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<document id="AB6BA5FBA2DA2B2995BBFE6FA5C46AF2" ID-DOI="10.11646/zootaxa.5012.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5158062" ID-ZooBank="56C6852D-AAE0-4B6B-AB57-919CD62DAEC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628057811575" checkinUser="plazi" docAuthor="Sim-Smith, Carina, Hickman, Cleveland &amp; Kelly, Michelle" docDate="2021" docId="D3075148FFD3FFF1FF678B6BB35BCBFB" docLanguage="en" docName="zootaxa.5012.1.1.pdf" docOrigin="Zootaxa 5012 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5012.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cacospongia hermanorum Sim-Smith &amp; Hickman &amp; Kelly 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="2F3E2930FFDBFFFBFFF08E7DB158CB02" masterDocTitle="New shallow-water sponges (Porifera) from the Galápagos Islands" masterLastPageNumber="71" masterPageNumber="1" pageNumber="9" updateTime="1699033904328" updateUser="plazi">
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<mods:title id="BE960655130CBD603C9BCFB9B3F44F5A">New shallow-water sponges (Porifera) from the Galápagos Islands</mods:title>
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<mods:namePart id="4AAFAC3495F71753DB689E9F107469BC">Sim-Smith, Carina</mods:namePart>
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<mods:namePart id="14F08AA37765094CA0BFF51F192150D1">Hickman, Cleveland</mods:namePart>
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<mods:nameIdentifier id="53DD823DC278B374D9B120B166A6DC19" type="email">hickman.c@rockbridge.net</mods:nameIdentifier>
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<mods:namePart id="CA6C31848AB9C80FE3DBD8DC86EB4396">Kelly, Michelle</mods:namePart>
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<mods:nameIdentifier id="EF4883CBD11485E01308FEE3D0AFBBD9" type="email">elly@niwa.co.nz</mods:nameIdentifier>
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<mods:title id="AEDA3899886BC72D13B4A59C8634B9B3">Zootaxa</mods:title>
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<mods:date id="716E53818467960CDEAA48DC9E8A6812">2021</mods:date>
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<treatment id="D3075148FFD3FFF1FF678B6BB35BCBFB" LSID="urn:lsid:plazi:treatment:D3075148FFD3FFF1FF678B6BB35BCBFB" httpUri="http://treatment.plazi.org/id/D3075148FFD3FFF1FF678B6BB35BCBFB" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="13B4B3D5FFD3FFF3FF678B6BB318CE32" box="[151,576,1302,1329]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="5B11E05EFFD3FFF3FF678B6BB318CE32" blockId="8.[151,576,1302,1364]" box="[151,576,1302,1329]" pageId="8" pageNumber="9">
<heading id="00595732FFD3FFF3FF678B6BB318CE32" bold="true" box="[151,576,1302,1329]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
<emphasis id="69DA3C4CFFD3FFF3FF678B6BB318CE32" bold="true" box="[151,576,1302,1329]" pageId="8" pageNumber="9">
<taxonomicName id="9CAE9BDDFFD3FFF3FF678B6BB083CE33" ID-CoL="8NFLP" authority="Sim-Smith &amp; Hickman &amp; Kelly, 2021" authorityName="Sim-Smith &amp; Hickman &amp; Kelly" authorityYear="2021" box="[151,475,1302,1329]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="8" pageNumber="9" phylum="Porifera" rank="species" species="hermanorum" status="sp. nov.">
<emphasis id="69DA3C4CFFD3FFF3FF678B6BB083CE33" bold="true" box="[151,475,1302,1329]" italics="true" pageId="8" pageNumber="9">Cacospongia hermanorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72E98137FFD3FFF3FE128B6BB318CE32" box="[482,576,1302,1328]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13B4B3D5FFD3FFF3FF678B47B1B3CE56" box="[151,235,1338,1364]" pageId="8" pageNumber="9" type="description">
