]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="9" pageNumber="59">Fig. 1</figureCitation>
A); the posterolateral process of the sixth abdominal somite is acutely pointed in
<taxonomicName id="4C1D91A6FFCBFFF6693C597EC717A675" box="[597,777,1573,1597]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF6693C597EC717A675" box="[597,777,1573,1597]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF6693C597EC672A675" box="[597,620,1574,1597]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFCBFFF66870597DC742A675" box="[793,860,1573,1597]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="9" pageNumber="59">Fig. 3</figureCitation>
B), rather than blunt in
<taxonomicName id="4C1D91A6FFCBFFF66F01597EC0D2A675" box="[1128,1228,1573,1597]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
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<collectingCountry id="F30AAAB5FFCBFFF66F01597EC061A675" box="[1128,1151,1574,1597]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
(Bruce 2011:
<figureCitation id="1326F6A0FFCBFFF66E06597DC4BDA628" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="9" pageNumber="59">
Fig.
<date id="FFA3CCE5FFCBFFF66BFE5910C4BDA628" box="[151,163,1608,1632]" pageId="9" pageNumber="59">1</date>
</figureCitation>
I); the lateral margin of the uropod is nearly straight in the posterior three-fourths in
<taxonomicName id="4C1D91A6FFCBFFF66F075910C13DA628" box="[1134,1315,1608,1632]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF66F075910C13DA628" box="[1134,1315,1608,1632]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66F075910C09BA617" box="[1134,1157,1608,1631]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFCBFFF66E5D5910C168A628" box="[1332,1398,1608,1632]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="9" pageNumber="59">Fig. 3</figureCitation>
N), whereas entirely convex in
<taxonomicName id="4C1D91A6FFCBFFF66ABF5933C622A6CB" box="[470,572,1643,1667]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF66ABF5933C622A6CB" box="[470,572,1643,1667]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66ABF5933C5F3A6CA" box="[470,493,1643,1666]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFCBFFF669275932C750A6CA" author="Bruce" box="[590,846,1642,1666]" pageId="9" pageNumber="59" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">
Bruce 2011b:
<figureCitation id="1326F6A0FFCBFFF6699A5932C72AA6CA" box="[755,820,1642,1666]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="9" pageNumber="59">Fig. 1</figureCitation>
K
</bibRefCitation>
); finally the incisor process of the mandible tapers distally and bears three or four minute subdistal teeth in
<taxonomicName id="4C1D91A6FFCBFFF6687A59D6C7D8A6ED" box="[787,966,1677,1701]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF6687A59D6C7D8A6ED" box="[787,966,1677,1701]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF6687A59D6C734A6ED" box="[787,810,1678,1701]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFCBFFF668BF59D5C004A6ED" box="[982,1050,1677,1701]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="9" pageNumber="59">Fig. 4</figureCitation>
F), whereas it is slightly expanded and is serrated with about 30 closely set teeth in
<taxonomicName id="4C1D91A6FFCBFFF669DE59E8C704A680" box="[695,794,1712,1736]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9693637FFCBFFF669DE59E8C704A680" box="[695,794,1712,1736]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF669DE59E8C6D0A68F" box="[695,718,1712,1735]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. balssi
</emphasis>
</taxonomicName>
(Bruce 2011:
<figureCitation id="1326F6A0FFCBFFF668D259E8C7E3A680" box="[955,1021,1712,1736]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="9" pageNumber="59">Fig. 5</figureCitation>
C).
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFCBFFF46BAE598BC148A5AD" lastPageId="11" lastPageNumber="61" pageId="9" pageNumber="59" type="reference_group">
<paragraph id="8BA2EA25FFCBFFF56BAE598BC188A15F" blockId="9.[151,1436,151,2013]" lastBlockId="10.[151,1436,151,280]" lastPageId="10" lastPageNumber="60" pageId="9" pageNumber="59">
<taxonomicName id="4C1D91A6FFCBFFF66BAE598BC5BFA6A3" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[199,417,1747,1771]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF66BAE598BC5BFA6A3" box="[199,417,1747,1771]" italics="true" pageId="9" pageNumber="59">Onycocaris trullata</emphasis>
</taxonomicName>
is also represented only by a single ovigerous female, of which the second pereopods are intact (
<bibRefCitation id="EF8C97D4FFCBFFF66B8B59ADC579A745" author="Bruce" box="[226,359,1781,1805]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978</bibRefCitation>
). It is readily distinguished from
<taxonomicName id="4C1D91A6FFCBFFF669BA59AEC79FA745" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[723,897,1781,1805]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFCBFFF669BA59AEC79FA745" box="[723,897,1781,1805]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF669BA59AEC6F4A745" box="[723,746,1782,1805]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFCBFFF668E059ADC7D9A745" box="[905,967,1781,1805]" pageId="9" pageNumber="59" rank="species">
<emphasis id="B9693637FFCBFFF668E059ADC7D9A745" bold="true" box="[905,967,1781,1805]" pageId="9" pageNumber="59">n. sp.</emphasis>
</taxonomicNameLabel>
by the following female characters (
<bibRefCitation id="EF8C97D4FFCBFFF66E3159ADC4CCA767" author="Bruce" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978</bibRefCitation>
): (1) the rostrum is much narrower in
