171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="D597B87917440AC4B1EC544336AE590E" ID-CLB-Dataset="46802" ID-DOI="10.11646/zootaxa.4078.1.20" ID-GBIF-Dataset="ba98412f-c54a-48f7-b1f3-7b38f56f8b35" ID-ISSN="1175-5326" ID-Zenodo-Dep="263680" ID-ZooBank="836FA2B8-7871-4DED-9646-BD39A4BAA031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460361344939" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho" docDate="2016" docId="0395E107FF9F680DFF60FD9F3682F887" docLanguage="en" docName="zootaxa.4078.1.20.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Montesinus machadoi Grossi, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="228" masterDocId="FFAC997FFF956807FFF7FFBB3272FF81" masterDocTitle="A new brachypterous genus of Brazilian stag beetle (Coleoptera: Lucanidae: Lucaninae) with description of three new species" masterLastPageNumber="229" masterPageNumber="218" pageNumber="228" updateTime="1698294582438" updateUser="ExternalLinkService">
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<mods:title id="A1C55F1B6DE63851A1BAE5FCF89435FB">A new brachypterous genus of Brazilian stag beetle (Coleoptera: Lucanidae: Lucaninae) with description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="D801652F9E73008D845B338D62B87A9A" type="personal">
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<mods:roleTerm id="241DB5099FF72A84DF6550D83D24BB88">Author</mods:roleTerm>
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<mods:namePart id="114BC6A3242D0CF6366E58B4CDB617F4">Grossi, Paschoal Coelho</mods:namePart>
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<mods:typeOfResource id="E844DA8AF8293F769ADFF001D1F2E756">text</mods:typeOfResource>
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<mods:title id="587B371807C368C36E5EBF977956D998">Zootaxa</mods:title>
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<mods:part id="F438751D0134176ABFA9905BB3170B56">
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<mods:date id="3DDEED19266AB24481D29AE76AF36FF6">2016</mods:date>
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<mods:number id="07DC1B71197D7552BB09331168C714C5">4078</mods:number>
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<mods:classification id="C25B86C27D4272E537160DDE42CD4140">journal article</mods:classification>
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<mods:identifier id="C4ACC59A6B24AA8754BABF0D7A4CA2B0" type="DOI">10.11646/zootaxa.4078.1.20</mods:identifier>
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<treatment id="0395E107FF9F680DFF60FD9F3682F887" ID-DOI="http://doi.org/10.5281/zenodo.6073660" ID-GBIF-Taxon="119409830" ID-Zenodo-Dep="6073660" LSID="urn:lsid:plazi:treatment:0395E107FF9F680DFF60FD9F3682F887" httpUri="http://treatment.plazi.org/id/0395E107FF9F680DFF60FD9F3682F887" lastPageNumber="228" pageId="10" pageNumber="228">
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<subSubSection id="C326039AFF9F680DFF60FD9F33E3FDDE" pageId="10" pageNumber="228" type="nomenclature">
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<paragraph id="8B835011FF9F680DFF60FD9F307AFDBF" blockId="10.[151,520,548,607]" box="[151,520,548,574]" pageId="10" pageNumber="228">
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<heading id="D0CBE77DFF9F680DFF60FD9F307AFDBF" bold="true" box="[151,520,548,574]" fontSize="11" level="1" pageId="10" pageNumber="228" reason="1">
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<emphasis id="B9488C03FF9F680DFF60FD9F307AFDBF" bold="true" box="[151,520,548,574]" pageId="10" pageNumber="228">
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<taxonomicName id="4C3C2B92FF9F680DFF60FD9F33D1FDBF" ID-CoL="73Q7M" box="[151,419,548,574]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
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<emphasis id="B9488C03FF9F680DFF60FD9F33D1FDBF" bold="true" box="[151,419,548,574]" italics="true" pageId="10" pageNumber="228">Montesinus machadoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B3178FF9F680DFE5DFD9F307AFDBF" box="[426,520,548,574]" pageId="10" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B835011FF9F680DFF60FDFD33E3FDDE" blockId="10.[151,520,548,607]" box="[151,401,582,607]" pageId="10" pageNumber="228">
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(
