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<document id="FA561AC565931D970865145023B54C70" ID-CLB-Dataset="46802" ID-DOI="10.11646/zootaxa.4078.1.20" ID-GBIF-Dataset="ba98412f-c54a-48f7-b1f3-7b38f56f8b35" ID-ISSN="1175-5326" ID-Zenodo-Dep="263680" ID-ZooBank="836FA2B8-7871-4DED-9646-BD39A4BAA031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460361344939" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho" docDate="2016" docId="0395E107FF9D680EFF60FF2C3622FC01" docLanguage="en" docName="zootaxa.4078.1.20.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Montesinus monnei Grossi, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="227" masterDocId="FFAC997FFF956807FFF7FFBB3272FF81" masterDocTitle="A new brachypterous genus of Brazilian stag beetle (Coleoptera: Lucanidae: Lucaninae) with description of three new species" masterLastPageNumber="229" masterPageNumber="218" pageNumber="226" updateTime="1698294582438" updateUser="ExternalLinkService">
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<mods:title id="A6BF9FE29F49D76F8AA0D74615C02D81">A new brachypterous genus of Brazilian stag beetle (Coleoptera: Lucanidae: Lucaninae) with description of three new species</mods:title>
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<mods:name id="B9C9370D09D545ACC1EB6687C8343A55" type="personal">
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<mods:namePart id="D07B506FF0C8032A2338EC0FC22FAF97">Grossi, Paschoal Coelho</mods:namePart>
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<mods:date id="7C07745DA9D34BFF57E8EBE0B136EA13">2016</mods:date>
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<treatment id="0395E107FF9D680EFF60FF2C3622FC01" ID-DOI="http://doi.org/10.5281/zenodo.6073654" ID-GBIF-Taxon="119409831" ID-Zenodo-Dep="6073654" LSID="urn:lsid:plazi:treatment:0395E107FF9D680EFF60FF2C3622FC01" httpUri="http://treatment.plazi.org/id/0395E107FF9D680EFF60FF2C3622FC01" lastPageId="9" lastPageNumber="227" pageId="8" pageNumber="226">
<subSubSection id="C326039AFF9D680FFF60FF2C33ECFF52" pageId="8" pageNumber="226" type="nomenclature">
<paragraph id="8B835011FF9D680FFF60FF2C3399FF33" blockId="8.[151,491,151,211]" box="[151,491,151,178]" pageId="8" pageNumber="226">
<heading id="D0CBE77DFF9D680FFF60FF2C3399FF33" bold="true" box="[151,491,151,178]" fontSize="11" level="1" pageId="8" pageNumber="226" reason="1">
<emphasis id="B9488C03FF9D680FFF60FF2C3399FF33" bold="true" box="[151,491,151,178]" pageId="8" pageNumber="226">
<taxonomicName id="4C3C2B92FF9D680FFF60FF2C33F5FF30" ID-CoL="7427M" box="[151,391,151,177]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="monnei" status="sp. nov.">
<emphasis id="B9488C03FF9D680FFF60FF2C33F5FF30" bold="true" box="[151,391,151,177]" italics="true" pageId="8" pageNumber="226">Montesinus monnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B3178FF9D680FFE79FF233399FF33" box="[398,491,152,178]" pageId="8" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B835011FF9D680FFF60FF0133ECFF52" blockId="8.[151,491,151,211]" box="[151,414,186,211]" pageId="8" pageNumber="226">
(
<figureCitation id="13074C94FF9D680FFF69FF013354FF52" box="[158,294,186,211]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1457,1479]" captionTargetBox="[375,1210,212,1418]" captionTargetId="figure@3.[355,1232,193,1435]" captionTargetPageId="3" captionText="FIGURES 1 5. Habitus of Montesinus species. 1 3. Montesinus monnei sp. nov., holotype male in dorsal (1) and lateral views (2), and allotype female in dorsal view (3); 4. M. tatula sp. nov., holotype female in dorsal view; 5. M. machadoi Grossi sp. nov., holotype female in dorsal view. Scale bars = 4 mm." httpUri="https://zenodo.org/record/263681/files/figure.png" pageId="8" pageNumber="226">Figures 13</figureCitation>