<taxonomicName id="4C1D91A6FFCBFFF6691D5840C6F9A778" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[628,743,1816,1840]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF6691D5840C6F9A778" box="[628,743,1816,1840]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF6691D5840C694A767" box="[628,650,1816,1839]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D91A6FFCBFFF668295840C7EEA778" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[832,1008,1816,1840]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF668295840C7EEA778" box="[832,1008,1816,1840]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF668295840C749A767" box="[832,855,1816,1839]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFCBFFF66897584FC176A767" author="Bruce" box="[1022,1384,1815,1839]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">
Bruce 1978: 37A versus
<figureCitation id="1326F6A0FFCBFFF66E78584FC14EA767" box="[1297,1360,1815,1839]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="9" pageNumber="59">Fig. 3</figureCitation>
A
</bibRefCitation>
); (2) the inferior orbital angle is acuminate in
<taxonomicName id="4C1D91A6FFCBFFF669095863C6CBA71B" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[608,725,1851,1875]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF669095863C6CBA71B" box="[608,725,1851,1875]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF669095863C668A71A" box="[608,630,1851,1874]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFCBFFF6698C5862C7D7A71A" author="Bruce" box="[741,969,1850,1874]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978: Fig. 36</bibRefCitation>
), but at most subacute in
<taxonomicName id="4C1D91A6FFCBFFF66F825863C185A71B" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1259,1435,1851,1875]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF66F825863C185A71B" box="[1259,1435,1851,1875]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66F825863C11CA71A" box="[1259,1282,1851,1874]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFCBFFF66BF65805C4FBA73D" box="[159,229,1885,1909]" captionStart="FIGURE 2" captionStartId="2.[151,250,1778,1801]" captionTargetBox="[184,1406,158,1777]" captionTargetId="figure@2.[170,1434,123,1803]" captionTargetPageId="2" captionText="FIGURE 2. Onycocaris hayamaensis n. sp. A – C, holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215; D, E, paratype, male (cl 2.2 mm), CBM-ZC 11216. A, D, carapace and cephalic appendages, lateral view; B, same, dorsal view; C, E, abdomen, lateral view (appendages omitted in E). Scale bars: 1 mm." httpUri="https://zenodo.org/record/214122/files/figure.png" pageId="9" pageNumber="59">Fig. 2</figureCitation>
A); (3) the telson is more strongly narrowed posteriorly, with lateral spines more anteriorly located in
<taxonomicName id="4C1D91A6FFCBFFF66EE85806C4F7A7D0" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF66EE85806C4F7A7D0" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66EE85806C186A73D" box="[1409,1432,1886,1909]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D91A6FFCBFFF66A2B58D8C5EEA7D0" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[322,496,1920,1944]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF66A2B58D8C5EEA7D0" box="[322,496,1920,1944]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66A2B58D8C547A7DF" box="[322,345,1920,1943]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(for example, the posterior pair is located at about the midlength in
<taxonomicName id="4C1D91A6FFCBFFF66FB258D8C151A7D0" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[1243,1359,1920,1944]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF66FB258D8C151A7D0" box="[1243,1359,1920,1944]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF66FB258D8C0ECA7DF" box="[1243,1266,1920,1943]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
, rather than distinctly posterior to the midlength in
<taxonomicName id="4C1D91A6FFCBFFF6691258FBC734A7F3" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[635,810,1955,1979]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFCBFFF6691258FBC734A7F3" box="[635,810,1955,1979]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF6691258FBC68CA7F2" box="[635,658,1955,1978]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
; (
<bibRefCitation id="EF8C97D4FFCBFFF6682858FAC0C4A7F2" author="Bruce" box="[833,1242,1954,1978]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">
Bruce 1978: Fig. 37E versus
<figureCitation id="1326F6A0FFCBFFF66FEC58FAC0DBA7F2" box="[1157,1221,1954,1978]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="9" pageNumber="59">Fig. 3</figureCitation>
C
</bibRefCitation>
); (4) the antennal scale has a small distolateral tooth exceeded by the distal lamella in
<taxonomicName id="4C1D91A6FFCBFFF6681E589EC7F7A795" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[887,1001,1989,2013]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="59" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFCBFFF6681E589EC7F7A795" box="[887,1001,1989,2013]" italics="true" pageId="9" pageNumber="59">
<collectingCountry id="F30AAAB5FFCBFFF6681E589EC790A795" box="[887,910,1990,2013]" name="Iceland" pageId="9" pageNumber="59">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFCBFFF6689E589DC119A795" author="Bruce" box="[1015,1287,1989,2013]" pageId="9" pageNumber="59" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978: Fig. 37C, D</bibRefCitation>
), whereas the distolateral tooth is strong and distinctly overreaches the lamella in
<taxonomicName id="4C1D91A6FFC8FFF568E15FC0C026A0F8" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[904,1080,152,176]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC8FFF568E15FC0C026A0F8" box="[904,1080,152,176]" italics="true" pageId="10" pageNumber="60">
<collectingCountry id="F30AAAB5FFC8FFF568E15FC0C781A0E7" box="[904,927,152,175]" name="Iceland" pageId="10" pageNumber="60">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC8FFF56F215FCFC094A0E7" box="[1096,1162,151,175]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="10" pageNumber="60">Fig. 3</figureCitation>
D); (5) the dactyli of the second pereopods have two cutting edges in
<taxonomicName id="4C1D91A6FFC8FFF569165FE3C6ECA09B" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[639,754,187,211]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFC8FFF569165FE3C6ECA09B" box="[639,754,187,211]" italics="true" pageId="10" pageNumber="60">
