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<document id="5EB1AD186DE9D738293F80273D31B26B" ID-DOI="10.5281/zenodo.5169054" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169054" ID-ZooBank="5265BC7E-7076-4225-B659-678E3990B42C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628273540546" checkinUser="felipe" docAuthor="Otto, Robert L." docDate="2017" docId="03AC8798FFFD2744FF5F821D3073FCE3" docLanguage="en" docName="InsectaMundi.0569.1-27.pdf" docOrigin="Insecta Mundi 2017 (569)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phlegon panamensis , Otto 2017, sp. nov." docType="treatment" docUuid="54A2463A-F70D-4F68-893C-98911655B643" docUuidSource="ZooBank" docVersion="4" lastPageNumber="10" masterDocId="FF95FFE0FFF7274FFF9F837C3376FFD9" masterDocTitle="A revision of Phlegoninae (Coleoptera: Eucnemidae), with descriptions of a new genus and four new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="9" updateTime="1699038921155" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="0BAED2C2822D24ADFF44A1D6A70FD148">A revision of Phlegoninae (Coleoptera: Eucnemidae), with descriptions of a new genus and four new species</mods:title>
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<mods:namePart id="7399886E513F40F74BD61A80DDA50F5A">Otto, Robert L.</mods:namePart>
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<mods:date id="C31E0455FBC880681205162E32C0F31C">2017</mods:date>
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<mods:number id="1E40B8A2899CF7887D56E54B03D21173">2017-08-25</mods:number>
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<treatment id="03AC8798FFFD2744FF5F821D3073FCE3" ID-DOI="http://doi.org/10.5281/zenodo.5185395" ID-Zenodo-Dep="5185395" LSID="urn:lsid:zoobank.org:act:54A2463A-F70D-4F68-893C-98911655B643" httpUri="http://treatment.plazi.org/id/03AC8798FFFD2744FF5F821D3073FCE3" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
<subSubSection id="C31F6505FFFD2745FF5F821D313AFEA3" box="[192,588,353,378]" pageId="10" pageNumber="9" type="nomenclature">
<paragraph id="8BBA368EFFFD2745FF5F821D313AFEA3" blockId="10.[192,588,353,410]" box="[192,588,353,378]" pageId="10" pageNumber="9">
<heading id="D0F281E2FFFD2745FF5F821D313AFEA3" bold="true" box="[192,588,353,378]" fontSize="10" level="2" pageId="10" pageNumber="9" reason="3">
<emphasis id="B971EA9CFFFD2745FF5F821D313AFEA3" bold="true" box="[192,588,353,378]" pageId="10" pageNumber="9">
<taxonomicName id="4C054D0DFFFD2745FF5F821D32AAFEA0" authority="Otto, 2017" authorityName=", Otto" authorityYear="2017" box="[192,476,353,377]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B971EA9CFFFD2745FF5F821D32AAFEA0" bold="true" box="[192,476,353,377]" italics="true" pageId="10" pageNumber="9">Phlegon panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257E7FFFD2745FE7B821E313AFEA3" box="[484,588,354,378]" pageId="10" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFD2745FF5F82FE3234FE40" box="[192,322,385,410]" pageId="10" pageNumber="9" type="description">
<paragraph id="8BBA368EFFFD2745FF5F82FE3234FE40" blockId="10.[192,588,353,410]" box="[192,322,385,410]" pageId="10" pageNumber="9">
<figureCitation id="133E2A0BFFFD2745FF5F82FE3234FE40" box="[192,322,385,410]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="10" pageNumber="9">Fig. 2025</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFD2745FF5F82BD30FCFD40" pageId="10" pageNumber="9" type="diagnosis">
<paragraph id="8BBA368EFFFD2745FF5F82BD30FCFD40" blockId="10.[192,1440,449,666]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF5F82BD3226FE00" bold="true" box="[192,336,449,473]" pageId="10" pageNumber="9">Diagnosis.</emphasis>
Rounded punctures along elytral sutures distinguish this species from both