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<figureCitation id="13074C94FF9F680DFF69FDFD3379FDDE" box="[158,267,582,607]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1457,1479]" captionTargetBox="[375,1210,212,1418]" captionTargetId="figure@3.[355,1232,193,1435]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Habitus of Montesinus species. 1 – 3. Montesinus monnei sp. nov., holotype male in dorsal (1) and lateral views (2), and allotype female in dorsal view (3); 4. M. tatula sp. nov., holotype female in dorsal view; 5. M. machadoi Grossi sp. nov., holotype female in dorsal view. Scale bars = 4 mm." httpUri="https://zenodo.org/record/263681/files/figure.png" pageId="10" pageNumber="228">Figures 5</figureCitation>
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,
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<figureCitation id="13074C94FF9F680DFEE0FDFC3347FDDE" box="[279,309,583,607]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1971,1993]" captionTargetBox="[211,1360,196,1937]" captionTargetId="figure@6.[187,1392,193,1957]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Left maxillae in dorsal view: 18 – 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263684/files/figure.png" pageId="10" pageNumber="228">21</figureCitation>
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,
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<figureCitation id="13074C94FF9F680DFEB6FDFD33FBFDDE" box="[321,393,582,607]" captionStart="FIGURES 22 – 27" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[237,1354,210,1914]" captionTargetId="figure@7.[217,1369,193,1924]" captionTargetPageId="7" captionText="FIGURES 22 – 27. Details of morphology of Altitatiayus Weinreich and Montesinus gen. nov. 22 – 24. right female protibiae (punctures and setae omitted); 26 – 27. wings in dorsal view; 22. Montesinus monnei sp. nov.; 23. M. tatula sp. nov.; 24 – 25. M. machadoi sp. nov., 25. right wing; 26. Altitatiayus ruficollis (Lüderwaldt); 27. A. rotundatus (Boileau)." httpUri="https://zenodo.org/record/263685/files/figure.png" pageId="10" pageNumber="228">24–25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C326039AFF9F680DFF60FD3533FDFC92" pageId="10" pageNumber="228" type="materials_examined">
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<paragraph id="8B835011FF9F680DFF60FD3533FDFC92" blockId="10.[151,1437,654,1798]" pageId="10" pageNumber="228">
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<emphasis id="B9488C03FF9F680DFF60FD3533F3FD26" bold="true" box="[151,385,654,679]" pageId="10" pageNumber="228">Material examined.</emphasis>
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<typeStatus id="5487EEB3FF9F680DFE7FFD353382FD26" box="[392,496,654,679]" pageId="10" pageNumber="228" type="holotype">Holotype</typeStatus>
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female labeled a) “
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<collectingCountry id="F32B1081FF9F680DFD3CFD353160FD26" box="[715,786,654,679]" name="Brazil" pageId="10" pageNumber="228">Brasil</collectingCountry>
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, Espírito Santo, Serra do Caparaó, Ibitirama,
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<quantity id="4CC4FDF4FF9F680DFAE4FD353715FD26" box="[1299,1383,654,679]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="228" unit="m" value="2600.0">2.600m</quantity>
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/
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<date id="FF8276D1FF9F680DFA80FD34336AFD4A" pageId="10" pageNumber="228" value="2012-04-10" valueMax="2012-04-12" valueMin="2012-04-10">10- 12-IV-2012</date>
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/ M.V.P. Simões col.”.b) “
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<taxonomicName id="4C3C2B92FF9F680DFDA5FD0F313CFD4A" box="[594,846,690,715]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
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<emphasis id="B9488C03FF9F680DFDA5FD0F313CFD4A" box="[594,846,690,715]" italics="true" pageId="10" pageNumber="228">Montesinus machadoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B3178FF9F680DFCACFD0831EFFD4B" box="[859,925,691,714]" pageId="10" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
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♀/
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<typeStatus id="5487EEB3FF9F680DFC23FD083615FD4A" box="[980,1127,691,715]" pageId="10" pageNumber="228" type="holotype">HOLOTYPE</typeStatus>
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/ P. C. Grossi det. 2015”.