<collectingCountry id="F30AAAB5FFC8FFF569165FE3C688A09A" box="[639,662,187,210]" name="Iceland" pageId="10" pageNumber="60">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC8FFF568695FE2C7FEA09A" author="Bruce" box="[768,992,186,210]" pageId="10" pageNumber="60" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978: Fig. 40</bibRefCitation>
), but there is only one cutting edge in
<taxonomicName id="4C1D91A6FFC8FFF56EE85FE3C53BA0BD" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC8FFF56EE85FE3C53BA0BD" italics="true" pageId="10" pageNumber="60">
<collectingCountry id="F30AAAB5FFC8FFF56EE85FE3C186A09A" box="[1409,1432,187,210]" name="Iceland" pageId="10" pageNumber="60">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC8FFF56A5D5F85C56BA0BD" box="[308,373,221,245]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="10" pageNumber="60">Fig. 5</figureCitation>
A–C); (6) the mesial cutting edges of the fixed fingers of the second pereopod chelae are armed with small teeth in
<taxonomicName id="4C1D91A6FFC8FFF56A015E58C5C4A150" ID-CoL="fc47d0a9-d4a6-4156-975b-7b4e85a3e43c" box="[360,474,256,280]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B9693637FFC8FFF56A015E58C5C4A150" box="[360,474,256,280]" italics="true" pageId="10" pageNumber="60">
<collectingCountry id="F30AAAB5FFC8FFF56A015E58C561A15F" box="[360,383,256,279]" name="Iceland" pageId="10" pageNumber="60">O</collectingCountry>
. trullata
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC8FFF56A805FA7C6E2A15F" author="Bruce" box="[489,764,255,279]" pageId="10" pageNumber="60" refString="Bruce, A. J. (1978) A report on a collection of pontoniine shrimps from Madagascar and adjacent seas. Zoological Journal of the Linnean Society, 62, 205 - 290." type="journal article" year="1978">Bruce 1978: Fig. 40B, D</bibRefCitation>
), whereas such teeth are absent in
<taxonomicName id="4C1D91A6FFC8FFF56F1C5E58C13AA150" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1141,1316,256,280]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC8FFF56F1C5E58C13AA150" box="[1141,1316,256,280]" italics="true" pageId="10" pageNumber="60">
<collectingCountry id="F30AAAB5FFC8FFF56F1C5E58C092A15F" box="[1141,1164,256,279]" name="Iceland" pageId="10" pageNumber="60">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC8FFF56E5A5FA7C16AA15F" box="[1331,1396,255,279]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1903]" captionTargetBox="[183,1388,182,1879]" captionTargetId="figure@7.[172,1416,173,1890]" captionTargetPageId="7" captionText="FIGURE 5. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, right major second pereopod, lateral view; B, same, chela and carpus, mesial view; C, same, distal part of fingers, lateral view, showing details of armature of cutting edges; D, same, distal part of fixed finger, ventral (flexor) view; E, left minor second pereopod, lateral view; F, same, chela and carpus, mesial view; G, same, distal part of fixed finger, ventral (flexor) view. Scale bars: 0.5 mm for A, B, D – F; 0.25 mm for C." httpUri="https://zenodo.org/record/214125/files/figure.png" pageId="10" pageNumber="60">Fig. 5</figureCitation>
B).
</paragraph>
<caption id="DF62BAADFFC8FFF56BFE58CDC76AA7A1" httpUri="https://zenodo.org/record/214127/files/figure.png" pageId="10" pageNumber="60" targetBox="[171,1414,317,1922]" targetPageId="10">
<paragraph id="8BA2EA25FFC8FFF56BFE58CDC76AA7A1" blockId="10.[151,1435,1941,2025]" pageId="10" pageNumber="60">
<emphasis id="B9693637FFC8FFF56BFE58CDC50CA7E3" bold="true" box="[151,274,1941,1964]" pageId="10" pageNumber="60">FIGURE 7.</emphasis>
Underwater photographs. A,
<taxonomicName id="4C1D91A6FFC8FFF5695558CFC724A7E4" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[572,826,1942,1964]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC8FFF5695558CFC724A7E4" box="[572,826,1942,1964]" italics="true" pageId="10" pageNumber="60">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC8FFF5685658CEC766A7E3" box="[831,888,1942,1963]" pageId="10" pageNumber="60" rank="species">
<emphasis id="B9693637FFC8FFF5685658CEC766A7E3" bold="true" box="[831,888,1942,1963]" pageId="10" pageNumber="60">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 3.4 mm) (indicated by red arrow), and host sponge
<taxonomicName id="4C1D91A6FFC8FFF56AAC58EDC634A783" ID-CoL="84NXG" box="[453,554,1973,1995]" class="Demospongiae" family="Chalinidae" genus="Haliclona" kingdom="Animalia" order="Haplosclerida" pageId="10" pageNumber="60" phylum="Porifera" rank="genus">
<emphasis id="B9693637FFC8FFF56AAC58EDC634A783" box="[453,554,1973,1995]" italics="true" pageId="10" pageNumber="60">Haliclona</emphasis>
</taxonomicName>
sp.; B,
<taxonomicName id="4C1D91A6FFC8FFF569E958EDC79FA783" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[640,897,1973,1995]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC8FFF569E958EDC79FA783" box="[640,897,1973,1995]" italics="true" pageId="10" pageNumber="60">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC8FFF568E358EDC7D8A782" box="[906,966,1973,1994]" pageId="10" pageNumber="60" rank="species">
<emphasis id="B9693637FFC8FFF568E358EDC7D8A782" bold="true" box="[906,966,1973,1994]" pageId="10" pageNumber="60">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 3.4 mm); C,
<taxonomicName id="4C1D91A6FFC8FFF56BFE588CC58BA7A1" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[151,405,2003,2025]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="60" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC8FFF56BFE588CC58BA7A1" box="[151,405,2003,2025]" italics="true" pageId="10" pageNumber="60">Onycocaris hayamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC8FFF56AF2588CC5CAA7A1" box="[411,468,2004,2025]" pageId="10" pageNumber="60" rank="species">
<emphasis id="B9693637FFC8FFF56AF2588CC5CAA7A1" bold="true" box="[411,468,2004,2025]" pageId="10" pageNumber="60">n. sp.</emphasis>
</taxonomicNameLabel>
, male, spouse of holotype, not collected.