<taxonomicName id="4C054D0DFFFD2745FA9782BE36D6FE00" authorityName=", Fleutiaux, Det. R. L. Otto" authorityYear="2005" box="[1288,1440,450,473]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B971EA9CFFFD2745FA9782BE36D6FE00" box="[1288,1440,450,473]" italics="true" pageId="10" pageNumber="9">P. giganteus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D0DFFFD2745FF6A829E32C2FE23" authorityName=", Otto" authorityYear="2017" box="[245,436,482,506]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chiriquiensis">
<emphasis id="B971EA9CFFFD2745FF6A829E32C2FE23" box="[245,436,482,506]" italics="true" pageId="10" pageNumber="9">P. chiriquiensis</emphasis>
</taxonomicName>
. Males are distinguished from those of
<taxonomicName id="4C054D0DFFFD2745FC07829E373BFE23" authorityName=", (Lacordaire" authorityYear="1857" box="[920,1101,482,506]" class="Insecta" family="Eucnemidae" genus="Euryptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="herculeanus">
<emphasis id="B971EA9CFFFD2745FC07829E373BFE23" box="[920,1101,482,506]" italics="true" pageId="10" pageNumber="9">P. herculeanus</emphasis>
</taxonomicName>
by shorter antennomere IX in relation to X; subequal in both segments for
<taxonomicName id="4C054D0DFFFD2745FC93817E30B5FDC3" authorityName=", (Lacordaire" authorityYear="1857" box="[780,963,514,538]" class="Insecta" family="Eucnemidae" genus="Euryptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="herculeanus">
P.
<emphasis id="B971EA9CFFFD2745FCB1817E30B5FDC3" box="[814,963,514,538]" italics="true" pageId="10" pageNumber="9">herculeanus</emphasis>
</taxonomicName>
. Males are further distinguished from male
<taxonomicName id="4C054D0DFFFD2745FE9B815E32F7FDE3" authorityName=", Laporte" authorityYear="1840" box="[260,385,546,570]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FE9B815E32F7FDE3" box="[260,385,546,570]" italics="true" pageId="10" pageNumber="9">P. buqueti</emphasis>
</taxonomicName>
by elongate, thinner antennal rami on antennomeres IXX; shorter and thicker in
<taxonomicName id="4C054D0DFFFD2745FA18815E326AFD83" authorityName=", Laporte" authorityYear="1840" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FA18815E326AFD83" italics="true" pageId="10" pageNumber="9">P. buqueti</emphasis>
</taxonomicName>
. Females of
<taxonomicName id="4C054D0DFFFD2745FE23813E3107FD80" authorityName=", Otto" authorityYear="2017" box="[444,625,578,601]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B971EA9CFFFD2745FE23813E3107FD80" box="[444,625,578,601]" italics="true" pageId="10" pageNumber="9">P. panamensis</emphasis>
</taxonomicName>
can be distinguished from both
<taxonomicName id="4C054D0DFFFD2745FB9A813E37F5FD83" authorityName=", Laporte" authorityYear="1840" box="[1029,1155,578,602]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FB9A813E37F5FD83" box="[1029,1155,578,602]" italics="true" pageId="10" pageNumber="9">P. buqueti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D0DFFFD2745FB5C813E360CFD83" authorityName=", (Lacordaire" authorityYear="1857" box="[1219,1402,578,602]" class="Insecta" family="Eucnemidae" genus="Euryptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="herculeanus">
<emphasis id="B971EA9CFFFD2745FB5C813E360CFD83" box="[1219,1402,578,602]" italics="true" pageId="10" pageNumber="9">P. herculeanus</emphasis>
</taxonomicName>
by relative length of antennomere IX in relation to X; antennomere IX longer than X in both
<taxonomicName id="4C054D0DFFFD2745FABD811E36D6FDA3" authorityName=", Laporte" authorityYear="1840" box="[1314,1440,610,634]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FABD811E36D6FDA3" box="[1314,1440,610,634]" italics="true" pageId="10" pageNumber="9">P. buqueti</emphasis>
</taxonomicName>
and
<emphasis id="B971EA9CFFFD2745FF6981FE32C5FD43" box="[246,435,642,666]" italics="true" pageId="10" pageNumber="9">