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<typeStatus id="5487EEB3FF9F680DFF60FD6D328DFD6E" box="[151,255,726,751]" pageId="10" pageNumber="228" type="holotype">Holotype</typeStatus>
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deposited at
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<collectionCode id="ED2DC8D4FF9F680DFE68FD6C339FFD6E" box="[415,493,727,751]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="10" pageNumber="228">MNRJ</collectionCode>
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.One
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<typeStatus id="5487EEB3FF9F680DFDDCFD6C30F9FD6E" box="[555,651,727,751]" pageId="10" pageNumber="228" type="paratype">paratype</typeStatus>
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female deposited at
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<collectionCode id="ED2DC8D4FF9F680DFC75FD6C31ABFD6E" box="[898,985,727,751]" pageId="10" pageNumber="228">CERPE</collectionCode>
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and labeled as
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<typeStatus id="5487EEB3FF9F680DFB61FD6D368BFD6E" box="[1174,1273,726,751]" pageId="10" pageNumber="228" type="holotype">holotype</typeStatus>
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,except by the yellow
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<typeStatus id="5487EEB3FF9F680DFF1DFD403338FC92" box="[234,330,763,787]" pageId="10" pageNumber="228" type="paratype">paratype</typeStatus>
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label.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326039AFF9F680DFF30FCA531B8FA1F" pageId="10" pageNumber="228" type="description">
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<paragraph id="8B835011FF9F680DFF30FCA533DEFAB3" blockId="10.[151,1437,654,1798]" pageId="10" pageNumber="228">
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<emphasis id="B9488C03FF9F680DFF30FCA5304CFCB6" bold="true" box="[199,574,798,823]" pageId="10" pageNumber="228">
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Description of female
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<typeStatus id="5487EEB3FF9F680DFE20FCA5304CFCB6" box="[471,574,798,823]" pageId="10" pageNumber="228" type="holotype">holotype</typeStatus>
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</emphasis>
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. Total length, 18.0 mm. Total pronotal width 6.9 mm.
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<emphasis id="B9488C03FF9F680DFB33FCA53777FCB6" box="[1220,1285,798,823]" italics="true" pageId="10" pageNumber="228">Color</emphasis>
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: Body shiny black, except by dark brownish red femora.
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<emphasis id="B9488C03FF9F680DFD62FCF9316AFCDA" box="[661,792,834,859]" italics="true" pageId="10" pageNumber="228">Mouthparts</emphasis>
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: Mandibles with dorsal longitudinal furrow distinct and more dorsally visible; internal tooth somewhat forward directed and forming a carina to base, left one broken. Labrum trapezoidal, anteriorly slightly lobed, surface with scattered setose punctures. Mentum transverse, densely punctate; punctures shallow, coarse, and coalescent, each with 1 long setae; setae longer near anterior and lateral sides. Maxillae with a hooklet on apex of galea; lacinial hook slightly backwardly curved (
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<figureCitation id="13074C94FF9F680DFB64FC693694FC6A" box="[1171,1254,978,1003]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1971,1993]" captionTargetBox="[211,1360,196,1937]" captionTargetId="figure@6.[187,1392,193,1957]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Left maxillae in dorsal view: 18 – 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263684/files/figure.png" pageId="10" pageNumber="228">Fig. 21</figureCitation>
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).
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<emphasis id="B9488C03FF9F680DFB0CFC69374AFC6A" box="[1275,1336,978,1003]" italics="true" pageId="10" pageNumber="228">Head</emphasis>
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: Surface almost totally punctate except clypeal area and vertex; punctures large to coarse, never coalescent. Ocular canthi simply rounded, and not expanded. Vertex convex. Gula concave, with scattered C-shaped punctures, tegument reticulate; anterior lateral angles not so projected, rounded. Gena near eyes with coalescent coarse punctures.
