treatments-xml/data/03/9F/F8/039FF85F9676D048B7A42204FCE8F9E2.xml

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<mods:title id="926F21CE0DAA851B3A0B768BB9BF4F7F">A new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from a granitic cavity in Serra da Estrela (Portugal)</mods:title>
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<mods:namePart id="91AB90DE416A7F7DFB85BD7EB5500A38">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="683164B37AA8AB2E2B69EBCFDC748D77">Conde, José</mods:namePart>
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<mods:namePart id="97CF45F0179EFEDAE9B2857D078E779D">Antunes, Sandra</mods:namePart>
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<mods:namePart id="EAAA996FFAB93B97E1D4A5F0A6CB3934">Aguiar, Carlos A. S.</mods:namePart>
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<mods:namePart id="F4249FE18E893A52B04D37C1CBB9877E">Boieiro, Mário</mods:namePart>
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<treatment id="039FF85F9676D048B7A42204FCE8F9E2" ID-DOI="http://doi.org/10.5281/zenodo.6096916" ID-GBIF-Taxon="119631432" ID-Zenodo-Dep="6096916" LSID="urn:lsid:plazi:treatment:039FF85F9676D048B7A42204FCE8F9E2" httpUri="http://treatment.plazi.org/id/039FF85F9676D048B7A42204FCE8F9E2" lastPageId="8" lastPageNumber="409" pageId="2" pageNumber="403">
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<paragraph id="8B8949499676D042B7A42204FC5CF95A" blockId="2.[151,917,1659,1719]" box="[151,917,1659,1686]" pageId="2" pageNumber="403">
<heading id="D0C1FE259676D042B7A42204FC5CF95A" bold="true" box="[151,917,1659,1686]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B942955B9676D042B7A42204FC5CF95A" bold="true" box="[151,917,1659,1686]" pageId="2" pageNumber="403">
<taxonomicName id="4C3632CA9676D042B7A42204FD31F95A" ID-CoL="3796R" authority="Serrano &amp; Boieiro" authorityName="Serrano &amp; Boieiro" box="[151,760,1659,1686]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov." subGenus="Lathromene">
<emphasis id="B942955B9676D042B7A42204FF35F959" bold="true" box="[151,252,1659,1685]" italics="true" pageId="2" pageNumber="403">Domene</emphasis>
(
<emphasis id="B942955B9676D042B63E2204FE68F959" bold="true" box="[269,417,1659,1685]" italics="true" pageId="2" pageNumber="403">Lathromene</emphasis>
) 
<emphasis id="B942955B9676D042B6812204FDC1F959" bold="true" box="[434,520,1659,1685]" italics="true" pageId="2" pageNumber="403">viriatoi</emphasis>
Serrano &amp; Boieiro
</taxonomicName>
, 
<taxonomicNameLabel id="A27128209676D042B4362203FC5CF95A" box="[773,917,1660,1686]" pageId="2" pageNumber="403" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8949499676D042B7A422E1FEB2F97B" blockId="2.[151,917,1659,1719]" box="[151,379,1694,1719]" pageId="2" pageNumber="403">
<emphasis id="B942955B9676D042B7A422E1FF69F97B" bold="true" box="[151,160,1694,1719]" pageId="2" pageNumber="403">(</emphasis>
<figureCitation id="130D55CC9676D042B79322E1FF2EF97B" box="[160,231,1694,1719]" captionStart="FIGURE 2" captionStartId="3.[151,250,1067,1089]" captionTargetBox="[151,1435,193,1045]" captionTargetId="figure@3.[151,1435,193,1046]" captionTargetPageId="3" captionText="FIGURE 2. Domene viriatoi n. sp. adult." httpUri="https://zenodo.org/record/240968/files/figure.png" pageId="2" pageNumber="403">Figs 2</figureCitation>
, 
<figureCitation id="130D55CC9676D042B7C022E0FECAF97B" box="[243,259,1695,1719]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="2" pageNumber="403">3</figureCitation>
, 
<figureCitation id="130D55CC9676D042B63C22E0FED6F97B" box="[271,287,1695,1719]" captionStart="FIGURE 4" captionStartId="5.[151,250,1143,1165]" captionText="FIGURE 4. Head of Domene viriatoi n. sp. (ventral view)." pageId="2" pageNumber="403">4</figureCitation>
, 
<figureCitation id="130D55CC9676D042B61822E1FEF2F97B" box="[299,315,1694,1719]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="2" pageNumber="403">5</figureCitation>
, 
<figureCitation id="130D55CC9676D042B67422E1FE9EF97B" box="[327,343,1694,1719]" captionStart="FIGURE 6" captionStartId="7.[151,250,1152,1174]" captionTargetBox="[169,1407,488,1107]" captionTargetId="figure@7.[151,1436,481,1131]" captionTargetPageId="7" captionText="FIGURE 6. Genital segment of Domene viriatoi n. sp. (ventral view): (a), male; (b), female." httpUri="https://zenodo.org/record/240971/files/figure.png" pageId="2" pageNumber="403">6</figureCitation>
, 
<figureCitation id="130D55CC9676D042B65022E0FEBBF97B" box="[355,370,1695,1719]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="2" pageNumber="403">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C1AC29676D042B7A4229EFF3DF862" pageId="2" pageNumber="403" type="materials_examined">
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<emphasis id="B942955B9676D042B7A4229EFE9CF936" bold="true" box="[151,341,1761,1786]" pageId="2" pageNumber="403">
<typeStatus id="548DF7EB9676D042B7A4229EFF1BF936" box="[151,210,1761,1786]" pageId="2" pageNumber="403">Type</typeStatus>
material:
</emphasis>
<typeStatus id="548DF7EB9676D042B659229EFE1CF936" box="[362,469,1761,1786]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
, ♂, 
<collectingCountry id="F32109D99676D042B52C229EFD48F936" box="[543,641,1761,1786]" name="Portugal" pageId="2" pageNumber="403">Portugal</collectingCountry>
, Buraco da Moura (Lapa dos Dinheiros) (U.T.M. coordinate: 29TPE103714), 
<date id="FF886F899676D042B6622379FE3DF8D2" box="[337,500,1798,1822]" pageId="2" pageNumber="403" value="2014-10-13" valueMax="2014-10-24" valueMin="2014-10-13">13‒24.X.2014</date>
, (J. Conde, M. Boieiro &amp; S. Antunes leg.). 