,
<figureCitation id="13074C94FF9D680FFEC4FF01331CFF52" box="[307,366,186,211]" captionStart-0="FIGURES 6 12" captionStart-1="FIGURES 13 17" captionStart-2="FIGURES 18 21" captionStartId-0="4.[151,264,1936,1958]" captionStartId-1="5.[151,264,1971,1993]" captionStartId-2="6.[151,264,1971,1993]" captionTargetBox-0="[230,1352,195,1913]" captionTargetBox-1="[188,1400,197,1917]" captionTargetBox-2="[211,1360,196,1937]" captionTargetId-0="figure@4.[230,1352,195,1913]" captionTargetId-1="figure@5.[172,1416,193,1945]" captionTargetId-2="figure@6.[187,1392,193,1957]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." captionText-1="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." captionText-2="FIGURES 18 21. Left maxillae in dorsal view: 18 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/263682/files/figure.png" httpUri-1="https://zenodo.org/record/263683/files/figure.png" httpUri-2="https://zenodo.org/record/263684/files/figure.png" pageId="8" pageNumber="226">619</figureCitation>
,
<figureCitation id="13074C94FF9D680FFE8EFF0033E4FF52" box="[377,406,187,211]" captionStart="FIGURES 22 27" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[237,1354,210,1914]" captionTargetId="figure@7.[217,1369,193,1924]" captionTargetPageId="7" captionText="FIGURES 22 27. Details of morphology of Altitatiayus Weinreich and Montesinus gen. nov. 22 24. right female protibiae (punctures and setae omitted); 26 27. wings in dorsal view; 22. Montesinus monnei sp. nov.; 23. M. tatula sp. nov.; 24 25. M. machadoi sp. nov., 25. right wing; 26. Altitatiayus ruficollis (Lüderwaldt); 27. A. rotundatus (Boileau)." httpUri="https://zenodo.org/record/263685/files/figure.png" pageId="8" pageNumber="226">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326039AFF9D680FFF60FEB93107F916" pageId="8" pageNumber="226" type="materials_examined">
<paragraph id="8B835011FF9D680FFF60FEB936C9FE07" blockId="8.[151,1437,258,2011]" pageId="8" pageNumber="226">
<emphasis id="B9488C03FF9D680FFF60FEB933F6FE9A" bold="true" box="[151,388,258,283]" pageId="8" pageNumber="226">Material examined.</emphasis>
<typeStatus id="5487EEB3FF9D680FFE79FEB93384FE9A" box="[398,502,258,283]" pageId="8" pageNumber="226" type="holotype">Holotype</typeStatus>
male (dissected) labeled as follows: Handwritten white label, a) “BRASIL-Esp. Santo/Parque Nac. Caparaó/ Próx. Pico da Bandeira/ (
<quantity id="4CC4FDF4FF9D680FFCCBFE9D31ECFEBE" box="[828,926,294,319]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="226" unit="m" value="2500.0">2500 m</quantity>
)
<date id="FF8276D1FF9D680FFC40FE9C3640FEBE" box="[951,1074,295,319]" pageId="8" pageNumber="226" value="2001-04-13">13.IV.2001</date>
/ Fabio Oliveira Correia”; b) “
<taxonomicName id="4C3C2B92FF9D680FFF54FEF7330CFEE2" box="[163,382,332,355]" class="Insecta" family="Lucanidae" genus="Montesinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="monnei" status="sp. nov.">
<emphasis id="B9488C03FF9D680FFF54FEF7330CFEE2" box="[163,382,332,355]" italics="true" pageId="8" pageNumber="226">Montesinus monnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B3178FF9D680FFE70FEF033B1FEE2" box="[391,451,331,355]" pageId="8" pageNumber="226" rank="species">n. sp.</taxonomicNameLabel>
/♂
<typeStatus id="5487EEB3FF9D680FFE1EFEF030FDFEE2" box="[489,655,331,355]" pageId="8" pageNumber="226" type="holotype">HOLOTYPUS</typeStatus>
/ P. C. Grossi det. 2015”. Allotype female labeled as
<typeStatus id="5487EEB3FF9D680FFB1EFEF13739FEE2" box="[1257,1355,330,355]" pageId="8" pageNumber="226" type="holotype">holotype</typeStatus>
except for the female symbol and the ALLOTYPUS label.