<taxonomicName id="4C054D0DFFFD2745FF6981FE32DBFD43" authorityName=", (Lacordaire" authorityYear="1857" box="[246,429,642,666]" class="Insecta" family="Eucnemidae" genus="Euryptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="herculeanus">P. herculeanus</taxonomicName>
,
</emphasis>
and IX as long as X in
<taxonomicName id="4C054D0DFFFD2745FD4F81FE30F5FD40" authorityName=", Otto" authorityYear="2017" box="[720,899,642,665]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B971EA9CFFFD2745FD4F81FE30F5FD40" box="[720,899,642,665]" italics="true" pageId="10" pageNumber="9">P. panamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFD2745FF5F81BE3078FBE3" pageId="10" pageNumber="9" type="materials_examined">
<paragraph id="8BBA368EFFFD2745FF5F81BE30B7FCE3" blockId="10.[192,1440,705,826]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF5F81BE312EFD03" bold="true" box="[192,600,705,730]" pageId="10" pageNumber="9">
Type Material. Male
<typeStatus id="54BE882CFFFD2745FE4081BE312EFD03" box="[479,600,706,730]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
</emphasis>
: “La Chorrera, 10.V.12 Pan, Aug. Busck” / “J. Muona det.,
<taxonomicName id="4C054D0DFFFD2745FAA981BE36EFFD03" authorityName="Laporte" authorityYear="1840" box="[1334,1433,706,730]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B971EA9CFFFD2745FAA981BE36EFFD03" box="[1334,1433,706,730]" italics="true" pageId="10" pageNumber="9">Phlegon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D0DFFFD2745FF5F819E322CFD23" authority="Bon." authorityName="Bon." box="[192,346,738,762]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FF5F819E326AFD23" box="[192,284,738,762]" italics="true" pageId="10" pageNumber="9">buqueti</emphasis>
Bon.
</taxonomicName>
” (genus, species and author handwritten) / “
<taxonomicName id="4C054D0DFFFD2745FCE9819E3798FD23" authority=", Laporte 1840" authorityName=", Laporte" authorityYear="1840" box="[886,1262,738,762]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B971EA9CFFFD2745FCE9819E374DFD23" box="[886,1083,738,762]" italics="true" pageId="10" pageNumber="9">Phlegon buqueti</emphasis>
,
<bibRefCitation id="EF944B7FFFFD2745FBD6819E3798FD23" author="Laporte, F. L. &amp; Comte de Castelnau" box="[1097,1262,738,762]" pageId="10" pageNumber="9" pagination="254 - 255" refId="ref16338" refString="Laporte, F. L. de (Comte de Castelnau). 1840. Histoire naturelle des insectes Coleopteres, p. 254 - 255. In: P. Dumenil (ed.), Histoire naturelle des animaux articules, annelides, crustaces, arachnids, myriapodes et insectes. P. Dumenil; Paris, France. 564 p." type="book chapter" year="1840">Laporte 1840</bibRefCitation>
</taxonomicName>
, J. Muona det. 2014” / “
<emphasis id="B971EA9CFFFD2745FEBC807D32A6FCC0" bold="true" box="[291,464,769,793]" pageId="10" pageNumber="9">
<typeStatus id="54BE882CFFFD2745FEBC807D32BDFCC0" box="[291,459,769,793]" pageId="10" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
,
<taxonomicName id="4C054D0DFFFD2745FE42807E3136FCC3" authorityName="Laporte" authorityYear="1840" box="[477,576,770,794]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B971EA9CFFFD2745FE42807E3136FCC3" box="[477,576,770,794]" italics="true" pageId="10" pageNumber="9">Phlegon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D0DFFFD2745FDD2807E3050FCC3" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[589,806,770,794]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B971EA9CFFFD2745FDD2807E3196FCC0" box="[589,736,770,793]" italics="true" pageId="10" pageNumber="9">panamensis</emphasis>
, Otto
</taxonomicName>
, det. R.L. Otto, 2016” (
<specimenCount id="9D03FD07FFFD2745FBA9807D373CFCC3" box="[1078,1098,769,794]" pageId="10" pageNumber="9" type="male"></specimenCount>
handwritten behind species name on label) [red printed label].