<paragraph id="5B11E05EFFD3FFF3FF678B47B1B3CE56" blockId="8.[151,576,1302,1364]" box="[151,235,1338,1364]" pageId="8" pageNumber="9">
(
<figureCitation id="C395FCDBFFD3FFF3FF508B47B1BBCE56" box="[160,227,1338,1364]" captionStart="FIGURE 4" captionStartId="9.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-91@9.[151,1436,345,1811]" captionTargetPageId="9" captionText="FIGURE 4. Cacospongia hermanorum sp. nov., holotype, MCCDRS9434: A. Holotype, deck photo. B. Close up of the surface of the holotype, deck photo. C. Cut side view of the holotype, preserved. D. Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation. E. A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow). F. Cross section of a fibre showing the concentric laminations." figureDoi="http://doi.org/10.5281/zenodo.5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13B4B3D5FFD3FFF3FF678BFFB2DCCEE6" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="5B11E05EFFD3FFF3FF678BFFB424CE9E" blockId="8.[151,1437,1410,2013]" box="[151,1404,1410,1437]" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF678BFFB0D9CE9E" bold="true" box="[151,385,1410,1436]" pageId="8" pageNumber="9">Material examined.</emphasis>
<materialsCitation id="EBC6EA03FFD3FFF3FE778BFFB424CE9E" box="[391,1404,1410,1437]" collectingDate="2004-02-03" elevation="6" latitude="-0.867" location="Los Hermanos Islands" longLatPrecision="78" longitude="-90.782" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="69DA3C4CFFD3FFF3FE778BFFB35FCE9E" box="[391,519,1410,1436]" italics="true" pageId="8" pageNumber="9">
<typeStatus id="84155EFCFFD3FFF3FE778BFFB0B2CE9E" box="[391,490,1410,1436]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
MCCDRS9434,
<location id="5E71B685FFD3FFF3FD328BFEB2E7CE9E" LSID="urn:lsid:plazi:treatment:D3075148FFD3FFF1FF678B6BB35BCBFB:5E71B685FFD3FFF3FD328BFEB2E7CE9E" box="[706,959,1410,1437]" latitude="-0.867" longLatPrecision="78" longitude="-90.782" name="Los Hermanos Islands" pageId="8" pageNumber="9">Los Hermanos Islands</location>
,
<geoCoordinate id="3E9A8699FFD3FFF3FC3A8BFFB574CE9E" box="[970,1068,1410,1436]" degrees="0.867" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="55" value="-0.867">0.867° S</geoCoordinate>
,
<geoCoordinate id="3E9A8699FFD3FFF3FBC98BFFB5E8CE9F" box="[1081,1200,1410,1437]" degrees="90.782" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="55" value="-90.782">90.782° W</geoCoordinate>
,
<quantity id="9C564DBBFFD3FFF3FB4D8BFFB5B2CE9F" box="[1213,1258,1410,1437]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="m" value="6.0">
<elevation id="D083076DFFD3FFF3FB4D8BFFB5B2CE9F" box="[1213,1258,1410,1437]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<date id="2F10C69EFFD3FFF3FB068BFFB420CE9E" box="[1270,1400,1410,1436]" pageId="8" pageNumber="9" value="2004-02-03">
<collectingDate id="3F543F76FFD3FFF3FB068BFFB420CE9E" box="[1270,1400,1410,1436]" pageId="8" pageNumber="9" value="2004-02-03">3 Feb 2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="5B11E05EFFD3FFF3FF378BDBB336CEC2" blockId="8.[151,1437,1410,2013]" box="[199,622,1446,1473]" pageId="8" pageNumber="9">
<materialsCitation id="EBC6EA03FFD3FFF3FF378BDBB332CEC2" box="[199,618,1446,1473]" location="Los Hermanos Islands" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="69DA3C4CFFD3FFF3FF378BDBB03ECEC2" bold="true" box="[199,358,1446,1472]" pageId="8" pageNumber="9">
<typeStatus id="84155EFCFFD3FFF3FF378BDBB05ACEC2" box="[199,258,1446,1472]" pageId="8" pageNumber="9">Type</typeStatus>
locality.