</paragraph>
</caption>
<paragraph id="8BA2EA25FFC9FFF46BAE5FC0C700A284" blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="61">
<taxonomicName id="4C1D91A6FFC9FFF46BAE5FC0C587A0F8" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[199,409,152,176]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46BAE5FC0C587A0F8" box="[199,409,152,176]" italics="true" pageId="11" pageNumber="61">Onycocaris fujinoi</emphasis>
</taxonomicName>
was named based on two specimens (one male and one ovigerous female, the former was designated as the
<typeStatus id="54A65487FFC9FFF46A355FE4C5DEA09C" box="[348,448,188,212]" pageId="11" pageNumber="61" type="holotype">holotype</typeStatus>
) from Yoron Islands, Ryukyu Islands,
<collectingCountry id="F30AAAB5FFC9FFF468045FE5C7AEA09C" box="[877,944,189,212]" name="Japan" pageId="11" pageNumber="61">Japan</collectingCountry>
, originally referred to
<taxonomicName id="4C1D91A6FFC9FFF46FC35FE5C182A09C" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[1194,1436,188,212]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFC9FFF46FC35FE5C182A09C" box="[1194,1436,188,212]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46FC35FE5C0DFA09C" box="[1194,1217,189,212]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. quadratophthalma
</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8C97D4FFC9FFF46BD55F87C5C7A0BF" author="Fujino" box="[188,473,223,247]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake (1969)</bibRefCitation>
, though
<bibRefCitation id="EF8C97D4FFC9FFF469545FB8C6FFA0BF" author="Bruce" box="[573,737,223,247]" pageId="11" pageNumber="61" refString="Bruce, A. J. (2011 b) Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balssi sp. nov., from northern Australia, with the designation of O. fujinoi sp. nov. Crustaceana, 84, 477 - 490." type="journal article" year="2011" yearSuffix="b">Bruce (2011b)</bibRefCitation>
did not examine directly those specimens. In Bruce’s (2011: 489) identification key, the tip of the fixed finger of the second pereopod chela is said to be simple in
<taxonomicName id="4C1D91A6FFC9FFF46E695E5DC174A154" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[1280,1386,260,284]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46E695E5DC174A154" box="[1280,1386,260,284]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46E695E5DC109A154" box="[1280,1303,261,284]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
, but
<bibRefCitation id="EF8C97D4FFC9FFF46BFE5E7FC5E1A177" author="Fujino" box="[151,511,295,319]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake (1969: 440)</bibRefCitation>
clearly stated that it was bifid in the minor cheliped of the male, as in
<taxonomicName id="4C1D91A6FFC9FFF46EE85E70C536A12C" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis" status="sp. nov.">
<emphasis id="B9693637FFC9FFF46EE85E70C536A12C" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46EE85E70C186A177" box="[1409,1432,296,319]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A8B4CFFC9FFF46A5A5E14C568A12C" box="[307,374,332,356]" pageId="11" pageNumber="61" rank="species">
<emphasis id="B9693637FFC9FFF46A5A5E14C568A12C" bold="true" box="[307,374,332,356]" pageId="11" pageNumber="61">n. sp.</emphasis>
</taxonomicNameLabel>
Nevertheless,
<taxonomicName id="4C1D91A6FFC9FFF4694C5E15C6E5A12C" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[549,763,332,356]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF4694C5E15C6E5A12C" box="[549,763,332,356]" italics="true" pageId="11" pageNumber="61">Onycocaris fujinoi</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C1D91A6FFC9FFF46F7F5E15C0D0A12C" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1046,1230,332,356]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46F7F5E15C0D0A12C" box="[1046,1230,332,356]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46F7F5E15C033A12C" box="[1046,1069,333,356]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
by the following characters (
<bibRefCitation id="EF8C97D4FFC9FFF46A715E37C63DA1CF" author="Fujino" box="[280,547,367,391]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
): (1) the posterior margin of the telson bears one median plumose seta flanked by two pairs of spines in
<taxonomicName id="4C1D91A6FFC9FFF46AD95ECDC602A1E4" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[432,540,404,428]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46AD95ECDC602A1E4" box="[432,540,404,428]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46AD95ECDC5D9A1E4" box="[432,455,405,428]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF469435ECCC782A1E4" author="Fujino" box="[554,924,404,428]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969" yearSuffix="b">Fujino &amp; Miyake 1969: Fig. 18b</bibRefCitation>
), whereas there are three pairs of spines in
<taxonomicName id="4C1D91A6FFC9FFF46EE85ECDC536A198" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46EE85ECDC536A198" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46EE85ECDC186A1E4" box="[1409,1432,405,428]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF46A5F5EEFC566A187" box="[310,376,439,463]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
C); (2) the eye is relatively narrower with the cornea more tapering distally in