<typeStatus id="54BE882CFFFD2745FDF4805E31ACFCE3" box="[619,730,802,826]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
deposited in JMC.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF5F801E3078FBE3" blockId="10.[192,1440,865,1082]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF5F801E323BFCA3" bold="true" box="[192,333,866,890]" pageId="10" pageNumber="9">
<typeStatus id="54BE882CFFFD2745FF5F801E323BFCA3" box="[192,333,866,890]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D03FD07FFFD2745FEC4801E32F3FCA3" box="[347,389,865,890]" pageId="10" pageNumber="9" type="male">1 ♂</specimenCount>
, “Barro Colorado I, CZ
<collectingCountry id="F312761EFFFD2745FD3B801E307CFCA0" box="[676,778,866,889]" name="Panama" pageId="10" pageNumber="9">Panama</collectingCountry>
V, 1822 1987, Hank Wolda” (month and dates handwritten) / “TRAP, 1A” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (greenframed white label) / “
<emphasis id="B971EA9CFFFD2745FE5180DD3101FC60" bold="true" box="[462,631,929,953]" pageId="10" pageNumber="9">
<typeStatus id="54BE882CFFFD2745FE5180DD3104FC60" box="[462,626,929,953]" pageId="10" pageNumber="9" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
,
<taxonomicName id="4C054D0DFFFD2745FD1A80DE319EFC63" authorityName="Laporte" authorityYear="1840" box="[645,744,930,954]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B971EA9CFFFD2745FD1A80DE319EFC63" box="[645,744,930,954]" italics="true" pageId="10" pageNumber="9">Phlegon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D0DFFFD2745FD6980DE30A6FC63" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[758,976,930,954]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B971EA9CFFFD2745FD6980DE30FFFC60" box="[758,905,930,953]" italics="true" pageId="10" pageNumber="9">panamensis</emphasis>
, Otto
</taxonomicName>
, det. R.L. Otto, 2016” (
<specimenCount id="9D03FD07FFFD2745FB7480DD3789FC63" box="[1259,1279,929,954]" pageId="10" pageNumber="9" type="male"></specimenCount>
handwritten on label) [yellow printed label];
<specimenCount id="9D03FD07FFFD2745FDDA80BE3119FC03" box="[581,623,961,986]" pageId="10" pageNumber="9" type="female">1 ♀</specimenCount>
, “PAN.
<collectingRegion id="49C1F86CFFFD2745FD4E80BE306EFC03" box="[721,792,962,986]" country="Panama" name="Colon" pageId="10" pageNumber="9">Colon</collectingRegion>
Prv, Ft Espanar vic Marg,
<date id="FFBB104EFFFD2745FBF880BE3789FC03" box="[1127,1279,962,986]" pageId="10" pageNumber="9" value="1992-06-02">2 June 1992</date>
, JE Wappes” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “
<emphasis id="B971EA9CFFFD2745FF44877D32F2FBC0" bold="true" box="[219,388,1025,1049]" pageId="10" pageNumber="9">
<typeStatus id="54BE882CFFFD2745FF44877D3209FBC0" box="[219,383,1025,1049]" pageId="10" pageNumber="9" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
,
<taxonomicName id="4C054D0DFFFD2745FE0A877E328EFBC3" authorityName="Laporte" authorityYear="1840" box="[405,504,1026,1050]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B971EA9CFFFD2745FE0A877E328EFBC3" box="[405,504,1026,1050]" italics="true" pageId="10" pageNumber="9">Phlegon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D0DFFFD2745FD96877E3190FBC3" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[521,742,1026,1050]" class="Insecta" family="Eucnemidae" genus="Phlegon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B971EA9CFFFD2745FD96877E31EAFBC0" box="[521,668,1026,1049]" italics="true" pageId="10" pageNumber="9">panamensis</emphasis>
, Otto
</taxonomicName>
, det. R.L. Otto, 2016” (
<specimenCount id="9D03FD07FFFD2745FB8D877D3750FBC3" box="[1042,1062,1025,1050]" pageId="10" pageNumber="9" type="female"></specimenCount>
handwritten on label) [yellow printed label].