</emphasis>
<location id="5E71B685FFD3FFF3FE9C8BDAB332CEC2" LSID="urn:lsid:plazi:treatment:D3075148FFD3FFF1FF678B6BB35BCBFB:5E71B685FFD3FFF3FE9C8BDAB332CEC2" box="[364,618,1446,1473]" name="Los Hermanos Islands" pageId="8" pageNumber="9">Los Hermanos Islands</location>
</materialsCitation>
.
</paragraph>
<paragraph id="5B11E05EFFD3FFF3FF378BB7B2DCCEE6" blockId="8.[151,1437,1410,2013]" box="[199,900,1482,1509]" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF378BB7B0ABCEE6" bold="true" box="[199,499,1482,1508]" pageId="8" pageNumber="9">Habitat and distribution.</emphasis>
Only known from the
<typeStatus id="84155EFCFFD3FFF3FD038BB6B27BCEE7" box="[755,803,1483,1509]" pageId="8" pageNumber="9">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="13B4B3D5FFD3FFF3FF378B93B5B2CC73" pageId="8" pageNumber="9" type="description">
<paragraph id="5B11E05EFFD3FFF3FF378B93B0B6CDBF" blockId="8.[151,1437,1410,2013]" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF378B93B001CD0A" bold="true" box="[199,345,1518,1544]" pageId="8" pageNumber="9">Description.</emphasis>
A flattened, cushion-shaped sponge,
<quantity id="9C564DBBFFD3FFF3FCF88B93B202CD0B" box="[776,858,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="50.0">50 mm</quantity>
long ×
<quantity id="9C564DBBFFD3FFF3FC478B93B551CD0B" box="[951,1033,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="40.0">40 mm</quantity>
wide ×
<quantity id="9C564DBBFFD3FFF3FB9B8B93B5E5CD0B" box="[1131,1213,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="15.0">15 mm</quantity>
thick (
<figureCitation id="C395FCDBFFD3FFF3FAE08B93B4D7CD0A" box="[1296,1423,1518,1544]" captionStart="FIGURE 4" captionStartId="9.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-91@9.[151,1436,345,1811]" captionTargetPageId="9" captionText="FIGURE 4. Cacospongia hermanorum sp. nov., holotype, MCCDRS9434: A. Holotype, deck photo. B. Close up of the surface of the holotype, deck photo. C. Cut side view of the holotype, preserved. D. Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation. E. A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow). F. Cross section of a fibre showing the concentric laminations." figureDoi="http://doi.org/10.5281/zenodo.5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, C</figureCitation>
). Surface is finely conulose, with conules around
<quantity id="9C564DBBFFD3FFF3FD5C886FB3B4CD2F" box="[684,748,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="1.0">1 mm</quantity>
high. The conules are formed by collagen fibrils that protrude beyond the surface of the sponge, coming together to form the apex of the conules (
<figureCitation id="C395FCDBFFD3FFF3FBBF884BB5F1CD52" box="[1103,1193,1590,1616]" captionStart="FIGURE 4" captionStartId="9.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-91@9.[151,1436,345,1811]" captionTargetPageId="9" captionText="FIGURE 4. Cacospongia hermanorum sp. nov., holotype, MCCDRS9434: A. Holotype, deck photo. B. Close up of the surface of the holotype, deck photo. C. Cut side view of the holotype, preserved. D. Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation. E. A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow). F. Cross section of a fibre showing the concentric laminations." figureDoi="http://doi.org/10.5281/zenodo.5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="8" pageNumber="9">Fig. 4B</figureCitation>
). A few small, inconspicuous oscules,
<quantity id="9C564DBBFFD3FFF3FE908827B0E5CD77" box="[352,445,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
in diameter, are present on the upper surface of the sponge. The surface is densely punctate below the surface reticulation of fibres. Texture is very firm and harsh, easily torn. Colour in life and ethanol is blackish brown, interior is tan.