<typeStatus id="5487EEB3FF9D680FFD2BFED53136FE06" box="[732,836,366,391]" pageId="8" pageNumber="226" type="holotype">Holotype</typeStatus>
and allotype deposited at
<collectionCode id="ED2DC8D4FF9D680FFB9EFED436C5FE07" box="[1129,1207,367,390]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="226">MNRJ</collectionCode>
.
</paragraph>
<paragraph id="8B835011FF9D680FFF30FE293107F916" blockId="8.[151,1437,258,2011]" pageId="8" pageNumber="226">
<emphasis id="B9488C03FF9D680FFF30FE2933B0FE2A" bold="true" box="[199,450,402,427]" pageId="8" pageNumber="226">
<typeStatus id="5487EEB3FF9D680FFF30FE293346FE2A" box="[199,308,402,427]" pageId="8" pageNumber="226" type="holotype">Holotype</typeStatus>
description
</emphasis>
. Total length, including mandibles, 19.0 mm. Greatest pronotal width 7.7 mm.
<emphasis id="B9488C03FF9D680FFAA6FE2937EEFE2A" box="[1361,1436,402,427]" italics="true" pageId="8" pageNumber="226">Color:</emphasis>
Dorsal surface dull black, pronotum and legs somewhat opaque ventrally with gular area, femora and trochanter reddish dark.
<emphasis id="B9488C03FF9D680FFEC1FE6133B0FE72" box="[310,450,474,499]" italics="true" pageId="8" pageNumber="226">Mouthparts:</emphasis>
Mandibles symmetrical, cylindrical in cross section, abruptly upturned; apex slightly incurved, acutely pointed and with an obscure internal tooth; integument finely reticulated; dorsal base with a convex process; basal tooth conical, slightly emarginate at posterior apex (
<figureCitation id="13074C94FF9D680FFC02FD99362FFDBA" box="[1013,1117,546,571]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1457,1479]" captionTargetBox="[375,1210,212,1418]" captionTargetId="figure@3.[355,1232,193,1435]" captionTargetPageId="3" captionText="FIGURES 1 5. Habitus of Montesinus species. 1 3. Montesinus monnei sp. nov., holotype male in dorsal (1) and lateral views (2), and allotype female in dorsal view (3); 4. M. tatula sp. nov., holotype female in dorsal view; 5. M. machadoi Grossi sp. nov., holotype female in dorsal view. Scale bars = 4 mm." httpUri="https://zenodo.org/record/263681/files/figure.png" pageId="8" pageNumber="226">Figs 12</figureCitation>
). Ventrally near basal tooth with a subtriangular punctate area. Labrum trapezoidal, with 3 anterior rounded processes; punctures concentrated at edge; anterior margin and anterior angles with yellowish setae. Mentum with sparse large punctures from middle to anterior edge. Maxillae with lacinia simple, not curved, and with no hooklets; palpi slender with 24 almost with the same length (
<figureCitation id="13074C94FF9D680FFE95FD09339AFD4A" box="[354,488,690,715]" captionStart-0="FIGURES 6 12" captionStart-1="FIGURES 13 17" captionStart-2="FIGURES 18 21" captionStartId-0="4.[151,264,1936,1958]" captionStartId-1="5.[151,264,1971,1993]" captionStartId-2="6.[151,264,1971,1993]" captionTargetBox-0="[230,1352,195,1913]" captionTargetBox-1="[188,1400,197,1917]" captionTargetBox-2="[211,1360,196,1937]" captionTargetId-0="figure@4.[230,1352,195,1913]" captionTargetId-1="figure@5.[172,1416,193,1945]" captionTargetId-2="figure@6.[187,1392,193,1957]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." captionText-1="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." captionText-2="FIGURES 18 21. Left maxillae in dorsal view: 18 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/263682/files/figure.png" httpUri-1="https://zenodo.org/record/263683/files/figure.png" httpUri-2="https://zenodo.org/record/263684/files/figure.png" pageId="8" pageNumber="226">Figs 1118</figureCitation>
).