<typeStatus id="54BE882CFFFD2745FEE9875E3285FBE3" box="[374,499,1058,1082]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
are retained in GERP.
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFD2744FF5F871D3058FDE0" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="description">
<paragraph id="8BBA368EFFFD2745FF5F871D30A0FB63" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF5F871D3141FBA3" bold="true" box="[192,567,1121,1146]" pageId="10" pageNumber="9">
Description. Male
<typeStatus id="54BE882CFFFD2745FE28871E3147FBA3" box="[439,561,1122,1146]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
:
</emphasis>
Length
<quantity id="4CFD9B6BFFFD2745FD06871D3072FBA0" box="[665,772,1121,1145]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="9" unit="mm" value="13.5">13.5 mm</quantity>
, width, 4.0 mm. Body stout, oblong, elongate; uniformly reddish brown; antennae reddish brown; femora, tibiae and tarsi reddish brown; head, pronotum and elytra clothed with elongate, semierect yellowish setae (
<figureCitation id="133E2A0BFFFD2745FCEF87DE30BFFB63" box="[880,969,1186,1210]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="10" pageNumber="9">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F87BE3045FAC3" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F87BE3236FB03" bold="true" box="[240,320,1218,1242]" pageId="10" pageNumber="9">Head:</emphasis>
Subspherical; frons convex, with triangular fovea above frontoclypeal region; surfaces shiny, densely punctate; punctures variably sized; apical margin of frontoclypeal region rounded, about 2 times wider than base; mandibles slender, without teeth.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F865D36D6FA43" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F865D3210FAE0" bold="true" box="[240,358,1313,1337]" pageId="10" pageNumber="9">Antenna</emphasis>
(
<figureCitation id="133E2A0BFFFD2745FEE9865E32BCFAE3" box="[374,458,1314,1338]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="10" pageNumber="9">Fig. 21</figureCitation>
): About 2/3 of body length; antennomere III longer than IV; antennomeres IVV slightly longer than wide; antennomeres VIVII quadrate; antennomere VIII wider than long; antennomeres IXX enlarged, pectinate; ramus thinner, at least 1.5 times longer than length of respective segment, arising near apices; antennomere IX shorter than X; antennomere XI simple, elongate, tubular.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F86DE3722FA23" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F86DE32F2FA63" bold="true" box="[240,388,1442,1466]" pageId="10" pageNumber="9">Pronotum:</emphasis>
Surfaces shiny, densely, shallowly punctate; punctures variably sized; slightly wider than long, with moderate, sharp hind angles; parallel-sided at hind angles, gradually narrowing anteriorly above hind angles; disc convex, without circular fovea; base sinuous.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F857D30A9F9C3" blockId="10.[192,1441,1121,1914]" box="[240,991,1537,1562]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F857D32F2F9C0" bold="true" box="[240,388,1537,1561]" pageId="10" pageNumber="9">Scutellum:</emphasis>
Setose, quadrate, elongate and distally rounded.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F855E37C4F980" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F855E3238F9E3" bold="true" box="[240,334,1570,1594]" pageId="10" pageNumber="9">Elytra:</emphasis>
Striae indicated, formed from round punctures; interstices slightly elevated; surfaces shiny, very shallowly punctate; apices (
<figureCitation id="133E2A0BFFFD2745FDCE853E31D1F983" box="[593,679,1602,1626]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="10" pageNumber="9">Fig. 22</figureCitation>
) with shallow cavities near elytral suture.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F851E3122F963" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F851E324EF9A3" bold="true" box="[240,312,1634,1658]" pageId="10" pageNumber="9">Legs:</emphasis>
First tarsomere shorter than combined length of remaining four on mesotarsi; tibiae rounded in cross section; metatarsomeres IIII simple; metatarsomere IV excavated, bilobed; mesotarsomere V elongate; pretarsal claws simple.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F85BE3204F8C3" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F85BE3220F903" bold="true" box="[240,342,1730,1754]" pageId="10" pageNumber="9">Venter:</emphasis>
Closely punctate, with elongate, semierect yellowish setae; hypomeron simple, without lateral antennal grooves; metepisterna parallel-sided; metacoxal plates medially 3.06.0 times wider than laterally.
</paragraph>
<paragraph id="8BBA368EFFFD2745FF6F845D30F3F8A3" blockId="10.[192,1441,1121,1914]" pageId="10" pageNumber="9">
<emphasis id="B971EA9CFFFD2745FF6F845D3203F8E0" bold="true" box="[240,373,1825,1849]" pageId="10" pageNumber="9">Aedeagus</emphasis>
(
<figureCitation id="133E2A0BFFFD2745FE19845E32ABF8E3" box="[390,477,1826,1850]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="10" pageNumber="9">Fig. 23</figureCitation>
): Basal piece short, rounded, dorsally open; remaining parts bulbous, basally narrowed, widest near base of lateral lobes; median lobe short, ventrally curved, dorsally hooked at tip; lateral lobes ventrally curved, elongate, apically rounded.
</paragraph>
<paragraph id="8BBA368EFFFC2744FF5F83BD3058FDE0" blockId="11.[192,1440,193,570]" pageId="11" pageNumber="10">
<emphasis id="B971EA9CFFFC2744FF5F83BD323AFF00" bold="true" box="[192,332,193,217]" pageId="11" pageNumber="10">Variation.</emphasis>
Two
<typeStatus id="54BE882CFFFC2744FE0E83BE317DFF03" box="[401,523,194,218]" pageId="11" pageNumber="10" type="paratype">paratypes</typeStatus>
were examined. The male
<typeStatus id="54BE882CFFFC2744FCC083BE30BBFF03" box="[863,973,194,218]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
was 15.0 mm long and 5.0 mm wide, larger and wider than the
<typeStatus id="54BE882CFFFC2744FD9E839E3118FF23" box="[513,622,226,250]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
. The female
<typeStatus id="54BE882CFFFC2744FC99839E3002FF23" box="[774,884,226,250]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
(
<figureCitation id="133E2A0BFFFC2744FC1B839E30AEFF23" box="[900,984,226,250]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="11" pageNumber="10">Fig. 24</figureCitation>
) was 18.0 mm long and 6.0 mm wide, also larger and wider than the
<typeStatus id="54BE882CFFFC2744FDDE827E31D8FEC3" box="[577,686,258,282]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
. There are no notable differences between the male
<typeStatus id="54BE882CFFFC2744FAAD827E36D6FEC3" box="[1330,1440,258,282]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
and the
<typeStatus id="54BE882CFFFC2744FEB4825E32EEFEE3" box="[299,408,290,314]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
. The female
<typeStatus id="54BE882CFFFC2744FDA2825E31DDFEE3" box="[573,683,290,314]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
differs from the male
<typeStatus id="54BE882CFFFC2744FC53825E3743FEE3" box="[972,1077,290,314]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
in four features. First color, the female
<typeStatus id="54BE882CFFFC2744FED5823E32CEFE83" box="[330,440,322,346]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
is much darker than the
<typeStatus id="54BE882CFFFC2744FD69823E3015FE83" box="[758,867,322,346]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
: reddish brown in the
<typeStatus id="54BE882CFFFC2744FBE6823E3793FE83" box="[1145,1253,322,346]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
, dark brown in the
<typeStatus id="54BE882CFFFC2744FF6F821E3214FEA3" box="[240,354,354,378]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
. Secondly, a pair of deep, circular foveae is present on the pronotal disc of the
<typeStatus id="54BE882CFFFC2744FAAD821E36D6FEA3" box="[1330,1440,354,378]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
(absent in the