</paragraph>
<paragraph id="5B11E05EFFD3FFF3FF3788BBB5B2CC73" blockId="8.[151,1437,1410,2013]" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF3788BBB06DCDE2" bold="true" box="[199,309,1734,1760]" pageId="8" pageNumber="9">Skeleton.</emphasis>
The skeleton is a well-developed, irregular reticulation of concentrically laminated, yellow-brown fibres that are slightly fasciculated just below the surface (
<figureCitation id="C395FCDBFFD3FFF3FCD28897B2C2CC07" box="[802,922,1770,1797]" captionStart="FIGURE 4" captionStartId="9.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-91@9.[151,1436,345,1811]" captionTargetPageId="9" captionText="FIGURE 4. Cacospongia hermanorum sp. nov., holotype, MCCDRS9434: A. Holotype, deck photo. B. Close up of the surface of the holotype, deck photo. C. Cut side view of the holotype, preserved. D. Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation. E. A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow). F. Cross section of a fibre showing the concentric laminations." figureDoi="http://doi.org/10.5281/zenodo.5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="8" pageNumber="9">Fig. 4D, F</figureCitation>
). Primary fibres are lightly cored with foreign spicules (
<figureCitation id="C395FCDBFFD3FFF3FEF18973B002CC2A" box="[257,346,1806,1832]" captionStart="FIGURE 4" captionStartId="9.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-91@9.[151,1436,345,1811]" captionTargetPageId="9" captionText="FIGURE 4. Cacospongia hermanorum sp. nov., holotype, MCCDRS9434: A. Holotype, deck photo. B. Close up of the surface of the holotype, deck photo. C. Cut side view of the holotype, preserved. D. Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation. E. A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow). F. Cross section of a fibre showing the concentric laminations." figureDoi="http://doi.org/10.5281/zenodo.5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="8" pageNumber="9">Fig. 4E</figureCitation>
); 70140 µm in diameter and separated by a distance of 4001300 µm. Secondary fibres are uncored, 30100 µm in diameter. They typically form polygonal meshes that are 80700 µm in size. Foreign spicules are lightly scattered throughout the choanosome, being more common at the surface of the sponge.
</paragraph>
</subSubSection>
<subSubSection id="13B4B3D5FFD3FFF3FF378907B5B2CC96" box="[199,1258,1914,1941]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="5B11E05EFFD3FFF3FF378907B5B2CC96" blockId="8.[151,1437,1410,2013]" box="[199,1258,1914,1941]" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF378907B016CC96" bold="true" box="[199,334,1914,1940]" pageId="8" pageNumber="9">Etymology.</emphasis>
Named for the
<typeStatus id="84155EFCFFD3FFF3FE0D8906B375CC97" box="[509,557,1915,1941]" pageId="8" pageNumber="9">type</typeStatus>
locality, Los Hermanos Islands in the
<collectingRegion id="996A2EBCFFD3FFF3FC2C8907B50ACC96" box="[988,1106,1914,1940]" country="Ecuador" name="Galapagos" pageId="8" pageNumber="9">Galápagos</collectingRegion>
Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="13B4B3D5FFD3FFF1FF3789E3B35BCBFB" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="discussion">
<paragraph id="5B11E05EFFD3FFF2FF3789E3B30DCA43" blockId="8.[151,1437,1410,2013]" lastBlockId="9.[151,1437,151,321]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="69DA3C4CFFD3FFF3FF3789E3B063CCBA" bold="true" box="[199,315,1950,1976]" pageId="8" pageNumber="9">Remarks.</emphasis>