<emphasis id="B9488C03FF9D680FFDF7FD093034FD4A" box="[512,582,690,715]" italics="true" pageId="8" pageNumber="226">Head:</emphasis>
Shape transverse, as wide as elytra; vertex broadly convex and scarcely punctate, punctures fine; disc depressed until anterior margin, more punctate, punctures fine to moderate. Canthi subtriangular in shape, externally projected. Anteriorly and above eyes with a convex process each side. Temporal process lacking, instead with a small convex area. Gena deeper than gula, disc convex and sparsely punctate; punctures fine. Gula anteriorly with integument microreticulate and sparse to moderately punctate; punctures fine to moderate; posteriorly with a subrectangular smooth area limited by two subparallel sutures.
<emphasis id="B9488C03FF9D680FFAD7FCDD37E9FCFE" box="[1312,1435,870,895]" italics="true" pageId="8" pageNumber="226">Prothorax:</emphasis>
Pronotum slightly convex, sparsely punctate elsewhere; punctures fine, more punctate near anterior angles; along posterior margin with moderate punctures; all pronotal angles rounded.
<taxonomicName id="4C3C2B92FF9D680FFC19FC1536CAFC46" authority="Shape" authorityName="Shape" box="[1006,1208,942,967]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="8" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="B9488C03FF9D680FFC19FC153617FC46" box="[1006,1125,942,967]" italics="true" pageId="8" pageNumber="226">Scutellum:</emphasis>
Shape
</taxonomicName>
subtriangular, very sparsely and finely punctate; surface anteriorly depressed.
<emphasis id="B9488C03FF9D680FFCDBFC69310EFC6A" box="[812,892,978,1003]" italics="true" pageId="8" pageNumber="226">Elytra:</emphasis>
Surface totally punctate; punctures fine to large (mostly moderate); near suture punctures fine, anteriorly larger. Each puncture with internal lateral wall, visible floor, and single setae. Punctures on elytral declivity with elongated setae, longer than puncture diameter. Humeri externally projected and rounded distally.
<emphasis id="B9488C03FF9D680FFD8BFB8430C9FBD7" box="[636,699,1087,1110]" italics="true" pageId="8" pageNumber="226">Legs:</emphasis>
Protibiae distally with 2 external strong teeth; medially to base with 68 smaller ones, 2 of these more developed. Mesotibiae with 3 external teeth, distal one much longer than the others; right tibia with the distal longer and left tibia with the proximal longer. Metatibiae with 1 external middle tooth.
<emphasis id="B9488C03FF9D680FFF29FB113323FB42" box="[222,337,1194,1219]" italics="true" pageId="8" pageNumber="226">Genitalia:</emphasis>
Genital capsule simple, rounded and sclerotized. Tergite with lateral paired plates quadrate, discal area more strongly sclerotized; between disc and lateral plates less membranous. Sternite setose at middle until discal area and with some scattered setae at posterior margin. Genitalia with simple genital capsule (
<figureCitation id="13074C94FF9D680FFAEEFB4937E2FA8A" box="[1305,1424,1266,1291]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 910</figureCitation>
). Aedeagus symmetrical, instead internal sac apex (
<figureCitation id="13074C94FF9D680FFD3FFAAD315CFAAE" box="[712,814,1302,1327]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 68</figureCitation>
,
<figureCitation id="13074C94FF9D680FFCCDFAAC31F6FAAE" box="[826,900,1303,1327]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1971,1993]" captionTargetBox="[188,1400,197,1917]" captionTargetId="figure@5.[172,1416,193,1945]" captionTargetPageId="5" captionText="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263683/files/figure.png" pageId="8" pageNumber="226">1317</figureCitation>
); basal piece strongly sclerotized ventrally with lateral apical sides directed backward being mostly membranous dorsally and slightly curved (
<figureCitation id="13074C94FF9D680FFB3EFA813761FAD2" box="[1225,1299,1338,1363]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 6</figureCitation>
,