<typeStatus id="548DF7EB9676D042B4D72379FB9DF8D2" box="[996,1108,1798,1822]" pageId="2" pageNumber="403" type="paratype">Paratypes</typeStatus>
, same locality of 
<typeStatus id="548DF7EB9676D042B22E237AFA4BF8D2" box="[1309,1410,1797,1822]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D3082C09676D042B2BD2379FF67F88D" pageId="2" pageNumber="403" type="generic">1 ♂</specimenCount>
, 
<date id="FF886F899676D042B78F2355FE98F88E" box="[188,337,1834,1858]" pageId="2" pageNumber="403" value="2013-10-07" valueMax="2013-10-31" valueMin="2013-10-07">7‒31.X.2013</date>
, 
<specimenCount id="9D3082C09676D042B66C2355FE4AF88E" box="[351,387,1834,1858]" pageId="2" pageNumber="403" type="generic">1♂</specimenCount>
, 
<date id="FF886F899676D042B6A12355FDB6F88E" box="[402,639,1834,1858]" pageId="2" pageNumber="403" value="2014-07-19" valueMax="2014-08-17" valueMin="2014-07-19">19.VII‒17.VIII.2014</date>
, (J. Conde &amp; S. Antunes leg.), 
<specimenCount id="9D3082C09676D042B4DF2355FBC6F88E" box="[1004,1039,1834,1858]" pageId="2" pageNumber="403" type="generic">2♂</specimenCount>
(
<specimenCount id="9D3082C09676D042B3112355FB8CF88E" box="[1058,1093,1834,1858]" pageId="2" pageNumber="403" type="generic">1♂</specimenCount>
gold coated), 
<date id="FF886F899676D042B3DF2355FAACF88E" box="[1260,1381,1834,1858]" pageId="2" pageNumber="403" value="2014-10-13">13.X.2014</date>
, 3♀, 
<date id="FF886F899676D042B7A42331FEF3F8AA" box="[151,314,1870,1894]" pageId="2" pageNumber="403" value="2014-10-13" valueMax="2014-10-24" valueMin="2014-10-13">13‒24.X.2014</date>
, (J. Conde, M. Boieiro &amp; S. Antunes leg.). The 
<typeStatus id="548DF7EB9676D042B4A22332FC3AF8AA" box="[913,1011,1869,1894]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
and 
<typeStatus id="548DF7EB9676D042B3052331FB6BF8AA" box="[1078,1186,1870,1894]" pageId="2" pageNumber="403" type="paratype">paratypes</typeStatus>
are deposited in the collection of the first author, at Departamento de Biologia Animal (Faculdade de Ciências da Universidade de Lisboa).
</paragraph>
</subSubSection>
<caption id="DF4919C19677D043B7A42054FDF7FB8D" box="[151,574,1067,1089]" httpUri="https://zenodo.org/record/240968/files/figure.png" pageId="3" pageNumber="404" targetBox="[151,1435,193,1045]" targetPageId="3">
<paragraph id="8B8949499677D043B7A42054FDF7FB8D" blockId="3.[151,574,1067,1089]" box="[151,574,1067,1089]" pageId="3" pageNumber="404">
<emphasis id="B942955B9677D043B7A42054FEDBFB8D" bold="true" box="[151,274,1067,1089]" pageId="3" pageNumber="404">FIGURE 2.</emphasis>
<taxonomicName id="4C3632CA9677D043B62A2053FE77FB8D" box="[281,446,1068,1089]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9677D043B62A2053FE77FB8D" box="[281,446,1068,1089]" italics="true" pageId="3" pageNumber="404">Domene viriatoi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A27128209677D043B6F72054FE37FB8D" box="[452,510,1067,1089]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
adult.
</paragraph>
</caption>
<subSubSection id="C32C1AC29677D043B7F42017FA58FB21" pageId="3" pageNumber="404" type="etymology">
<paragraph id="8B8949499677D043B7F42017FA58FB21" blockId="3.[151,1437,1128,2028]" pageId="3" pageNumber="404">
<emphasis id="B942955B9677D043B7F42017FE87FB4D" bold="true" box="[199,334,1128,1153]" pageId="3" pageNumber="404">Etymology.</emphasis>
The new species is dedicated to Viriato known as a Benefactor and Savior by the Lusitanians. He was originally from the high Montes Hermínios, nowadays Serra da Estrela, and, according to some theories, he became a soldier after being shepherd and a huntsman. Viriato is known as the greatest military leader of the Lusitanians, having won many battles against the Roman occupation being named after this “ 
<emphasis id="B942955B9677D043B39F20A9FAB4FB21" box="[1196,1405,1238,1261]" italics="true" pageId="3" pageNumber="404">Terror romanorum</emphasis>
”.
</paragraph>
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<subSubSection id="C32C1AC29677D048B7F42087FCE8F9E2" lastPageId="8" lastPageNumber="409" pageId="3" pageNumber="404" type="description">
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<emphasis id="B942955B9677D043B7F42087FE90FADD" bold="true" box="[199,345,1272,1297]" pageId="3" pageNumber="404">Description.</emphasis>
Microphtalmous, wingless and elongate body (
<figureCitation id="130D55CC9677D043B44E2087FC0CFADD" box="[893,965,1272,1297]" captionStart="FIGURE 2" captionStartId="3.[151,250,1067,1089]" captionTargetBox="[151,1435,193,1045]" captionTargetId="figure@3.[151,1435,193,1046]" captionTargetPageId="3" captionText="FIGURE 2. Domene viriatoi n. sp. adult." httpUri="https://zenodo.org/record/240968/files/figure.png" pageId="3" pageNumber="404">Fig. 2</figureCitation>
). Coloration: body reddish and abdomen dark-reddish. Body length: 
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(males), 
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(females).
</paragraph>
<paragraph id="8B8949499677D043B7F4213FFA54F84C" blockId="3.[151,1437,1128,2028]" pageId="3" pageNumber="404">
<emphasis id="B942955B9677D043B7F4213FFECEFA95" bold="true" box="[199,263,1344,1369]" pageId="3" pageNumber="404">Head</emphasis>
(
<figureCitation id="130D55CC9677D043B624213FFE96FA95" box="[279,351,1344,1369]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Figs 3</figureCitation>
a, 4): subquadrate/orbicular, 1.1 times longer than wide (males and females) [length: 
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(males) and 
<quantity id="4CCEE4AC9677D043B66F211BFDCBFAB0" box="[348,514,1380,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.31" metricValueMin="1.15" pageId="3" pageNumber="404" unit="mm" value="1.23" valueMax="1.31" valueMin="1.15">1.15‒1.31 mm</quantity>
(females), width: 
<quantity id="4CCEE4AC9677D043B5E4211AFCB5FAB0" box="[727,892,1381,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.18" metricValueMin="1.1" pageId="3" pageNumber="404" unit="mm" value="1.14" valueMax="1.18" valueMin="1.1">1.10‒1.18 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9677D043B326211AFB72FAB0" box="[1045,1211,1381,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="404" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.10‒1.20 mm</quantity>
(females)], slightly wider than pronotum (1.1‒1.2 times); eyes reduced, with about 22 ommatidia (
<figureCitation id="130D55CC9677D043B31A21F7FBA7FA6D" box="[1065,1134,1416,1441]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
b); long and thin seta in a supraocular, small circular depression; a patch of yellowish minor pubescence above the depression and behind the eyes (
<figureCitation id="130D55CC9677D043B7EA21AFFED5FA25" box="[217,284,1488,1513]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
b); two dark spots in the vertex signal the insertions of the dorsal arms of the tentorium; gular sutures well defined, converging towards neck (
<figureCitation id="130D55CC9677D043B55B218BFD79F9C1" box="[616,688,1524,1549]" captionStart="FIGURE 4" captionStartId="5.[151,250,1143,1165]" captionText="FIGURE 4. Head of Domene viriatoi n. sp. (ventral view)." pageId="3" pageNumber="404">Fig. 4</figureCitation>
); dorsal surface without any isodiametric microreticulation, with well-defined dense punctuation, similar to that of pronotum, on lateral sides and behind vertex; vertex and front with sparse punctuation, almost smooth (
<figureCitation id="130D55CC9677D043B5582243FD66F999" box="[619,687,1596,1621]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
a); antennae filiform (
<figureCitation id="130D55CC9677D043B49A2243FC27F999" box="[937,1006,1596,1621]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
d), 
<quantity id="4CCEE4AC9677D043B3262242FBA5F998" box="[1045,1132,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="404" unit="mm" value="2.7">2.7 mm</quantity>
long, reaching the base of pronotum when directed backwards; all antennomeres longer than wide, the 1st longest of all [length: 
<quantity id="4CCEE4AC9677D043B21F221CFF0AF96E" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.25" metricValueMin="1.13" pageId="3" pageNumber="404" unit="mm" value="1.19" valueMax="1.25" valueMin="1.13">1.13‒1.25 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9677D043B66E22F5FDCAF96E" box="[349,515,1674,1698]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.29" metricValueMin="1.21" pageId="3" pageNumber="404" unit="mm" value="1.25" valueMax="1.29" valueMin="1.21">1.21‒1.29 mm</quantity>
(females)]; 3rd antennomere in average 1.2-1.3 times longer than the 4th and 5th antennomeres; the 6th to 10th antennomeres are subequal and the 11th is 1.3‒1.4 times longer than the five precedents; labrum deeply emarginated with 9‒11 large setae spread over the disk and the anterior margin (
<figureCitation id="130D55CC9677D043B20E22ABFAB6F921" box="[1341,1407,1748,1773]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
c); mandibles symmetrical with four distinct teeth in the inner edge, two of them are more pronounced (
<figureCitation id="130D55CC9677D043B2772286FA47F8DD" box="[1348,1422,1785,1810]" captionStart="FIGURE 4" captionStartId="5.[151,250,1143,1165]" captionText="FIGURE 4. Head of Domene viriatoi n. sp. (ventral view)." pageId="3" pageNumber="404">Fig. 4</figureCitation>
); maxillary palpus with four articles, second and third subequal, more than 3 times as long as broad, apical joint reduced and conical; internal lobe of maxilla with clusters of setae; labium with bilobed glossae, circumscribed by hairy paraglossae; labial palpus with three joints, the 2nd being the longest and more robust, and the 3rd the thinnest.