<taxonomicName id="4C1D91A6FFC9FFF46FB75EE0C157A198" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[1246,1353,440,464]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46FB75EE0C157A198" box="[1246,1353,440,464]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46FB75EE0C0EBA187" box="[1246,1269,440,463]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D91A6FFC9FFF46BFE5E85C553A1BC" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[151,333,476,500]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46BFE5E85C553A1BC" box="[151,333,476,500]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46BFE5E85C4B0A1BC" box="[151,174,477,500]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF46A375E84C78BA1BC" author="Fujino" box="[350,917,476,500]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969" yearSuffix="b">
Fujino &amp; Miyake 1969: Fig. 16b versus
<figureCitation id="1326F6A0FFC9FFF468505E84C760A1BC" box="[825,894,476,500]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
A
</bibRefCitation>
); (3) the chela of the first pereopod is much slenderer in
<taxonomicName id="4C1D91A6FFC9FFF46A4F5D58C58AA250" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[294,404,512,536]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46A4F5D58C58AA250" box="[294,404,512,536]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46A4F5D58C523A25F" box="[294,317,512,535]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D91A6FFC9FFF46A9F5D58C6B2A250" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[502,684,512,536]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46A9F5D58C6B2A250" box="[502,684,512,536]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46A9F5D58C613A25F" box="[502,525,512,535]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(about 8.5 times longer than the greatest width versus 5.5 times) (
<bibRefCitation id="EF8C97D4FFC9FFF46BF75D7CC6F7A274" author="Fujino" box="[158,745,548,572]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969" yearSuffix="h">
Fujino &amp; Miyake 1969: Fig. 18h versus
<figureCitation id="1326F6A0FFC9FFF469055D7CC6AEA274" box="[620,688,548,572]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="11" pageNumber="61">Fig. 4</figureCitation>
C, D
</bibRefCitation>
); (4) fingers of the chela of the male major second pereopod are armed with one (dactylus) and two (fixed finger) teeth on respective cutting edges in
<taxonomicName id="4C1D91A6FFC9FFF46FD95D10C100A228" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[1200,1310,584,608]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46FD95D10C100A228" box="[1200,1310,584,608]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46FD95D10C0D9A217" box="[1200,1223,584,607]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF46E595D1FC582A2CC" author="Fujino" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969" yearSuffix="c">Fujino &amp; Miyake 1969: Fig. 16c</bibRefCitation>
), rather than two (dactylus) and three (fixed finger) teeth in
<taxonomicName id="4C1D91A6FFC9FFF46F255D35C11FA2CC" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1100,1281,620,644]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46F255D35C11FA2CC" box="[1100,1281,620,644]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46F255D35C07DA2CC" box="[1100,1123,621,644]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF46E785D34C14BA2CC" box="[1297,1365,620,644]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="11" pageNumber="61">Fig. 6</figureCitation>
A, B); the palm of the male major second pereopod is smooth on the ventral margin in
<taxonomicName id="4C1D91A6FFC9FFF46F7B5DC8C063A2E0" ID-CoL="58d89622-31f6-42c7-8688-555c856ab32f" box="[1042,1149,656,680]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="fujinoi">
<emphasis id="B9693637FFC9FFF46F7B5DC8C063A2E0" box="[1042,1149,656,680]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46F7B5DC8C037A2EF" box="[1042,1065,656,679]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. fujinoi
</emphasis>
</taxonomicName>
(Funino &amp;
<bibRefCitation id="EF8C97D4FFC9FFF46E695DD7C186A2EF" author="Miyake" box="[1280,1432,655,679]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Miyake 1969</bibRefCitation>
: Fig. 16c), but tuberculate in
<taxonomicName id="4C1D91A6FFC9FFF46ABC5DEDC696A284" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[469,648,692,716]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46ABC5DEDC696A284" box="[469,648,692,716]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46ABC5DEDC5F2A284" box="[469,492,693,716]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF469FE5DECC6C7A284" box="[663,729,692,716]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1924]" captionTargetBox="[211,1383,217,1854]" captionTargetId="figure@8.[176,1410,193,1880]" captionTargetPageId="8" captionText="FIGURE 6. Onycocaris hayamaensis n. sp., paratype, male (cl 2.9 mm), CBM-ZC 11216. A, major right second pereopod, lateral view; B, same, mesial view; C, minor left second pereopod, lateral view; D, same, chela and carpus, mesial view; E, left first pleopod, dorsal view; F, appendices interna and masculina on endopod of left second pleopod, mesial view. Scale bars: 1 mm for A – D; 0.5 mm for E; 0.25 mm for I." httpUri="https://zenodo.org/record/214126/files/figure.png" pageId="11" pageNumber="61">Fig. 6</figureCitation>
A, B).