<figureCitation id="13074C94FF9D680FFAD6FA80374DFAD2" box="[1313,1343,1339,1363]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1971,1993]" captionTargetBox="[188,1400,197,1917]" captionTargetId="figure@5.[172,1416,193,1945]" captionTargetPageId="5" captionText="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263683/files/figure.png" pageId="8" pageNumber="226">13</figureCitation>
); paired struts present, as long as basal piece length with apex wider and less sclerotized. Parameres large, leaf-like shaped and apically broad, 1.3x shorter than basal piece; ventrally in its curvature with a two-folded flange (
<figureCitation id="13074C94FF9D680FFAD8FA3937EAFA1A" box="[1327,1432,1410,1435]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 67</figureCitation>
,
<figureCitation id="13074C94FF9D680FFF60FA1D3292FA3E" box="[151,224,1446,1471]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1971,1993]" captionTargetBox="[188,1400,197,1917]" captionTargetId="figure@5.[172,1416,193,1945]" captionTargetPageId="5" captionText="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263683/files/figure.png" pageId="8" pageNumber="226">1315</figureCitation>
); internal ventral base with an outcurved process; dorsally with internal base projected (
<figureCitation id="13074C94FF9D680FFB11FA1D3740FA3E" box="[1254,1330,1446,1471]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 6</figureCitation>
,
<figureCitation id="13074C94FF9D680FFAB6FA1C37FCFA3E" box="[1345,1422,1446,1471]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1971,1993]" captionTargetBox="[188,1400,197,1917]" captionTargetId="figure@5.[172,1416,193,1945]" captionTargetPageId="5" captionText="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263683/files/figure.png" pageId="8" pageNumber="226">13, 15</figureCitation>
); external surface near apex with small, dense setae. Median lobe trapezoidal, strongly sclerotized and weakly, irregularly emarginate at apex; base with one symmetrical emargination. Dorsal cross bar transverse, straight. Internal sac permanently everted, shorter than tegmen (basal piece + parameres), almost membranous and partly coated by dense short setae; middle expanded laterally and apex with two asymmetrical, lateral projections covered by minute sclerotized spicules; internally with 4 paired sclerites, 2 elongate from base until base of apical part, and two subtriangularly shaped located at apex (
<figureCitation id="13074C94FF9D680FFD73F9C5309BF916" box="[644,745,1662,1687]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 68</figureCitation>
,
<figureCitation id="13074C94FF9D680FFD02F9C4311AF916" box="[757,872,1662,1687]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1971,1993]" captionTargetBox="[188,1400,197,1917]" captionTargetId="figure@5.[172,1416,193,1945]" captionTargetPageId="5" captionText="FIGURES 13 17. Montesinus monnei sp. nov. aedeagus. 13. ventral view; 14. dorsal view; 15. lateral view with internal sac omitted; 16. apex of internal sac in ventral view; 17. internal sac in dorsal view with setae omitted. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263683/files/figure.png" pageId="8" pageNumber="226">13, 1617</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C326039AFF9D680EFF30F919304CFD96" lastPageId="9" lastPageNumber="227" pageId="8" pageNumber="226" type="description">
<paragraph id="8B835011FF9D680EFF30F919304CFD96" blockId="8.[151,1437,258,2011]" lastBlockId="9.[151,1437,151,896]" lastPageId="9" lastPageNumber="227" pageId="8" pageNumber="226">
<emphasis id="B9488C03FF9D680FFF30F919307BF93A" bold="true" box="[199,521,1698,1723]" pageId="8" pageNumber="226">
Female (