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<emphasis id="B9488C03FF9F680DFEFCFBD9330FFBFA" box="[267,381,1122,1147]" italics="true" pageId="10" pageNumber="228">Prothorax</emphasis>
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: Surface finely punctate; punctures fine; posterior angles absent; posterior declivity strong.
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<taxonomicName id="4C3C2B92FF9F680DFF60FB3D3324FB1E" authority="Sides" authorityName="Sides" box="[151,342,1158,1183]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="10" pageNumber="228" phylum="Arthropoda" rank="genus">
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<emphasis id="B9488C03FF9F680DFF60FB3D3377FB1E" box="[151,261,1158,1183]" italics="true" pageId="10" pageNumber="228">Scutellum</emphasis>
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: Sides
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</taxonomicName>
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rounded; surface with scattered shallow punctures.
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<emphasis id="B9488C03FF9F680DFC36FB3D3675FB1E" box="[961,1031,1158,1183]" italics="true" pageId="10" pageNumber="228">Elytra</emphasis>
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: Surface less irregularly punctate; almost punctures fine; elytral scratches wider, distance from suture and first scratch wider. Humeri truncate, but more projected.
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<emphasis id="B9488C03FF9F680DFEBAFB7433F0FB67" box="[333,386,1231,1254]" italics="true" pageId="10" pageNumber="228">Legs</emphasis>
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: Protibiae with 4 spines, 2 anterior and 2 at middle; second spine wider; distance between 3rd and 4th spines smaller (
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<figureCitation id="13074C94FF9F680DFE61FB4E3398FA8F" box="[406,490,1269,1294]" captionStart="FIGURES 22 – 27" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[237,1354,210,1914]" captionTargetId="figure@7.[217,1369,193,1924]" captionTargetPageId="7" captionText="FIGURES 22 – 27. Details of morphology of Altitatiayus Weinreich and Montesinus gen. nov. 22 – 24. right female protibiae (punctures and setae omitted); 26 – 27. wings in dorsal view; 22. Montesinus monnei sp. nov.; 23. M. tatula sp. nov.; 24 – 25. M. machadoi sp. nov., 25. right wing; 26. Altitatiayus ruficollis (Lüderwaldt); 27. A. rotundatus (Boileau)." httpUri="https://zenodo.org/record/263685/files/figure.png" pageId="10" pageNumber="228">Fig. 24</figureCitation>
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). Mesotibiae with 3 spines, the middle one much smaller.
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<emphasis id="B9488C03FF9F680DFB85FB4D36C4FA8C" box="[1138,1206,1270,1293]" italics="true" pageId="10" pageNumber="228">Venter</emphasis>
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: All ventrites finely punctate; punctures fine.