<bibRefCitation id="3F3F9DAFFFD3FFF3FEB089E3B276CCBA" author="Desqueyroux-Faundez, R. &amp; Van Soest, R. W. M." box="[320,814,1950,1977]" pageId="8" pageNumber="9" pagination="379 - 467" refId="ref39327" refString="Desqueyroux-Faundez, R. &amp; Van Soest, R. W. M. (1997) Shallow water demosponges of the Galapagos Islands. Revue Suisse de Zoologie, 104, 379 - 467. https: // doi. org / 10.5962 / bhl. part. 80003" type="journal article" year="1997">Desqueyroux-Faúndez and Van Soest (1997)</bibRefCitation>
described two species of
<taxonomicName id="9CAE9BDDFFD3FFF3FBB789E2B583CCBB" authorityName="Schmidt" authorityYear="1862" box="[1095,1243,1951,1977]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="8" pageNumber="9" phylum="Porifera" rank="genus">
<emphasis id="69DA3C4CFFD3FFF3FBB789E2B583CCBB" box="[1095,1243,1951,1977]" italics="true" pageId="8" pageNumber="9">Cacospongia</emphasis>
</taxonomicName>
from the
<collectingRegion id="996A2EBCFFD3FFF3FAB489E3B04ECCDE" country="Ecuador" name="Galapagos" pageId="8" pageNumber="9">Galápagos Islands</collectingRegion>
,
<taxonomicName id="9CAE9BDDFFD3FFF3FED089BEB34CCCDF" authority="Thiele 1905" authorityName="Thiele" authorityYear="1905" box="[288,532,1986,2013]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="8" pageNumber="9" phylum="Porifera" rank="species" species="similis">
<emphasis id="69DA3C4CFFD3FFF3FED089BEB0D1CCDE" box="[288,393,1986,2013]" italics="true" pageId="8" pageNumber="9">C. similis</emphasis>
<bibRefCitation id="3F3F9DAFFFD3FFF3FE7E89BFB34CCCDF" author="Thiele, J." box="[398,532,1986,2013]" pageId="8" pageNumber="9" pagination="407 - 496" refId="ref42156" refString="Thiele, J. (1905) Die Kiesel- und Hornschwamme der Sammlung Plate. Zoologische Jahrbucher, Supplement 6, 407 - 496. [in German]" type="journal article" year="1905">Thiele 1905</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="9CAE9BDDFFD3FFF3FDB889BEB5E6CCDE" authority="Desqueyroux-Faundez and Van Soest, 1997" authorityName="Desqueyroux-Faundez and Van Soest" authorityYear="1997" box="[584,1214,1986,2013]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="8" pageNumber="9" phylum="Porifera" rank="species" species="incognita">
<emphasis id="69DA3C4CFFD3FFF3FDB889BEB389CCDE" box="[584,721,1987,2013]" italics="true" pageId="8" pageNumber="9">C. incognita</emphasis>
<bibRefCitation id="3F3F9DAFFFD3FFF3FD2789BFB5E6CCDE" author="Desqueyroux-Faundez, R. &amp; Van Soest, R. W. M." box="[727,1214,1986,2013]" pageId="8" pageNumber="9" pagination="379 - 467" refId="ref39327" refString="Desqueyroux-Faundez, R. &amp; Van Soest, R. W. M. (1997) Shallow water demosponges of the Galapagos Islands. Revue Suisse de Zoologie, 104, 379 - 467. https: // doi. org / 10.5962 / bhl. part. 80003" type="journal article" year="1997">Desqueyroux-Faúndez and Van Soest, 1997</bibRefCitation>
</taxonomicName>
, both of which have subsequently been transferred to the genus
<taxonomicName id="9CAE9BDDFFD2FFF2FD7A8EEAB26CCBB3" authorityName="Cook &amp; Bergquist" authorityYear="2000" box="[650,820,151,177]" class="Demospongiae" family="Thorectidae" genus="Scalarispongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="genus">
<emphasis id="69DA3C4CFFD2FFF2FD7A8EEAB26CCBB3" box="[650,820,151,177]" italics="true" pageId="9" pageNumber="10">Scalarispongia</emphasis>
</taxonomicName>
due to their regular, ladder-like reticulation of fibres.