<figureCitation id="13074C94FF9D680FFED9F919330AF93A" box="[302,376,1698,1723]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1457,1479]" captionTargetBox="[375,1210,212,1418]" captionTargetId="figure@3.[355,1232,193,1435]" captionTargetPageId="3" captionText="FIGURES 1 5. Habitus of Montesinus species. 1 3. Montesinus monnei sp. nov., holotype male in dorsal (1) and lateral views (2), and allotype female in dorsal view (3); 4. M. tatula sp. nov., holotype female in dorsal view; 5. M. machadoi Grossi sp. nov., holotype female in dorsal view. Scale bars = 4 mm." httpUri="https://zenodo.org/record/263681/files/figure.png" pageId="8" pageNumber="226">Figs 3</figureCitation>
,
<figureCitation id="13074C94FF9D680FFE70F91933D6F93A" box="[391,420,1698,1723]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">12</figureCitation>
,
<figureCitation id="13074C94FF9D680FFE45F91933BDF93A" box="[434,463,1698,1723]" captionStart="FIGURES 18 21" captionStartId="6.[151,264,1971,1993]" captionTargetBox="[211,1360,196,1937]" captionTargetId="figure@6.[187,1392,193,1957]" captionTargetPageId="6" captionText="FIGURES 18 21. Left maxillae in dorsal view: 18 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263684/files/figure.png" pageId="8" pageNumber="226">19</figureCitation>
,
<figureCitation id="13074C94FF9D680FFE2AF919338EF93A" box="[477,508,1698,1723]" captionStart="FIGURES 22 27" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[237,1354,210,1914]" captionTargetId="figure@7.[217,1369,193,1924]" captionTargetPageId="7" captionText="FIGURES 22 27. Details of morphology of Altitatiayus Weinreich and Montesinus gen. nov. 22 24. right female protibiae (punctures and setae omitted); 26 27. wings in dorsal view; 22. Montesinus monnei sp. nov.; 23. M. tatula sp. nov.; 24 25. M. machadoi sp. nov., 25. right wing; 26. Altitatiayus ruficollis (Lüderwaldt); 27. A. rotundatus (Boileau)." httpUri="https://zenodo.org/record/263685/files/figure.png" pageId="8" pageNumber="226">22</figureCitation>
).
</emphasis>
As male in general aspect, except by the following characters. Total length 15.0 mm. Greatest pronotal width 5.6 mm.
<emphasis id="B9488C03FF9D680FFDC8F97D30B9F95E" box="[575,715,1734,1759]" italics="true" pageId="8" pageNumber="226">Mouthparts:</emphasis>
Mandibles shorter than head, incurved; basal teeth lacking, apex acute with an internal concavity; each mandible with a single forward projected tooth, the left tooth somewhat sinuous, posteriorly to it abruptly curved; internal dorsal carina present, above it concave; external dorsal carina present only at mandibular base, incomplete and making with internal carina a weak depression. Labrum trapezoidal, irregularly sinuous in anterior margin, sparsely punctate and setose only anteriorly. Maxillae with lacinia hook-shaped right angled, apex slender, and with one spiral-shaped hooklet (
<figureCitation id="13074C94FF9D680FFB9EF8C136B1F812" box="[1129,1219,1914,1939]" captionStart="FIGURES 6 12" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[230,1352,195,1913]" captionTargetId="figure@4.[230,1352,195,1913]" captionTargetPageId="4" captionText="FIGURES 6 12. Montesinus monnei sp. nov. 6. ventral view of aedeagus; 7. dorsal view of parameres and internal sac; 8. internal sac in ventral view; 9 10. genital capsule in dorsal view (9) and ventral view (10); 11 12. left maxillae in dorsal view of male (11) and female (12). scale bars = 1 mm." httpUri="https://zenodo.org/record/263682/files/figure.png" pageId="8" pageNumber="226">Figs 12</figureCitation>
,
<figureCitation id="13074C94FF9D680FFB25F8C03683F812" box="[1234,1265,1915,1939]" captionStart="FIGURES 18 21" captionStartId="6.[151,264,1971,1993]" captionTargetBox="[211,1360,196,1937]" captionTargetId="figure@6.[187,1392,193,1957]" captionTargetPageId="6" captionText="FIGURES 18 21. Left maxillae in dorsal view: 18 19. M. monnei sp. nov., male (18) and female (19); 20. M. tatula sp. nov.; 21. M. machadoi sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/263684/files/figure.png" pageId="8" pageNumber="226">19</figureCitation>
). Mentum less transverse, anteriorly slightly sinuous; each lateral convex; moderately punctate, punctures large to coalescent, each puncture with a single elongate setae, more evident near margins.