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</paragraph>
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<paragraph id="8B835011FF9F680DFF30FA8631B8FA1F" blockId="10.[151,1437,654,1798]" pageId="10" pageNumber="228">
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<emphasis id="B9488C03FF9F680DFF30FA863347FAD7" bold="true" box="[199,309,1341,1366]" pageId="10" pageNumber="228">Variation</emphasis>
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. The other female seems to be a little damaged but some characters are distinct as follows: body wider; posterior angles of pronotum a little more pronounced; anterior tibiae with 5 (left) to 6 spines (right); right mesotibia with four spines, with basal ones larger than those of
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<typeStatus id="5487EEB3FF9F680DFC97FA3E31B4FA1F" box="[864,966,1413,1438]" pageId="10" pageNumber="228" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326039AFF9F680DFF30FA1233C8F98B" pageId="10" pageNumber="228" type="etymology">
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<paragraph id="8B835011FF9F680DFF30FA1233C8F98B" blockId="10.[151,1437,654,1798]" pageId="10" pageNumber="228">
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<emphasis id="B9488C03FF9F680DFF30FA123334FA43" bold="true" box="[199,326,1449,1474]" pageId="10" pageNumber="228">Etymology</emphasis>
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. The specific name is an homage to another great friend, Dr. Angelo Barbosa Monteiro Machado, one of the great zoologists in
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<collectingCountry id="F32B1081FF9F680DFE29FA763050FA67" box="[478,546,1485,1510]" name="Brazil" pageId="10" pageNumber="228">Brazil</collectingCountry>
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and an Odonata specialist who always helped my research through his donation of stag beetles specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326039AFF9F680DFF30F9AE3682F887" pageId="10" pageNumber="228" type="discussion">
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<paragraph id="8B835011FF9F680DFF30F9AE3682F887" blockId="10.[151,1437,654,1798]" pageId="10" pageNumber="228">
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<emphasis id="B9488C03FF9F680DFF30F9AE3346F9AF" bold="true" box="[199,308,1557,1582]" pageId="10" pageNumber="228">Remarks</emphasis>
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. Both
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<typeStatus id="5487EEB3FF9F680DFE77F9AE3390F9AF" box="[384,482,1557,1582]" pageId="10" pageNumber="228" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="5487EEB3FF9F680DFDEDF9AD3009F9AF" box="[538,635,1558,1582]" pageId="10" pageNumber="228" type="paratype">paratype</typeStatus>
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have missing tarsi, and were collected alive walking during a cold and rainy day along the track to Pico da Bandeira (Marianna V. P. Simões, pers. comm.). Initially, I believed that these two specimens belonged to one of the other two species, but after dissecting the maxillae, the characters proved to be distinct, so revealing the existence of this new species. It is closer to
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<taxonomicName id="4C3C2B92FF9F680DFC21F939363DF918" box="[982,1103,1666,1689]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="monnei" status="sp. nov.">
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<emphasis id="B9488C03FF9F680DFC21F939363DF918" box="[982,1103,1666,1689]" italics="true" pageId="10" pageNumber="228">M. monnei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B3178FF9F680DFBAEF93A36C6F91B" box="[1113,1204,1665,1690]" pageId="10" pageNumber="228" rank="species">
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<emphasis id="B9488C03FF9F680DFBAEF93A36C6F91B" bold="true" box="[1113,1204,1665,1690]" pageId="10" pageNumber="228">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in sharing the same elytral punctures and the shape of maxillae, and the presence of hooklets on it, but differs from this species in the armature of the galea and lacinia. The species can be distinguished by the large size of
|
||
<taxonomicName id="4C3C2B92FF9F680DFB6AF9713747F963" box="[1181,1333,1737,1762]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
|
||
<emphasis id="B9488C03FF9F680DFB6AF9713747F963" box="[1181,1333,1737,1762]" italics="true" pageId="10" pageNumber="228">M. machadoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B3178FF9F680DFAC8F97237EEF963" box="[1343,1436,1737,1762]" pageId="10" pageNumber="228" rank="species">
|
||
<emphasis id="B9488C03FF9F680DFAC8F97237EEF963" bold="true" box="[1343,1436,1737,1762]" pageId="10" pageNumber="228">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(similar to that of
|
||
<taxonomicName id="4C3C2B92FF9F680DFE96F95433B5F887" box="[353,455,1773,1798]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="tatula" status="sp. nov.">
|
||
<emphasis id="B9488C03FF9F680DFE96F95433B5F887" box="[353,455,1773,1798]" italics="true" pageId="10" pageNumber="228">M. tatula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9488C03FF9F680DFE38F956305AF887" bold="true" box="[463,552,1773,1798]" pageId="10" pageNumber="228">
|
||
<taxonomicNameLabel id="A27B3178FF9F680DFE38F956305AF887" box="[463,552,1773,1798]" pageId="10" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), shape of head, and number of spines on meso and metatibiae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |