135 lines
21 KiB
XML
135 lines
21 KiB
XML
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<mods:title id="0451E36EE58D9183360EBC4934F6F059">Review of the subtribe Gyrophaenina Kraatz 1856 (Coleoptera: Staphylinidae: Aleocharinae: Homalotini) of Middle Asia, with emphasis on the fauna of Kazakhstan</mods:title>
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<emphasis id="26D7EAEFFFA00E7DC7C1FBCCFC66FB20" bold="true" box="[151,970,1150,1177]" pageId="9" pageNumber="388">
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Key to the
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<taxonomicName id="D3A34D7EFFA00E7DC649FBCDFE19FB20" ID-CoL="8VRQ3" authorityName="Mannerheim" authorityYear="1830" box="[287,437,1150,1177]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="26D7EAEFFFA00E7DC649FBCDFE19FB20" bold="true" box="[287,437,1150,1177]" italics="true" pageId="9" pageNumber="388">Gyrophaena</emphasis>
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species of the
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<taxonomicName id="D3A34D7EFFA00E7DC53EFBCDFD17FB20" authorityName="Casey" authorityYear="1906" box="[616,699,1150,1177]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="388" phylum="Arthropoda" rank="species" species="laetula">
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<emphasis id="26D7EAEFFFA00E7DC53EFBCDFD17FB20" bold="true" box="[616,699,1150,1177]" italics="true" pageId="9" pageNumber="388">laetula</emphasis>
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</taxonomicName>
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group of
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<collectingCountry id="6CB4766DFFA00E7DC462FBCCFC66FB20" box="[820,970,1151,1177]" name="Kazakhstan" pageId="9" pageNumber="388">Kazakhstan</collectingCountry>
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</emphasis>
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</heading>
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<subSubSection id="5CB96576FFA00E7DC7C1FB70FA30F9EF" pageId="9" pageNumber="388" type="key">
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<keyStep id="AF572E58FFA00E7DC7C1FB70FA30FA1C" pageId="9" pageNumber="388">
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<paragraph id="141C36FDFFA00E7DC7C1FB70FA30FA89" blockId="9.[151,1437,1219,1622]" pageId="9" pageNumber="388">
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<keyLead id="AF5295C8FFA00E7DC7C1FB70FA30FA89" pageId="9" pageNumber="388">
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1 Pronotum with several small, scattered punctures not forming longitudinal rows and two rather large punctures near base of pronotum. Elytra between cells of microsculpture without micropunctures, but with several distinct tubercles. Medial teeth of male abdominal tergite VIII quite wide and long (
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<figureCitation id="8C982A78FFA00E7DC597FB4EFC9AFAAD" box="[705,822,1277,1300]" captionStart="FIGURES 34–38" captionStartId="10.[151,264,1943,1968]" captionTargetBox="[173,1405,196,1906]" captionTargetId="figure@10.[168,1419,181,1923]" captionTargetPageId="10" captionText="FIGURES 34–38. Gyrophaena aryanamensis, sp. nov. (Figs 34: Holotype, TAJIKISTAN, Sharak; 35–38: Paratype, KYR- GYZSTAN, Dzhany-Bazar): 34—habitus, dorsal view, 35, 37—aedeagus, lateral view, 36, 38—male abdominal tergite VIII, dorsal view. Scale bars: 1 mm (34), 0.1 mm (35–38)." httpUri="https://zenodo.org/record/3238880/files/figure.png" pageId="9" pageNumber="388">Figs. 36, 38</figureCitation>
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). Ventral plate of aedeagus markedly wider and short, distinctly angled near the base, quite blunted apically (
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<figureCitation id="8C982A78FFA00E7DC5D8FAA8FCADFA88" box="[654,769,1307,1329]" captionStart="FIGURES 34–38" captionStartId="10.[151,264,1943,1968]" captionTargetBox="[173,1405,196,1906]" captionTargetId="figure@10.[168,1419,181,1923]" captionTargetPageId="10" captionText="FIGURES 34–38. Gyrophaena aryanamensis, sp. nov. (Figs 34: Holotype, TAJIKISTAN, Sharak; 35–38: Paratype, KYR- GYZSTAN, Dzhany-Bazar): 34—habitus, dorsal view, 35, 37—aedeagus, lateral view, 36, 38—male abdominal tergite VIII, dorsal view. Scale bars: 1 mm (34), 0.1 mm (35–38)." httpUri="https://zenodo.org/record/3238880/files/figure.png" pageId="9" pageNumber="388">Figs. 35, 37</figureCitation>
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)..................................