</paragraph>
<paragraph id="8BA2EA25FFC9FFF46BAE5D80C51CA544" blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="61">
<taxonomicName id="4C1D91A6FFC9FFF46BAE5D80C63AA2B8" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[199,548,728,752]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFC9FFF46BAE5D80C63AA2B8" box="[199,548,728,752]" italics="true" pageId="11" pageNumber="61">Onycocaris quadratophthalma</emphasis>
</taxonomicName>
is represented with certainty only by the female
<typeStatus id="54A65487FFC9FFF46F1E5D8FC0C7A2A7" box="[1143,1241,727,751]" pageId="11" pageNumber="61" type="holotype">holotype</typeStatus>
and one female specimen subsequently reported from
<collectingCountry id="F30AAAB5FFC9FFF4693F5DA5C6C0A35D" box="[598,734,765,789]" name="Hong Kong" pageId="11" pageNumber="61">Hong Kong</collectingCountry>
(
<bibRefCitation id="EF8C97D4FFC9FFF4699B5DA4C760A35C" author="Bruce" box="[754,894,764,788]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">Bruce 1992</bibRefCitation>
). The second pereopods of the
<typeStatus id="54A65487FFC9FFF46F915DA4C144A35C" box="[1272,1370,764,788]" pageId="11" pageNumber="61" type="holotype">holotype</typeStatus>
were missing (
<bibRefCitation id="EF8C97D4FFC9FFF46B975C47C561A37F" author="Balss" box="[254,383,799,823]" pageId="11" pageNumber="61" refString="Balss, H. (1921) Results of Dr. E. Mjobergs Swedish Scientific Expeditions to Australia 1910 - 13. XXIX: Stomatopoda, Macrura, Paguridea und Galatheidea. Kungliga Svenska Vetenskapsakademiens Handlinger, 61, 1 - 24." type="journal article" year="1921">Balss 1921</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFC9FFF46AE25C47C68DA37F" author="Fujino" box="[395,659,799,823]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
); in the specimen from
<collectingCountry id="F30AAAB5FFC9FFF468F75C79C038A371" box="[926,1062,801,825]" name="Hong Kong" pageId="11" pageNumber="61">Hong Kong</collectingCountry>
, only the major second pereopod was preserved.
<taxonomicName id="4C1D91A6FFC9FFF46A275C1DC6B2A314" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[334,684,836,860]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFC9FFF46A275C1DC6B2A314" box="[334,684,836,860]" italics="true" pageId="11" pageNumber="61">Onycocaris quadratophthalma</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C1D91A6FFC9FFF46F7D5C1DC0D3A314" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1044,1229,836,860]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46F7D5C1DC0D3A314" box="[1044,1229,836,860]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46F7D5C1DC035A314" box="[1044,1067,837,860]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
by the following characters (
<bibRefCitation id="EF8C97D4FFC9FFF46A7F5C3FC605A337" author="Fujino" box="[278,539,871,895]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFC9FFF4694F5C31C6B3A337" author="Bruce" box="[550,685,871,896]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">Bruce 1992</bibRefCitation>
): (1) the eyes are quadrate with flat and transverse anterior surfaces and contiguous mesial surfaces in
<taxonomicName id="4C1D91A6FFC9FFF469745CD5C70EA3EC" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[541,784,908,932]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
<emphasis id="B9693637FFC9FFF469745CD5C70EA3EC" box="[541,784,908,932]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF469745CD5C62AA3EC" box="[541,564,909,932]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. quadratophthalma
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF468495CD4C035A3EC" author="Fujino" box="[800,1067,908,932]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
: Fig. 16a;
<bibRefCitation id="EF8C97D4FFC9FFF46FCD5CD5C135A3EC" author="Bruce" box="[1188,1323,908,932]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">Bruce 1992</bibRefCitation>
:
<figureCitation id="1326F6A0FFC9FFF46E5F5CD4C168A3EC" box="[1334,1398,908,932]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="11" pageNumber="61">Fig. 1</figureCitation>
B), whereas they are less quadrate with anterior surfaces sloping to the non-contiguous mesial surfaces in
<taxonomicName id="4C1D91A6FFC9FFF46EE85CE8C536A3A4" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46EE85CE8C536A3A4" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46EE85CE8C186A38F" box="[1409,1432,944,967]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF46A5E5C8CC567A3A4" box="[311,377,980,1004]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
A); (2) the antennal scale has a small distolateral tooth exceeding as far as the distal lamella in
<taxonomicName id="4C1D91A6FFC9FFF46BFE5CA0C58EA458" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[151,400,1016,1040]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
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. quadratophthalma
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF46ACD5CAFC6A0A447" author="Fujino" box="[420,702,1015,1039]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
: Fig. 16a;
<bibRefCitation id="EF8C97D4FFC9FFF4682F5CA1C7CCA447" author="Bruce" box="[838,978,1015,1040]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">Bruce 1992</bibRefCitation>
:
<figureCitation id="1326F6A0FFC9FFF4688A5CAFC039A447" box="[995,1063,1015,1039]" captionStart="FIGURE 1" captionStartId="1.[151,250,1448,1471]" captionTargetBox="[151,1435,765,1424]" captionTargetId="figure@1.[151,1436,765,1424]" captionTargetPageId="1" captionText="FIGURE 1. Onycocaris hayamaensis n. sp., paratype, ovigerous female (cl 3.8 mm), entire animal in lateral view, CBM-ZC 11216." httpUri="https://zenodo.org/record/214121/files/figure.png" pageId="11" pageNumber="61">Fig. 1</figureCitation>
F), rather than having a strong distolateral tooth distinctly overreaching the distal lamella in
<taxonomicName id="4C1D91A6FFC9FFF4682D5B45C7E6A47C" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[836,1016,1052,1076]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF4682D5B45C7E6A47C" box="[836,1016,1052,1076]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF4682D5B45C745A47C" box="[836,859,1053,1076]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF46F6F5B44C057A47C" box="[1030,1097,1052,1076]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
D); (3) the fingers of the chela of the female major second pereopods bear two blunt teeth (dactylus) and one prominent middle tooth (fixed finger) on respective cutting edges in
<taxonomicName id="4C1D91A6FFC9FFF4692D5B3DC727A434" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[580,825,1124,1148]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
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<collectingCountry id="F30AAAB5FFC9FFF4692D5B3DC645A434" box="[580,603,1125,1148]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. quadratophthalma
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF468215B3CC04AA434" author="Bruce" box="[840,1108,1124,1148]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">
Bruce 1992:
<figureCitation id="1326F6A0FFC9FFF468B55B3CC000A434" box="[988,1054,1124,1148]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
D, E
</bibRefCitation>
), but such conspicuous teeth are absent on the fingers of the major second pereopod in
<taxonomicName id="4C1D91A6FFC9FFF468455BD0C7FEA4E8" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[812,992,1160,1184]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF468455BD0C7FEA4E8" box="[812,992,1160,1184]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF468455BD0C75DA4D7" box="[812,835,1160,1183]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF468995BD0C02AA4E8" box="[1008,1076,1160,1184]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="11" pageNumber="61">Fig. 4</figureCitation>
A–C); the dactyli of the third to fifth pereopods are devoid of accessory denticles proximal to the distal accessory tooth in
<taxonomicName id="4C1D91A6FFC9FFF46FCE5BF5C182A48C" ID-CoL="133da176-8194-4d79-8d16-f4a4571837c0" box="[1191,1436,1196,1220]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="quadratophthalma">
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. quadratophthalma
</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C97D4FFC9FFF46BF75B88C596A4A0" author="Bruce" box="[158,392,1232,1256]" pageId="11" pageNumber="61" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ED.) The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 11 - 29 April 1989, 329 - 343." type="journal article" year="1992">
Bruce 1992:
<figureCitation id="1326F6A0FFC9FFF46A595B88C56EA4A0" box="[304,368,1232,1256]" captionStart="FIGURE 3" captionStartId="3.[151,250,1857,1880]" captionTargetBox="[226,1348,185,1831]" captionTargetId="figure@3.[201,1387,147,1845]" captionTargetPageId="3" captionText="FIGURE 3. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, anterior part of carapace and cephalic appendages, dorsal view; B, sixth abdominal somite, dorsal view; C, telson, dorsal view; D, left antenna, ventral view (flagellum missing); E, left mandible, outer view; F, same, distal part of incisor process, inner view; G, same, distal part of molar process, outer view; H, left maxillule, outer view; I, same, distal part of basial endite, inner view; J, same, palp, inner view; K, left maxilla, outer view (endites broken); L, left first maxilliped, outer view; M, left second maxilliped, outer view; N, left uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm for A – E, H, K – N; 0.25 mm for I, J; 0.1 mm for F, G." httpUri="https://zenodo.org/record/214123/files/figure.png" pageId="11" pageNumber="61">Fig. 3</figureCitation>
G
</bibRefCitation>
), whereas there are several denticles proximal to the distal accessory tooth in
<taxonomicName id="4C1D91A6FFC9FFF46F805B88C182A4A0" ID-CoL="70d71967-a273-43bc-b15d-e9cea94461ef" box="[1257,1436,1232,1256]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="hayamaensis">
<emphasis id="B9693637FFC9FFF46F805B88C182A4A0" box="[1257,1436,1232,1256]" italics="true" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46F805B88C11EA4AF" box="[1257,1280,1232,1255]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. hayamaensis
</emphasis>
</taxonomicName>
(
<figureCitation id="1326F6A0FFC9FFF46BF65BACC4FDA544" box="[159,227,1268,1292]" captionStart="FIGURE 4" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[181,1406,180,1904]" captionTargetId="figure@5.[151,1436,173,1912]" captionTargetPageId="5" captionText="FIGURE 4. Onycocaris hayamaensis n. sp., holotype, ovigerous female (cl 3.4 mm), CBM-ZC 11215. A, left third maxilliped, lateral view; B, same, antepenultimate segment, coxa, and exopod, dorsal view; C, left first pereopod, lateral view; D, same, chela; E, left third pereopod, lateral view; F, same, dactylus, lateral view; G, left fourth pereopod, lateral view; H, left fifth pereopod, lateral view. Scale bars: 0.5 mm for A – C, E, G, H; 0.25 mm for D; 0.1 mm for F." httpUri="https://zenodo.org/record/214124/files/figure.png" pageId="11" pageNumber="61">Fig. 4</figureCitation>
F).
</paragraph>
<paragraph id="8BA2EA25FFC9FFF46BAE5A40C148A5AD" blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="61">
<collectingCountry id="F30AAAB5FFC9FFF46BAE5A40C4F8A578" box="[199,230,1304,1328]" name="American Samoa" pageId="11" pageNumber="61">As</collectingCountry>
mentioned before, six previously described species have been recorded from Japanese waters, of them the record of
<taxonomicName id="4C1D91A6FFC9FFF46A6B5A65C5A8A51C" ID-CoL="c1c15a6f-8586-4f5f-b7f3-fc8bba28a89e" box="[258,438,1340,1364]" class="Malacostraca" family="Palaemonidae" genus="Onycocaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="61" phylum="Arthropoda" rank="species" species="seychellensis">
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<collectingCountry id="F30AAAB5FFC9FFF46A6B5A65C507A51C" box="[258,281,1341,1364]" name="Iceland" pageId="11" pageNumber="61">O</collectingCountry>
. seychellensis
</emphasis>
</taxonomicName>
mentioned by
<bibRefCitation id="EF8C97D4FFC9FFF469325A64C71CA51C" author="Miyake" box="[603,770,1340,1364]" pageId="11" pageNumber="61" refString="Miyake, S. (1982) Japanese Crustacean Decapods and Stomatopods in Color. Vol. I. Macrura, Anomura and Stomatopoda. Hoikusha Publishing Co., Ltd., Osaka, 261 pp., 56 pls. [in Japanese]" type="book" year="1982">Miyake (1982)</bibRefCitation>
and
<bibRefCitation id="EF8C97D4FFC9FFF4685E5A64C02DA51C" author="Chace" box="[823,1075,1340,1364]" pageId="11" pageNumber="61" refString="Chace, F. A., Jr. &amp; Bruce, A. J. (1993) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition 1907 - 1910, Part 6: Superfamily Palaemonoidea. Smithsonian Contributions to Zoology, 543, i - vii, 1 - 152." type="journal article" year="1993">Chace &amp; Bruce (1993)</bibRefCitation>
needs to be verified, because no voucher specimen was indicated or available (cf.
<bibRefCitation id="EF8C97D4FFC9FFF469A25A38C772A530" author="Hayashi" box="[715,876,1376,1400]" pageId="11" pageNumber="61" refString="Hayashi, K. (2003) Shrimps and lobsters from Japan (127). Family Palaemonidae, subfamily Pontoniinae - Genus Onycocaris. Aquabiology, 144, 21 - 26." type="journal article" year="2003">Hayashi 2003</bibRefCitation>
). Other five species have been recorded from the southwestern part of the Japanese Archipelago (
<bibRefCitation id="EF8C97D4FFC9FFF469D05ADCC7DAA5D4" author="Fujino" box="[697,964,1412,1436]" pageId="11" pageNumber="61" refString="Fujino, T. &amp; Miyake, S. (1969) Studies on the genus Onycocaris with descriptions of five new species (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 15, 403 - 448." type="journal article" year="1969">Fujino &amp; Miyake 1969</bibRefCitation>
;
<bibRefCitation id="EF8C97D4FFC9FFF468B85ADCC06CA5D4" author="Hayashi" box="[977,1138,1412,1436]" pageId="11" pageNumber="61" refString="Hayashi, K. (2003) Shrimps and lobsters from Japan (127). Family Palaemonidae, subfamily Pontoniinae - Genus Onycocaris. Aquabiology, 144, 21 - 26." type="journal article" year="2003">Hayashi 2003</bibRefCitation>
), although they have been rarely collected. The scarce records of the genus in local waters may reflect scarce sampling efforts. In fact, the present new species was discovered from the shallow subtidal depths where access for collection is not easy.
</paragraph>
</subSubSection>
<subSubSection id="C307B9AEFFC9FFF46BAE5AA8C06AA640" box="[199,1140,1520,1545]" pageId="11" pageNumber="61" type="etymology">
<paragraph id="8BA2EA25FFC9FFF46BAE5AA8C06AA640" blockId="11.[151,1437,151,1545]" box="[199,1140,1520,1545]" pageId="11" pageNumber="61">
<emphasis id="B9693637FFC9FFF46BAE5AA8C558A640" bold="true" box="[199,326,1520,1544]" pageId="11" pageNumber="61">Etymology</emphasis>
. Named after the
<typeStatus id="54A65487FFC9FFF469795AA9C65EA641" box="[528,576,1521,1545]" pageId="11" pageNumber="61">type</typeStatus>
locality, Hayama, in Kanagawa Prefecture,
<collectingCountry id="F30AAAB5FFC9FFF46F445AA9C06EA640" box="[1069,1136,1521,1544]" name="Japan" pageId="11" pageNumber="61">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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