</paragraph>
<paragraph id="8B8949499677D045B7F423F4FE8EFAD7" blockId="3.[151,1437,1128,2028]" lastBlockId="5.[151,1437,1210,2027]" lastPageId="5" lastPageNumber="406" pageId="3" pageNumber="404">
<emphasis id="B942955B9677D043B7F423F4FE89F868" bold="true" box="[199,320,1931,1956]" pageId="3" pageNumber="404">Pronotum</emphasis>
(
<figureCitation id="130D55CC9677D043B66023F4FE52F868" box="[339,411,1931,1956]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="3" pageNumber="404">
Fig. 
<quantity id="4CCEE4AC9677D043B6BF23F3FE52F868" box="[396,411,1932,1956]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="3" pageNumber="404" unit="g" value="3.0">3</quantity>
</figureCitation>
g): 1.3 times longer than wide (males and females) [length: 
<quantity id="4CCEE4AC9677D043B35123F3FAC3F86F" box="[1122,1290,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" metricValueMax="1.31" metricValueMin="1.23" pageId="3" pageNumber="404" unit="mm" value="1.27" valueMax="1.31" valueMin="1.23">1.23‒1.31 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9677D043B7A423CFFEF0F80B" box="[151,313,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" metricValueMax="1.34" metricValueMin="1.22" pageId="3" pageNumber="404" unit="mm" value="1.28" valueMax="1.34" valueMin="1.22">1.22‒1.34 mm</quantity>
(females), width: 
<quantity id="4CCEE4AC9677D043B53523CFFD60F80B" box="[518,681,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" metricValueMax="1.04" metricValueMin="0.98" pageId="3" pageNumber="404" unit="mm" value="1.01" valueMax="1.04" valueMin="0.98">0.98‒1.04 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9677D043B40A23CFFC15F80B" box="[825,988,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.07" metricValueMin="0.94" pageId="3" pageNumber="404" unit="mm" value="1.005" valueMax="1.07" valueMin="0.94">0.94‒1.07 mm</quantity>
(females)], slightly narrower than head, widest anteriorly and distinctly tapering posteriad; anterior angles weakened, posterior angles marked but largely obtuse; basal margin well defined; punctuation similar to that of head, evenly distributed except for a smooth midline ranging from the anterior to the posterior parts; absence of any middle sulcus, interstices without microsculpture.
</paragraph>
<caption id="DF4919C19670D044B7A4232BFEEDF80A" httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="4" pageNumber="405" targetBox="[188,1398,193,1853]" targetPageId="4">
<paragraph id="8B8949499670D044B7A4232BFEEDF80A" blockId="4.[151,1436,1876,1990]" pageId="4" pageNumber="405">
<emphasis id="B942955B9670D044B7A4232BFEDBF8A6" bold="true" box="[151,274,1876,1898]" pageId="4" pageNumber="405">FIGURE 3.</emphasis>
SEM photographs of structural features of 
<taxonomicName id="4C3632CA9670D044B5F4232AFCA4F8A6" box="[711,877,1877,1898]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9670D044B5F4232AFCA4F8A6" box="[711,877,1877,1898]" italics="true" pageId="4" pageNumber="405">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9670D044B441232BFC6CF8A6" bold="true" box="[882,933,1876,1898]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="A27128209670D044B441232BFC6CF8A6" box="[882,933,1876,1898]" pageId="4" pageNumber="405" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B942955B9670D044B481232BFC19F8A6" bold="true" box="[946,976,1876,1898]" pageId="4" pageNumber="405">(a)</emphasis>
, head (dorsal view); 
<emphasis id="B942955B9670D044B397232BFB0DF8A6" bold="true" box="[1188,1220,1876,1898]" pageId="4" pageNumber="405">(b)</emphasis>
, detail of the eye and chaetotaxy (lateral view); 
<emphasis id="B942955B9670D044B69B230CFE0DF845" bold="true" box="[424,452,1907,1929]" pageId="4" pageNumber="405">(c)</emphasis>
, labrum (dorsal view); 
<emphasis id="B942955B9670D044B58E230CFD15F845" bold="true" box="[701,732,1907,1929]" pageId="4" pageNumber="405">(d)</emphasis>
, antennae (dorsal view); 
<emphasis id="B942955B9670D044B4D4230CFBCAF845" bold="true" box="[999,1027,1907,1929]" pageId="4" pageNumber="405">(e)</emphasis>
, part of the cleaning organ in foretibia (latero‒ventral view); 
<emphasis id="B942955B9670D044B64A23EEFE5BF86B" bold="true" box="[377,402,1937,1959]" pageId="4" pageNumber="405">(f)</emphasis>
, part of the cleaning organ in forefemur (latero‒ventral view); 
<emphasis id="B942955B9670D044B32323EEFBE7F86B" bold="true" box="[1040,1070,1937,1959]" pageId="4" pageNumber="405">(g)</emphasis>
, pronotum (dorsal view); 
<emphasis id="B942955B9670D044B20423EEFA9FF86B" bold="true" box="[1335,1366,1937,1959]" pageId="4" pageNumber="405">(h)</emphasis>
, elytra (dorsal view).