<typeStatus id="54BE882CFFFC2744FEE982FE3295FE43" box="[374,483,386,410]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
). Thirdly, the triangular fovea is shallowly indicated above the frontoclypeal region in the
<typeStatus id="54BE882CFFFC2744FEFC82DE32A3FE63" box="[355,469,418,442]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
. Lastly, the female antennal structures (
<figureCitation id="133E2A0BFFFC2744FC5682DE3768FE60" box="[969,1054,417,442]" captionStart="Figures 2025" captionStartId="24.[192,287,1817,1839]" captionTargetBox="[192,1440,192,1775]" captionTargetId="figure-76@24.[192,1440,192,1776]" captionTargetPageId="24" captionText="Figures 2025. Phlegon panamensis sp. nov. 20) Male holotype (JMC), dorsal habitus. 21) Male paratype antenna (GERP), dorsal view. 22) Male holotype elytral apices (JMC), dorsal view. 23) Aedeagus of holotype, dorsal view. 24) Female paratype (GERP), dorsal habitus. 25) Female paratype antenna (GERP), lateral view. (Scale: 20 = 5.0 mm; 2123, 25 = 1.0 mm; 24 = 10.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.5169068" httpUri="https://zenodo.org/record/5169068/files/figure.png" pageId="11" pageNumber="10">Fig. 25</figureCitation>
) differ from either the
<typeStatus id="54BE882CFFFC2744FAA882DE36D6FE63" box="[1335,1440,418,442]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
or male
<typeStatus id="54BE882CFFFC2744FEBB82BE32E0FE03" box="[292,406,450,474]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
. For the female
<typeStatus id="54BE882CFFFC2744FDC282BE31B8FE03" box="[605,718,450,474]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
, antennomere III is slightly longer than IV; antennomeres IVV are longer than wide; antennomeres VIVIII are slightly longer than wide; antennomeres IXX are strongly dentate, subequal, each as long as antennomeres VIIVIII combined; antennomere XI is simple, cylindrical, and longer than either IX or X.
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFC2744FF5F811D37B3FDA3" box="[192,1221,609,634]" pageId="11" pageNumber="10" type="distribution">
<paragraph id="8BBA368EFFFC2744FF5F811D37B3FDA3" blockId="11.[192,1221,609,634]" box="[192,1221,609,634]" pageId="11" pageNumber="10">
<emphasis id="B971EA9CFFFC2744FF5F811D3202FDA0" bold="true" box="[192,372,609,633]" pageId="11" pageNumber="10">Distribution.</emphasis>
This eucnemid has been taken from several localities in the country.
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFC2744FF5F81DD3127FD03" pageId="11" pageNumber="10" type="biology_ecology">
<paragraph id="8BBA368EFFFC2744FF5F81DD3127FD03" blockId="11.[192,1441,673,730]" pageId="11" pageNumber="10">
<emphasis id="B971EA9CFFFC2744FF5F81DD3247FD60" bold="true" box="[192,305,673,697]" pageId="11" pageNumber="10">Biology.</emphasis>
One specimen was taken presumably from a flight intercept trap placed in the Canal Zone. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31F6505FFFC2744FF5F807E3073FCE3" pageId="11" pageNumber="10" type="etymology">
<paragraph id="8BBA368EFFFC2744FF5F807E3073FCE3" blockId="11.[192,1440,770,826]" pageId="11" pageNumber="10">
<emphasis id="B971EA9CFFFC2744FF5F807E322AFCC3" bold="true" box="[192,348,770,794]" pageId="11" pageNumber="10">Etymology.</emphasis>
The specific epithet is derived from a combination of two words, Panama and -ensis, a Latin adjectival suffix meaning “pertaining to”.
</paragraph>
</subSubSection>
</treatment>
</document>