<taxonomicName id="9CAE9BDDFFD2FFF2FF678EC6B36BCBD7" authority="(Thiele 1905)" baseAuthorityName="Thiele" baseAuthorityYear="1905" box="[151,563,186,213]" class="Demospongiae" family="Thorectidae" genus="Scalarispongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="similis">
<emphasis id="69DA3C4CFFD2FFF2FF678EC6B0C9CBD7" box="[151,401,187,213]" italics="true" pageId="9" pageNumber="10">Scalarispongia similis</emphasis>
(
<bibRefCitation id="3F3F9DAFFFD2FFF2FE508EC6B372CBD7" author="Thiele, J." box="[416,554,186,213]" pageId="9" pageNumber="10" pagination="407 - 496" refId="ref42156" refString="Thiele, J. (1905) Die Kiesel- und Hornschwamme der Sammlung Plate. Zoologische Jahrbucher, Supplement 6, 407 - 496. [in German]" type="journal article" year="1905">Thiele 1905</bibRefCitation>
)
</taxonomicName>
also has much thinner primary (4070 µm) and secondary (3040 µm) fibres than
<taxonomicName id="9CAE9BDDFFD2FFF2FF218EA2B0DECBFB" authority="Sim-Smith &amp; Hickman &amp; Kelly, 2021" authorityName="Sim-Smith &amp; Hickman &amp; Kelly" authorityYear="2021" box="[209,390,223,249]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="hermanorum" status="sp. nov.">
<emphasis id="69DA3C4CFFD2FFF2FF218EA2B0DECBFB" box="[209,390,223,249]" italics="true" pageId="9" pageNumber="10">C. hermanorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72E98137FFD2FFF2FE7F8EA2B0B5CBFB" box="[399,493,223,249]" pageId="9" pageNumber="10" rank="species">
<emphasis id="69DA3C4CFFD2FFF2FE7F8EA2B0BECBFB" bold="true" box="[399,486,223,249]" pageId="9" pageNumber="10">sp. nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="9CAE9BDDFFD2FFF2FE078EA2B24CCBFA" baseAuthorityName="Desqueyroux-Faundez &amp; van Soest" baseAuthorityYear="1997" box="[503,788,223,249]" class="Demospongiae" family="Thorectidae" genus="Scalarispongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="incognita">
<emphasis id="69DA3C4CFFD2FFF2FE078EA2B24CCBFA" box="[503,788,223,249]" italics="true" pageId="9" pageNumber="10">Scalarispongia incognita</emphasis>
</taxonomicName>
has similar sized primary (120150 µm) and secondary fibres (50100 µm) to
<taxonomicName id="9CAE9BDDFFD2FFF2FE678F7EB311CA1F" authority="Sim-Smith &amp; Hickman &amp; Kelly, 2021" authorityName="Sim-Smith &amp; Hickman &amp; Kelly" authorityYear="2021" box="[407,585,259,285]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="hermanorum" status="sp. nov.">
<emphasis id="69DA3C4CFFD2FFF2FE678F7EB311CA1F" box="[407,585,259,285]" italics="true" pageId="9" pageNumber="10">C. hermanorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72E98137FFD2FFF2FDA08F7EB3F2CA1F" box="[592,682,259,285]" pageId="9" pageNumber="10" rank="species">
<emphasis id="69DA3C4CFFD2FFF2FDA08F7EB3FCCA1F" bold="true" box="[592,676,259,285]" pageId="9" pageNumber="10">sp. nov</emphasis>
.
</taxonomicNameLabel>
, but
<taxonomicName id="9CAE9BDDFFD2FFF2FD148F7EB233CA1E" baseAuthorityName="Desqueyroux-Faundez &amp; van Soest" baseAuthorityYear="1997" box="[740,875,259,285]" class="Demospongiae" family="Thorectidae" genus="Scalarispongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="incognita">
<emphasis id="69DA3C4CFFD2FFF2FD148F7EB233CA1E" box="[740,875,259,285]" italics="true" pageId="9" pageNumber="10">S. incognita</emphasis>
</taxonomicName>
has a thick organic, heavily pigmented ectosomal membrane, and a creamy white interior.