</paragraph>
</caption>
<caption id="DF4919C19671D045B7A42008FD38FB41" box="[151,753,1143,1165]" pageId="5" pageNumber="406">
<paragraph id="8B8949499671D045B7A42008FD38FB41" blockId="5.[151,753,1143,1165]" box="[151,753,1143,1165]" pageId="5" pageNumber="406">
<emphasis id="B942955B9671D045B7A42008FEDBFB41" bold="true" box="[151,274,1143,1165]" pageId="5" pageNumber="406">FIGURE 4.</emphasis>
Head of 
<taxonomicName id="4C3632CA9671D045B6432007FDDFFB41" box="[368,534,1144,1165]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9671D045B6432007FDDFFB41" box="[368,534,1144,1165]" italics="true" pageId="5" pageNumber="406">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9671D045B52F2008FD9CFB41" bold="true" box="[540,597,1143,1165]" pageId="5" pageNumber="406">
<taxonomicNameLabel id="A27128209671D045B52F2008FD9CFB41" box="[540,597,1143,1165]" pageId="5" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(ventral view).
</paragraph>
</caption>
<paragraph id="8B8949499671D045B7F42159FC16FA3F" blockId="5.[151,1437,1210,2027]" pageId="5" pageNumber="406">
<emphasis id="B942955B9671D045B7F42159FEDAFAF3" bold="true" box="[199,275,1318,1343]" pageId="5" pageNumber="406">Elytra</emphasis>
(
<figureCitation id="130D55CC9671D045B6152159FEA4FAF3" box="[294,365,1318,1343]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="5" pageNumber="406">Fig. 3</figureCitation>
h): 1.2 times wider than long [length: 
<quantity id="4CCEE4AC9671D045B4072158FC12FAF2" box="[820,987,1319,1343]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.2" metricValueMin="7.4" pageId="5" pageNumber="406" unit="mm" value="0.78" valueMax="0.82" valueMin="0.74">0.74‒0.82 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9671D045B34A2159FAE9FAF2" box="[1145,1312,1318,1343]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.299999999999999" metricValueMin="7.5" pageId="5" pageNumber="406" unit="mm" value="0.79" valueMax="0.83" valueMin="0.75">0.75‒0.83 mm</quantity>
(females), width: 
<quantity id="4CCEE4AC9671D045B7DE2135FE5FFAAE" box="[237,406,1354,1378]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="9.7" metricValueMin="9.3" pageId="5" pageNumber="406" unit="mm" value="0.95" valueMax="0.97" valueMin="0.93">0.93‒0.97 mm</quantity>
(males) and 
<quantity id="4CCEE4AC9671D045B50E2135FD2EFAAE" box="[573,743,1354,1378]" metricMagnitude="-4" metricUnit="m" metricValue="9.35" metricValueMax="9.9" metricValueMin="8.8" pageId="5" pageNumber="406" unit="mm" value="0.935" valueMax="0.99" valueMin="0.88">0.88‒0.99 mm</quantity>
(females)], approximately 0.6 times shorter than head or pronotum [average elytra length/head length: 0.63 (males and females), average elytra length/pronotum length: 0.61 (males) and 0.62 (females)]; only slightly narrower than head (males and females: 0.8 times) or pronotum (males: 0.9 times, females: 1.0); flat in dorsal view; humeral angles obsolete; lateral margins slightly divergent posteriad; surface with coarse, rugose and wrinkled punctuation. Wingless.
</paragraph>
<paragraph id="8B8949499671D045B7F42181FBBBF827" blockId="5.[151,1437,1210,2027]" pageId="5" pageNumber="406">
<emphasis id="B942955B9671D045B7F42181FEF5F9DB" bold="true" box="[199,316,1534,1559]" pageId="5" pageNumber="406">Abdomen</emphasis>
(
<figureCitation id="130D55CC9671D045B67D2181FE53F9DB" box="[334,410,1534,1559]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="5" pageNumber="406">Figs 5</figureCitation>
, 
<figureCitation id="130D55CC9671D045B69A2181FE73F9DB" box="[425,442,1534,1559]" captionStart="FIGURE 6" captionStartId="7.[151,250,1152,1174]" captionTargetBox="[169,1407,488,1107]" captionTargetId="figure@7.[151,1436,481,1131]" captionTargetPageId="7" captionText="FIGURE 6. Genital segment of Domene viriatoi n. sp. (ventral view): (a), male; (b), female." httpUri="https://zenodo.org/record/240971/files/figure.png" pageId="5" pageNumber="406">6</figureCitation>
): Maximum width at segment VI, approximately 1.3 times wider than elytra; tergites IV‒VI with fine and dense punctuation within a microsculpture of transverse meshes, these more pronounced in the intersegmental membranes; sternite III (males and females) with a middle carina in the first two thirds surrounded in both sides by a small depression to accommodate the resting metacoxae, another minor carina in the middle of the posterior intersegmental membrane (
<figureCitation id="130D55CC9671D045B54F22F1FD70F96B" box="[636,697,1678,1703]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="5" pageNumber="406">Fig.5</figureCitation>
a); sternites III‒VI with fine and dense punctuation within a microsculpture of transverse meshes, substituted by irregular rows of scale-shaped microtrichia anteriad near the intersegmental membranes and by regular posteriad quadrate meshes (e.g. 
<figureCitation id="130D55CC9671D045B4EB22A9FBD3F923" box="[984,1050,1750,1775]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="5" pageNumber="406">Fig. 5</figureCitation>
b). Male: sternite VII with shallow median impression of triangular shape posteriorly, with some lateral scattered short, and stout striated setae, some of them black (2‒5), interspersed among normal setae (
<figureCitation id="130D55CC9671D045B4212361FC91F8FB" box="[786,856,1822,1847]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="5" pageNumber="406">Fig. 5</figureCitation>
c); sternite VIII transverse, anteriorly with shallow median impression of semicircular shape, posteriorly with a distinct incision in an inverted U‒shaped surrounded by a small glabrous area with a microsculpture of transverse meshes; on either side of the apex of this impression there is a cluster of 15‒21 modified, short and stout black striated setae (
<figureCitation id="130D55CC9671D045B4D823F5FBFAF86F" box="[1003,1075,1930,1955]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="5" pageNumber="406">Figs 5</figureCitation>
e‒5f, 6a), looking like a brush; sternite IX slightly arcuate apically; genital segment in ventral view as in figure 6a. Female: genital segment (dorsal view) with a large glabrous area in the center and in ventral view as in figure 6b.
</paragraph>
<caption id="DF4919C19672D046B7A421D2FA5CF9D3" httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="6" pageNumber="407" targetBox="[188,1398,193,1431]" targetPageId="6">
<paragraph id="8B8949499672D046B7A421D2FA5CF9D3" blockId="6.[151,1436,1453,1567]" pageId="6" pageNumber="407">
<emphasis id="B942955B9672D046B7A421D2FEDBFA0F" bold="true" box="[151,274,1453,1475]" pageId="6" pageNumber="407">FIGURE 5.</emphasis>
SEM photographs of structural features of 
<taxonomicName id="4C3632CA9672D046B5F521D1FCA2FA0F" box="[710,875,1454,1475]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9672D046B5F521D1FCA2FA0F" box="[710,875,1454,1475]" italics="true" pageId="6" pageNumber="407">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9672D046B44221D2FC63FA0F" bold="true" box="[881,938,1453,1475]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="A27128209672D046B44221D2FC63FA0F" box="[881,938,1453,1475]" pageId="6" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B942955B9672D046B48521D2FC1DFA0F" bold="true" box="[950,980,1453,1475]" pageId="6" pageNumber="407">(a)</emphasis>
, III abdominal sternite (latero‒ventral view); 
<emphasis id="B942955B9672D046B7A421B3FF7FFA2E" bold="true" box="[151,182,1484,1506]" pageId="6" pageNumber="407">(b)</emphasis>
, detail of intersegmental area between III and IV abdominal sternites (latero‒ventral view); 
<emphasis id="B942955B9672D046B36821B3FBBEFA2E" bold="true" box="[1115,1143,1484,1506]" pageId="6" pageNumber="407">(c)</emphasis>
, VII, VIII and IX abdominal sternites (ventral view); 
<emphasis id="B942955B9672D046B6BD2195FE64F9CC" bold="true" box="[398,429,1514,1536]" pageId="6" pageNumber="407">(d)</emphasis>
, detail of middle posteriad region of VII abdominal sternite (male, ventral view); 
<emphasis id="B942955B9672D046B3DE2195FAC0F9CC" bold="true" box="[1261,1289,1514,1536]" pageId="6" pageNumber="407">(e)</emphasis>
, detail of VIII abdominal sternite (male, ventral view); 
<emphasis id="B942955B9672D046B5002276FD85F9D3" bold="true" box="[563,588,1545,1567]" pageId="6" pageNumber="407">(f)</emphasis>
, detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view).