<emphasis id="B9488C03FF9D680FFC32F8793679F85A" box="[965,1035,1986,2011]" italics="true" pageId="8" pageNumber="226">Head:</emphasis>
Smaller in size, ½ width of males head; broader than 1 elytron width. Disc at each side with concave impression, almost totally punctate; punctures large to coarse. Vertex posteriorly impunctate, anteriorly convex and strongly punctate; punctures large to coarse. Canthi not externally projected, just rounded, intruding eyes less than a half of eye length. Eyes closer to anterior pronotal margin. Gena not convex and more punctate, posteriorly setose and reddish. Gula broader with semicircular punctures, a transverse deep suture and reddish brown coloration.
<emphasis id="B9488C03FF9C680EFBB9FE9C36BBFEC1" box="[1102,1225,295,320]" italics="true" pageId="9" pageNumber="227">Prothorax:</emphasis>
Pronotum not as transverse, rounded at lateral edges, slightly convex and almost reddish except near lateral margins, black just before anterior and posterior angles; surface moderately punctate; punctures fine.
<taxonomicName id="4C3C2B92FF9C680EFBD9FED436EEFE09" box="[1070,1180,367,392]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="B9488C03FF9C680EFBD9FED436EEFE09" box="[1070,1180,367,392]" italics="true" pageId="9" pageNumber="227">Scutellum</emphasis>
</taxonomicName>
: smaller, less punctate and surface nearly smooth.
<emphasis id="B9488C03FF9C680EFE23FE2F3051FE2C" box="[468,547,404,429]" italics="true" pageId="9" pageNumber="227">Elytra:</emphasis>
Shape oblong and more elongate. Surface totally punctate; punctures smaller and shallow.
<emphasis id="B9488C03FF9C680EFEDEFE02331AFE51" box="[297,360,441,464]" italics="true" pageId="9" pageNumber="227">Legs:</emphasis>
Shape smaller. Protibiae wider, with 2 distal and only 2 proximal teeth. Mesotibiae less dentate, left tibia with only one external tooth, and right tibia with two teeth, the proximal tooth smaller. Metatibiae with teeth less acute, apex almost truncate.
</paragraph>
</subSubSection>
<subSubSection id="C326039AFF9C680EFF30FD9F305EFD05" pageId="9" pageNumber="227" type="etymology">
<paragraph id="8B835011FF9C680EFF30FD9F305EFD05" blockId="9.[151,1437,151,896]" pageId="9" pageNumber="227">
<emphasis id="B9488C03FF9C680EFF30FD9F333CFDBC" bold="true" box="[199,334,548,573]" pageId="9" pageNumber="227">Etymology.</emphasis>
The specific name is an homage to my friend Dr. Miguel Angel Monné Barrios, one of the greatest longhorn beetles specialist, and who kindly showed me the specimens here described, that were among the miscellaneous
<taxonomicName id="4C3C2B92FF9C680EFECAFDD733CAFD04" box="[317,440,620,645]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
at MNRJ.
</paragraph>
</subSubSection>
<subSubSection id="C326039AFF9C680EFF30FD343622FC01" pageId="9" pageNumber="227" type="discussion">
<paragraph id="8B835011FF9C680EFF30FD343622FC01" blockId="9.[151,1437,151,896]" pageId="9" pageNumber="227">
<emphasis id="B9488C03FF9C680EFF30FD343346FD29" bold="true" box="[199,308,655,680]" pageId="9" pageNumber="227">Remarks</emphasis>
. The specimens described here are the only two specimens known, and were apparently found dead, as the male has the right part of head, including canthus, eye and antennae, missing, as is the last tarsomere of the right metatarsus. The female is also damaged, since the right antennae, four last tarsomeres of left protarsus and two last tarsomeres of left mesotarsus are missing. This species can be distinguished from the other species by its smaller body size, opaque body surface and deeper, more distinct elytral punctures. It was also found near the highest place in the Serra do Caparaó, called Pico da Bandeira, while the second species described here is from another formation (Serra do Espinhaço), and the third from a lower place at Caparaó.
</paragraph>
</subSubSection>
</treatment>
</document>