</paragraph>
<caption id="0FD1B0D6FFD2FFF2FF678957B430CCAC" ID-DOI="http://doi.org/10.5281/zenodo.5158075" ID-Zenodo-Dep="5158075" httpUri="https://zenodo.org/record/5158075/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1834,1858]" targetBox="[151,1436,345,1811]" targetPageId="9">
<paragraph id="5B11E05EFFD2FFF2FF678957B430CCAC" blockId="9.[151,1437,1834,1966]" pageId="9" pageNumber="10">
<emphasis id="69DA3C4CFFD2FFF2FF678957B04ACC40" bold="true" box="[151,274,1834,1858]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
<taxonomicName id="9CAE9BDDFFD2FFF2FEE88957B37ECC40" authority="Sim-Smith &amp; Hickman &amp; Kelly, 2021" authorityName="Sim-Smith &amp; Hickman &amp; Kelly" authorityYear="2021" box="[280,550,1834,1858]" class="Demospongiae" family="Thorectidae" genus="Cacospongia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="10" phylum="Porifera" rank="species" species="hermanorum" status="sp. nov.">
<emphasis id="69DA3C4CFFD2FFF2FEE88957B37ECC40" box="[280,550,1834,1858]" italics="true" pageId="9" pageNumber="10">Cacospongia hermanorum</emphasis>
</taxonomicName>
<emphasis id="69DA3C4CFFD2FFF2FDDC8957B3DACC40" bold="true" box="[556,642,1834,1858]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="72E98137FFD2FFF2FDDC8957B326CC40" box="[556,638,1834,1858]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype, MCCDRS9434:
<emphasis id="69DA3C4CFFD2FFF2FC658957B2F6CC40" bold="true" box="[917,942,1834,1858]" pageId="9" pageNumber="10">A.</emphasis>
Holotype, deck photo.
<emphasis id="69DA3C4CFFD2FFF2FB698957B5E8CC40" bold="true" box="[1177,1200,1834,1858]" pageId="9" pageNumber="10">B.</emphasis>
Close up of the surface of the holotype, deck photo.
<emphasis id="69DA3C4CFFD2FFF2FE4C8933B08CCC64" bold="true" box="[444,468,1870,1894]" pageId="9" pageNumber="10">C.</emphasis>
Cut side view of the holotype, preserved.
<emphasis id="69DA3C4CFFD2FFF2FC758933B2C6CC64" bold="true" box="[901,926,1870,1894]" pageId="9" pageNumber="10">D.</emphasis>
Vertical cross section showing the fibres that protrude through the surface of the sponge and the well-developed fibre reticulation.
<emphasis id="69DA3C4CFFD2FFF2FC3D890FB2BCCC88" bold="true" box="[973,996,1906,1930]" pageId="9" pageNumber="10">E.</emphasis>
A primary fibre cored with foreign spicules (white arrow) and uncored secondary fibres (black arrow).
<emphasis id="69DA3C4CFFD2FFF2FD1E89EBB259CCAC" bold="true" box="[750,769,1942,1966]" pageId="9" pageNumber="10">F.</emphasis>
Cross section of a fibre showing the concentric laminations.
</paragraph>
</caption>
<paragraph id="5B11E05EFFD1FFF1FF678EEAB2E7CBB3" blockId="10.[151,959,150,177]" box="[151,959,150,177]" pageId="10" pageNumber="11">
<emphasis id="69DA3C4CFFD1FFF1FF678EEAB2E7CBB3" bold="true" box="[151,959,150,177]" pageId="10" pageNumber="11">
Subclass
<taxonomicName id="9CAE9BDDFFD1FFF1FEFA8EEAB2E7CBB3" authority="Cardenas, Perez &amp; Boury-Esnault" authorityName="Cardenas, Perez &amp; Boury-Esnault" box="[266,959,150,177]" pageId="10" pageNumber="11" rank="subClass" subClass="Heteroscleromorpha">Heteroscleromorpha Cárdenas, Pérez &amp; Boury-Esnault</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="5B11E05EFFD1FFF1FF678EA2B35BCBFB" blockId="10.[151,515,223,249]" box="[151,515,223,249]" pageId="10" pageNumber="11">
<emphasis id="69DA3C4CFFD1FFF1FF678EA2B35BCBFB" bold="true" box="[151,515,223,249]" pageId="10" pageNumber="11">
<collectingRegion id="996A2EBCFFD1FFF1FF678EA2B1BDCBFB" box="[151,229,223,249]" country="American Samoa" name="order" pageId="10" pageNumber="11">Order</collectingRegion>
<taxonomicName id="9CAE9BDDFFD1FFF1FF1C8EA2B35BCBFB" authority="Topsent" authorityName="Topsent" box="[236,515,223,249]" class="Demospongiae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplosclerida" pageId="10" pageNumber="11" phylum="Porifera" rank="order">Haplosclerida Topsent</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>