</paragraph>
</caption>
<paragraph id="8B8949499672D046B7F42239FF2EF94F" blockId="6.[151,1437,1606,2027]" pageId="6" pageNumber="407">
<emphasis id="B942955B9672D046B7F42239FF36F993" bold="true" box="[199,255,1606,1631]" pageId="6" pageNumber="407">Legs</emphasis>
: Long and slender; forelegs with antennal cleaning organ following the general pattern of the genus (
<figureCitation id="130D55CC9672D046B2592239FF6DF94F" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="6" pageNumber="407">Figs 3</figureCitation>
e‒3f).
</paragraph>
<paragraph id="8B8949499672D046B7F422F1FABFF8FB" blockId="6.[151,1437,1606,2027]" pageId="6" pageNumber="407">
<emphasis id="B942955B9672D046B7F422F1FEF3F96B" bold="true" box="[199,314,1678,1703]" pageId="6" pageNumber="407">Aedeagus</emphasis>
(
<figureCitation id="130D55CC9672D046B67E22F1FE5FF96B" box="[333,406,1678,1703]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
): Sclerotized ventral blade exceeding largely the edge of the aedeagus (
<figureCitation id="130D55CC9672D046B3D622F1FAE5F96B" box="[1253,1324,1678,1703]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
a), with a straight apex, slightly truncate in the middle, in each side with a small tooth, and a small lateral tooth‒shaped expansion in midway between the apex and the base (
<figureCitation id="130D55CC9672D046B5D822A9FCE7F923" box="[747,814,1750,1775]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
a); fused lateral lobes (
<figureCitation id="130D55CC9672D046B31822A9FBA4F923" box="[1067,1133,1750,1775]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
c); ellipse‒shaped in dorsal view, presenting a V‒shaped sclerotized blade covering the upper part and the apical edge of the aedeagus (
<figureCitation id="130D55CC9672D046B25C2285FF6FF8FB" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
b); median lobe with two internal small finger‒like structures and one plate of scale in each side (
<figureCitation id="130D55CC9672D046B3DB2361FAE6F8FB" box="[1256,1327,1822,1847]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="6" pageNumber="407">Figs 7</figureCitation>
a‒7b).
</paragraph>
<paragraph id="8B8949499672D047B7F4233DFD54FE60" blockId="6.[151,1437,1606,2027]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
<emphasis id="B942955B9672D046B7F4233DFE77F897" bold="true" box="[199,446,1858,1883]" pageId="6" pageNumber="407">Taxonomic remarks.</emphasis>
According to the taxonomic criteria defined by 
<bibRefCitation id="EFA734B89672D046B4D0233DFB46F897" author="Coiffait" box="[995,1167,1858,1883]" pageId="6" pageNumber="407" refString="Coiffait, H. (1982) Coleopteres Staphylinidae de la region palearctique occidentale. IV. Sous famille Paederinae. Tribu Paederini 1 (Paederi, Lathrobii). Supplement a la Nouvelle Revue d'Entomologie, 12 (4), 1 - 440." type="journal article" year="1982">Coiffait (1982)</bibRefCitation>
, 
<taxonomicName id="4C3632CA9672D046B3AF233CFA9DF896" box="[1180,1364,1859,1882]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9672D046B3AF233CFA9DF896" box="[1180,1364,1859,1882]" italics="true" pageId="6" pageNumber="407">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9672D046B26F233DFA55F897" bold="true" box="[1372,1436,1858,1883]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="A27128209672D046B26F233DFA55F897" box="[1372,1436,1858,1883]" pageId="6" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
should be included in the subgenus 
<taxonomicName id="4C3632CA9672D046B51A2319FD66F8B3" box="[553,687,1894,1919]" class="Insecta" family="Staphylinidae" genus="Lathromene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B942955B9672D046B51A2319FD66F8B3" box="[553,687,1894,1919]" italics="true" pageId="6" pageNumber="407">Lathromene</emphasis>
</taxonomicName>
. The species shows distinctive morphological characteristics from the other congeners of this subgenus, namely the shape of the aedeagus and the characters of the male sternite VIII. Taking into account the pieces of aedeagus, the new species can be easily segregated from 
<taxonomicName id="4C3632CA9672D046B3E323D0FAA6F80B" box="[1232,1391,1966,1991]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="scopaeella">
<emphasis id="B942955B9672D046B3E323D0FAA6F80B" box="[1232,1391,1966,1991]" italics="true" pageId="6" pageNumber="407">D. scopaeella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3632CA9672D046B2B223D0FE63F827" authority="Outerelo, 1977" authorityName="Outerelo" authorityYear="1977" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B942955B9672D046B2B223D0FF2FF827" italics="true" pageId="6" pageNumber="407">D. subiasi</emphasis>
(Outerelo, 1977)
</taxonomicName>
, 
<taxonomicName id="4C3632CA9672D046B68523ACFC65F827" authority="Feldmann &amp; Hernando, 2005" authorityName="Feldmann &amp; Hernando" authorityYear="2005" box="[438,940,2002,2027]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="gallaeciana">
<emphasis id="B942955B9672D046B68523ACFD94F827" box="[438,605,2002,2027]" italics="true" pageId="6" pageNumber="407">D. gallaeciana</emphasis>
<bibRefCitation id="EFA734B89672D046B55023ADFC65F827" author="Feldmann" box="[611,940,2002,2027]" pageId="6" pageNumber="407" refString="Feldmann, B. &amp; Hernando, C. (2005) Two new species of Domene Fauvel 1873 from Spain, with a new combination and a catalogue of the Iberian species of the genus (Coleoptera: Staphylinidae, Paederinae). Linzer Biologische Beitrage, 37 (1), 399 - 406." type="journal article" year="2005">Feldmann &amp; Hernando, 2005</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3632CA9672D047B4D123ACFF1BFF7C" authority="Outerelo &amp; Gamarra, 2012" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" lastPageNumber="408" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B942955B9672D046B4D123ACFB6FF827" box="[994,1190,2002,2027]" italics="true" pageId="6" pageNumber="407">D. barraganensis</emphasis>
<bibRefCitation id="EFA734B89672D047B39E23ADFF1BFF7C" author="Outerelo" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" refString="Outerelo, R. &amp; Gamarra, P. (2012) Domene (Lathromene) barraganensis, nueva especie de estafilinido de la Peninsula Iberica (Coleoptera: Staphylinidae, Paederinae). Arquivos Entomoloxicos, 7, 123 - 128." type="journal article" year="2012">Outerelo &amp; Gamarra, 2012</bibRefCitation>
</taxonomicName>
, by the symmetry 
<typeStatus id="548DF7EB9673D047B68124E7FE2BFF7C" box="[434,482,152,176]" pageId="7" pageNumber="408">type</typeStatus>
of ventral blade (symmetric in the new species 
<emphasis id="B942955B9673D047B31124E6FBF0FF7C" box="[1058,1081,153,176]" italics="true" pageId="7" pageNumber="408">vs</emphasis>
. asymmetric in the other four species). Among the species with symmetric ventral blade, the closest species to 
<taxonomicName id="4C3632CA9673D047B30624C2FB65FF18" box="[1077,1196,189,212]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9673D047B30624C2FB65FF18" box="[1077,1196,189,212]" italics="true" pageId="7" pageNumber="408">D. viriatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27128209673D047B38724C3FB3DFF18" box="[1204,1268,188,213]" pageId="7" pageNumber="408" rank="species">
<emphasis id="B942955B9673D047B38724C3FB24FF19" bold="true" box="[1204,1261,188,213]" pageId="7" pageNumber="408">n. sp</emphasis>
.
</taxonomicNameLabel>
seem to be 
<taxonomicName id="4C3632CA9673D047B2B224C2FDE7FF34" authority="Reboleira &amp; Oromi, 2011" authorityName="Reboleira &amp; Oromi" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="B942955B9673D047B2B224C2FECFFF34" italics="true" pageId="7" pageNumber="408">D. lusitanica</emphasis>
Reboleira &amp; Oromi, 2011
</taxonomicName>
and 
<taxonomicName id="4C3632CA9673D047B555249EFB64FF34" authority="Outerelo, Gamarra &amp; Salgado, 2000" authorityName="Outerelo, Gamarra &amp; Salgado" authorityYear="2000" box="[614,1197,223,248]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="caurelensis">
<emphasis id="B942955B9673D047B555249EFCCEFF34" box="[614,775,223,248]" italics="true" pageId="7" pageNumber="408">D. caurelensis</emphasis>
<bibRefCitation id="EFA734B89673D047B42324A0FB64FF34" author="Outerelo" box="[784,1197,223,248]" pageId="7" pageNumber="408" refString="Outerelo, R., Gamarra, P. &amp; Salgado, J. M. (2000) Los Staphylinidae (Coleoptera) cavernicolas del Noroeste de la Peninsula Iberica. II. Campanas de 1985 a 1996. Memoires de Biospeologie, 27, 107 - 121." type="journal article" year="2000">Outerelo, Gamarra &amp; Salgado, 2000</bibRefCitation>
</taxonomicName>
. However, unlike the new species, the ventral blade in 
<taxonomicName id="4C3632CA9673D047B521257AFD6DFED1" box="[530,676,260,285]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="B942955B9673D047B521257AFD6DFED1" box="[530,676,260,285]" italics="true" pageId="7" pageNumber="408">D. lusitanica</emphasis>
</taxonomicName>
does not exceed the edge of the aedeagus, presents a pointed apex and lacks the minor middle lateral hook-like expansions between the apex and base (cf. 
<figureCitation id="130D55CC9673D047B3B52558FB05FEF3" box="[1158,1228,295,320]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="7" pageNumber="408">Figs 7</figureCitation>
a‒7c and 
<figureCitation id="130D55CC9673D047B2082558FAB7FE8C" box="[1339,1406,295,320]" captionStart="FIGURE 6" captionStartId="7.[151,250,1152,1174]" captionTargetBox="[169,1407,488,1107]" captionTargetId="figure@7.[151,1436,481,1131]" captionTargetPageId="7" captionText="FIGURE 6. Genital segment of Domene viriatoi n. sp. (ventral view): (a), male; (b), female." httpUri="https://zenodo.org/record/240971/files/figure.png" pageId="7" pageNumber="408">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="EFA734B89673D047B7A42533FE5AFEA9" author="Reboleira" box="[151,403,332,357]" pageId="7" pageNumber="408" refString="Reboleira A. S. P. S., Goncalves, F. &amp; Oromi, P. (2011 b) On the Iberian endemic subgenus Lathromene Koch (Coleoptera: Staphylinidae: Paederinae): description of the first hypogean Domene Fauvel, 1872 from Portugal. Zootaxa, 2780, 48 - 56." type="journal article" year="2011" yearSuffix="b">
Reboleira 
<emphasis id="B942955B9673D047B6382532FE8AFEA9" box="[267,323,332,357]" italics="true" pageId="7" pageNumber="408">et al.</emphasis>
2011b
</bibRefCitation>
). On the other hand, the ventral blade of 
<taxonomicName id="4C3632CA9673D047B46F2532FC35FEA9" box="[860,1020,332,357]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="caurelensis">
<emphasis id="B942955B9673D047B46F2532FC35FEA9" box="[860,1020,332,357]" italics="true" pageId="7" pageNumber="408">D. caurelensis</emphasis>
</taxonomicName>
only exceeds slightly the edge of the aedeagus, the apex is strongly bifurcated and the minor middle lateral hook-like expansions are also absent (cf. 
<figureCitation id="130D55CC9673D047B7A425EBFF12FE61" box="[151,219,404,429]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="7" pageNumber="408">Figs 5</figureCitation>
a‒5c and 
<figureCitation id="130D55CC9673D047B67425EBFE40FE60" box="[327,393,404,429]" captionStart="FIGURE 7" captionStartId="7.[151,250,2012,2034]" captionTargetBox="[273,1325,1235,1982]" captionTargetId="figure@7.[251,1336,1224,1990]" captionTargetPageId="7" captionText="FIGURE 7. Aedeagus of Domene viriatoi n. sp.: (a), lateral view; (b), dorsal view; (c), ventral view." httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="7" pageNumber="408">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="EFA734B89673D047B69E25EBFD58FE60" author="Outerelo" box="[429,657,404,429]" pageId="7" pageNumber="408" refString="Outerelo, R., Gamarra, P. &amp; Salgado, J. M. (2000) Los Staphylinidae (Coleoptera) cavernicolas del Noroeste de la Peninsula Iberica. II. Campanas de 1985 a 1996. Memoires de Biospeologie, 27, 107 - 121." type="journal article" year="2000">
Outerelo 
<emphasis id="B942955B9673D047B52625EAFD87FE61" box="[533,590,404,429]" italics="true" pageId="7" pageNumber="408">et al.</emphasis>
2000
</bibRefCitation>
).
</paragraph>
<caption id="DF4919C19673D047B7A420FFFB8EFB5A" box="[151,1095,1152,1175]" httpUri="https://zenodo.org/record/240971/files/figure.png" pageId="7" pageNumber="408" targetBox="[169,1407,488,1107]" targetPageId="7">
<paragraph id="8B8949499673D047B7A420FFFB8EFB5A" blockId="7.[151,1095,1152,1175]" box="[151,1095,1152,1175]" pageId="7" pageNumber="408">
<emphasis id="B942955B9673D047B7A420FFFEDBFB5A" bold="true" box="[151,274,1152,1174]" pageId="7" pageNumber="408">FIGURE 6.</emphasis>
Genital segment of 
<taxonomicName id="4C3632CA9673D047B6D320FDFD4FFB5A" box="[480,646,1153,1175]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9673D047B6D320FDFD4FFB5A" box="[480,646,1153,1175]" italics="true" pageId="7" pageNumber="408">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9673D047B5B820FFFD0CFB5A" bold="true" box="[651,709,1152,1174]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="A27128209673D047B5B820FFFD0CFB5A" box="[651,709,1152,1174]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(ventral view): 
<emphasis id="B942955B9673D047B45420FFFC4DFB5A" bold="true" box="[871,900,1152,1174]" pageId="7" pageNumber="408">(a)</emphasis>
, male; 
<emphasis id="B942955B9673D047B4FC20FFFC27FB5A" bold="true" box="[975,1006,1152,1174]" pageId="7" pageNumber="408">(b)</emphasis>
, female.
</paragraph>
</caption>
<caption id="DF4919C19673D047B7A423A3FB50F83E" box="[151,1177,2012,2034]" httpUri="https://zenodo.org/record/240972/files/figure.png" pageId="7" pageNumber="408" targetBox="[273,1325,1235,1982]" targetPageId="7">
<paragraph id="8B8949499673D047B7A423A3FB50F83E" blockId="7.[151,1177,2012,2034]" box="[151,1177,2012,2034]" pageId="7" pageNumber="408">
<emphasis id="B942955B9673D047B7A423A3FEDBF83E" bold="true" box="[151,274,2012,2034]" pageId="7" pageNumber="408">FIGURE 7.</emphasis>
Aedeagus of 
<taxonomicName id="4C3632CA9673D047B6AD23A2FD8DF83E" box="[414,580,2013,2034]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B9673D047B6AD23A2FD8DF83E" box="[414,580,2013,2034]" italics="true" pageId="7" pageNumber="408">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B9673D047B57923A3FD4AF83E" bold="true" box="[586,643,2012,2034]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="A27128209673D047B57923A3FD4AF83E" box="[586,643,2012,2034]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B942955B9673D047B5A323A3FD67F83E" bold="true" box="[656,686,2012,2034]" pageId="7" pageNumber="408">(a)</emphasis>
, lateral view; 
<emphasis id="B942955B9673D047B40C23A3FC97F83E" bold="true" box="[831,862,2012,2034]" pageId="7" pageNumber="408">(b)</emphasis>
, dorsal view; 
<emphasis id="B942955B9673D047B4DE23A3FBC0F83E" bold="true" box="[1005,1033,2012,2034]" pageId="7" pageNumber="408">(c)</emphasis>
, ventral view.
</paragraph>
</caption>
<caption id="DF4919C1967CD048B7A4270BFDF7FC46" box="[151,574,884,907]" pageId="8" pageNumber="409">
<paragraph id="8B894949967CD048B7A4270BFDF7FC46" blockId="8.[151,574,884,907]" box="[151,574,884,907]" pageId="8" pageNumber="409">
<emphasis id="B942955B967CD048B7A4270BFEDBFC46" bold="true" box="[151,274,884,906]" pageId="8" pageNumber="409">FIGURE 8.</emphasis>
<taxonomicName id="4C3632CA967CD048B62A2709FE77FC47" box="[281,446,886,907]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B62A2709FE77FC47" box="[281,446,886,907]" italics="true" pageId="8" pageNumber="409">Domene viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B967CD048B6F7270BFE37FC46" bold="true" box="[452,510,884,906]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A2712820967CD048B6F7270BFE37FC46" box="[452,510,884,906]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
larva.
</paragraph>
</caption>
<paragraph id="8B894949967CD048B7F427CDFCCEFB6F" blockId="8.[151,1437,946,1582]" pageId="8" pageNumber="409">
Furthermore, the differences concerning the male sternite VIII between the new species and its closest relatives are even more evident. 
<taxonomicName id="4C3632CA967CD048B69A27A8FDABFC22" box="[425,610,983,1006]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B69A27A8FDABFC22" box="[425,610,983,1006]" italics="true" pageId="8" pageNumber="409">Domene viriatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2712820967CD048B55F27AAFD67FC22" box="[620,686,981,1006]" pageId="8" pageNumber="409" rank="species">
<emphasis id="B942955B967CD048B55F27AAFD67FC22" bold="true" box="[620,686,981,1006]" pageId="8" pageNumber="409">n. sp.</emphasis>
</taxonomicNameLabel>
exhibits posteriorly a distinct inverted U‒shaped incision with a cluster of 15‒21 modified, short and stout black striated setae on either side of the apex (
<figureCitation id="130D55CC967CD048B35C2785FB7CFBDF" box="[1135,1205,1018,1043]" captionStart="FIGURE 5" captionStartId="6.[151,250,1453,1475]" captionTargetBox="[188,1398,193,1431]" captionTargetId="figure@6.[188,1398,193,1431]" captionTargetPageId="6" captionText="FIGURE 5. SEM photographs of structural features of Domene viriatoi n. sp.: (a), III abdominal sternite (latero ‒ ventral view); (b), detail of intersegmental area between III and IV abdominal sternites (latero ‒ ventral view); (c), VII, VIII and IX abdominal sternites (ventral view); (d), detail of middle posteriad region of VII abdominal sternite (male, ventral view); (e), detail of VIII abdominal sternite (male, ventral view); (f), detail of the cluster of stout striate setae in VIII abdominal sternite (ventral view)." httpUri="https://zenodo.org/record/240970/files/figure.png" pageId="8" pageNumber="409">Figs 5</figureCitation>
e, 6a). The other two species, although presenting a posterior incision, it is not so pronounced as in the new species and lack short and stout black striated setae. Interestingly, some representatives of 
<taxonomicName id="4C3632CA967CD048B450203CFC08FB96" box="[867,961,1091,1114]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="B942955B967CD048B450203CFC08FB96" box="[867,961,1091,1114]" italics="true" pageId="8" pageNumber="409">Domene</emphasis>
</taxonomicName>
(
<emphasis id="B942955B967CD048B4E2203CFBCFFB96" box="[977,1030,1091,1114]" italics="true" pageId="8" pageNumber="409">s. str</emphasis>
.), like 
<taxonomicName id="4C3632CA967CD048B365203CFF06FBB2" authority="Hernando &amp; Baena, 2006" authorityName="Hernando &amp; Baena" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="gevia">
<emphasis id="B942955B967CD048B365203CFB7DFB96" box="[1110,1204,1091,1114]" italics="true" pageId="8" pageNumber="409">D. gevia</emphasis>
<bibRefCitation id="EFA734B8967CD048B38F203DFF06FBB2" author="Hernando" pageId="8" pageNumber="409" refString="Hernando, C. &amp; Baena, M. (2006) Un nuevo estafilinido de Andalucia: Domene (s. str.) gevia sp. nov. (Coleoptera, Staphylinidae, Paederinae). Heteropterus. Revista de Entomologia, 6, 29 - 33." type="journal article" year="2006">Hernando &amp; Baena, 2006</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3632CA967CD048B6342018FDC4FBB2" authority="Assing, 2012" authorityName="Assing" authorityYear="2012" box="[263,525,1125,1150]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="perezi">
<emphasis id="B942955B967CD048B6342018FEA4FBB2" box="[263,365,1127,1150]" italics="true" pageId="8" pageNumber="409">D. perezi</emphasis>
Assing, 2012
</taxonomicName>
, present a pair of clusters of stout setae in sternite VIII also, but near the base or along the edges of the inverted U or V‒shaped incision.
</paragraph>
<paragraph id="8B894949967CD048B7F420D2FAFFFAB7" blockId="8.[151,1437,946,1582]" pageId="8" pageNumber="409">
The congeners with populations closer to the new species are 
<taxonomicName id="4C3632CA967CD048B44C20D0FBD8FB0A" box="[895,1041,1197,1222]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="B942955B967CD048B44C20D0FBD8FB0A" box="[895,1041,1197,1222]" italics="true" pageId="8" pageNumber="409">D. lusitanica</emphasis>
</taxonomicName>
(from Serra de Sicó, 
<collectingCountry id="F32109D9967CD048B23720D2FAACFB0A" box="[1284,1381,1197,1222]" name="Portugal" pageId="8" pageNumber="409">Portugal</collectingCountry>
) and 
<taxonomicName id="4C3632CA967CD048B7A420ACFE14FB27" authority="Outerelo, 1985" authorityName="Outerelo" authorityYear="1985" box="[151,477,1234,1259]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B942955B967CD048B7A420ACFEE0FB27" box="[151,297,1234,1259]" italics="true" pageId="8" pageNumber="409">D. hispanica</emphasis>
Outerelo, 1985
</taxonomicName>
(from Peña de 
<collectingCountry id="F32109D9967CD048B5A620ADFD25FB27" box="[661,748,1234,1259]" name="France" pageId="8" pageNumber="409">Francia</collectingCountry>
, 
<collectingCountry id="F32109D9967CD048B5CA20ADFCF7FB27" box="[761,830,1234,1259]" name="Spain" pageId="8" pageNumber="409">Spain</collectingCountry>
). The latter species, which male is unknown, can be segregated from 
<taxonomicName id="4C3632CA967CD048B6602088FE0EFAC2" box="[339,455,1271,1294]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B6602088FE0EFAC2" box="[339,455,1271,1294]" italics="true" pageId="8" pageNumber="409">D. viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B967CD048B6FC208AFDC7FAC2" bold="true" box="[463,526,1269,1294]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A2712820967CD048B6FC208AFDC7FAC2" box="[463,526,1269,1294]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the presence of a pronotal sulcus in the posteriad middle region (absent in the new species), the flattening of the posterior region of pronotum (in 
<taxonomicName id="4C3632CA967CD048B4922164FBD1FAFE" box="[929,1048,1307,1330]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B4922164FBD1FAFE" box="[929,1048,1307,1330]" italics="true" pageId="8" pageNumber="409">D. viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B967CD048B3122165FBABFAFF" bold="true" box="[1057,1122,1306,1331]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A2712820967CD048B3122165FBABFAFF" box="[1057,1122,1306,1331]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is slightly convex) and the elytral length (equal to pronotum in 
<taxonomicName id="4C3632CA967CD048B5062140FD0CFA9A" box="[565,709,1341,1366]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B942955B967CD048B5062140FD0CFA9A" box="[565,709,1341,1366]" italics="true" pageId="8" pageNumber="409">D. hispanica</emphasis>
</taxonomicName>
and smaller than pronotum in 
<taxonomicName id="4C3632CA967CD048B3162140FB55FA9A" box="[1061,1180,1343,1366]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B3162140FB55FA9A" box="[1061,1180,1343,1366]" italics="true" pageId="8" pageNumber="409">D. viriatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2712820967CD048B3972142FB2BFA9A" box="[1188,1250,1341,1366]" pageId="8" pageNumber="409" rank="species">
<emphasis id="B942955B967CD048B3972142FB14FA9A" bold="true" box="[1188,1245,1341,1366]" pageId="8" pageNumber="409">n. sp</emphasis>
.
</taxonomicNameLabel>
). Moreover, the number of ommatidia in 
<taxonomicName id="4C3632CA967CD048B69E211CFDF5FAB7" box="[429,572,1378,1403]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B942955B967CD048B69E211CFDF5FAB7" box="[429,572,1378,1403]" italics="true" pageId="8" pageNumber="409">D. hispanica</emphasis>
</taxonomicName>
is 15 (Outerelo 1985), much less than in 
<taxonomicName id="4C3632CA967CD048B33F211CFB49FAB6" box="[1036,1152,1379,1402]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="viriatoi" status="sp. nov.">
<emphasis id="B942955B967CD048B33F211CFB49FAB6" box="[1036,1152,1379,1402]" italics="true" pageId="8" pageNumber="409">D. viriatoi</emphasis>
</taxonomicName>
<emphasis id="B942955B967CD048B3B4211DFB0FFAB7" bold="true" box="[1159,1222,1378,1403]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A2712820967CD048B3B4211DFB0FFAB7" box="[1159,1222,1378,1403]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130D55CC967CD048B3E5211DFAD1FAB7" box="[1238,1304,1378,1403]" captionStart="FIGURE 3" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[188,1398,193,1853]" captionTargetId="figure@4.[188,1398,193,1854]" captionTargetPageId="4" captionText="FIGURE 3. SEM photographs of structural features of Domene viriatoi n. sp.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in foretibia (latero ‒ ventral view); (f), part of the cleaning organ in forefemur (latero ‒ ventral view); (g), pronotum (dorsal view); (h), elytra (dorsal view)." httpUri="https://zenodo.org/record/240969/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
b).
</paragraph>
<paragraph id="8B894949967CD048B7F421FAFCE8F9E2" blockId="8.[151,1437,946,1582]" pageId="8" pageNumber="409">
The morphological resemblance between some species of 
<taxonomicName id="4C3632CA967CD048B46421F8FC7DFA52" box="[855,948,1415,1438]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="B942955B967CD048B46421F8FC7DFA52" box="[855,948,1415,1438]" italics="true" pageId="8" pageNumber="409">Domene</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3632CA967CD048B4DF21FAFBBAFA52" box="[1004,1139,1413,1438]" class="Insecta" family="Staphylinidae" genus="Lathromene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="B942955B967CD048B4DF21FAFBBAFA52" box="[1004,1139,1413,1438]" italics="true" pageId="8" pageNumber="409">Lathromene</emphasis>
</taxonomicName>
subgenera (e.g. shape and presence of stout setae in the sternite VIII incision) shows how controversial these supraspecific taxonomic entities are. Nevertheless, although practical, the subgeneric classification of genus 
<taxonomicName id="4C3632CA967CD048B4C521B0FB9DFA2A" box="[1014,1108,1487,1510]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="B942955B967CD048B4C521B0FB9DFA2A" box="[1014,1108,1487,1510]" italics="true" pageId="8" pageNumber="409">Domene</emphasis>
</taxonomicName>
is at present highly artificial and in urgent need for revision following a comprehensive phylogenetic analysis (
<bibRefCitation id="EFA734B8967CD048B36E218DFA58F9C6" author="Assing" box="[1117,1425,1522,1547]" pageId="8" pageNumber="409" refString="Assing, V. &amp; Feldmann, B. (2014) On Domene scabripennis Rougemont and its close relatives (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage, 46, 499 - 514." type="journal article" year="2014">Assing &amp; Feldmann 2014</bibRefCitation>
). However, this task is out of the scope of the present work.
</paragraph>
</subSubSection>
</treatment>
</document>