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<taxonomicName id="D3A34D7EFFA00E7DC3F0FAA8FAE4FA88" ID-CoL="8Y45Z" authority="Enushchenko, 2019" authorityName="Enushchenko" authorityYear="2019" box="[1190,1352,1307,1329]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="388" phylum="Arthropoda" rank="species" species="aryanamensis" status="sp. nov.">
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<emphasis id="26D7EAEFFFA00E7DC3F0FAA8FAE4FA88" box="[1190,1352,1307,1329]" italics="true" pageId="9" pageNumber="388">G. aryanamensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3DE45794FFA00E7DC218FAA8FA30FA89" box="[1358,1436,1307,1328]" pageId="9" pageNumber="388" rank="species">
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<emphasis id="26D7EAEFFFA00E7DC218FAA8FAC9FA89" bold="true" box="[1358,1381,1307,1328]" pageId="9" pageNumber="388">sp</emphasis>
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.
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<emphasis id="26D7EAEFFFA00E7DC227FAA8FA3AFA89" bold="true" box="[1393,1430,1307,1328]" pageId="9" pageNumber="388">nov</emphasis>
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.
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="141C36FDFFA00E7DC7C1FA8AFA30FA1C" blockId="9.[151,1437,1219,1622]" pageId="9" pageNumber="388">
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<keyLead id="AF5295C8FFA00E7DC7C1FA8AFA30FA1C" pageId="9" pageNumber="388">
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- Pronotum with two longitudinal rows of punctures, usually with several additional punctures closer to lateral margins of pronotum. Elytra between cells of microsculpture with sparse, small shallow punctation and few tubercles only on external posterior angles. Median teeth of male abdominal tergite VIII narrower and rather small. Ventral process of aedeagus more elongated, arched, quite acute apically (
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<figureCitation id="8C982A78FFA00E7DC6A0FA23FDEAFA1C" box="[502,582,1424,1446]" captionStart="FIGURES 39–46" captionStartId="15.[151,264,1884,1909]" captionTargetBox="[224,1372,204,1780]" captionTargetId="figure@15.[205,1382,181,1864]" captionTargetPageId="15" captionText="FIGURES 39–46. Gyrophaena aryanamensis, sp. nov. (Figs. 39, 40: KYRGYZSTAN, Dzhany-Bazar) and Gyrophaena hochhuthi Bernhauer (Figs. 41–46: KYRGYZSTAN, Dzhany-Bazar): 41—habitus, dorsal view, 42—aedeagus, lateral view, 43, 44—male abdominal tergite VIII, dorsal view, 39, 45—female abdominal tergite VIII, dorsal view, 40, 46—female abdominal sternite VIII, dorsal view. Scale bars: 1 mm (41), 0.1 mm (39, 40, 42–46)." httpUri="https://zenodo.org/record/3238882/files/figure.png" pageId="9" pageNumber="388">Figs. 42</figureCitation>
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,
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<figureCitation id="8C982A78FFA00E7DC506FA23FDC4FA1C" box="[592,616,1424,1445]" captionStart="FIGURES 47–50" captionStartId="16.[151,264,1927,1952]" captionTargetBox="[186,1362,200,1802]" captionTargetId="figure@16.[186,1400,181,1907]" captionTargetPageId="16" captionText="FIGURES 47–50. Gyrophaena hochhuthi Bernhauer: 47—aedeagus, lateral view, 48—male abdominal tergite VIII, dorsal view (Figs. 47, 48: G. hochhuthi, KYRGYZSTAN, Terskei-Alatau), 49—habitus of holotype, dorsal view, 50—labels of holotype. Scale bars: 1 mm (49), 0.1 mm (47, 48, 50)." httpUri="https://zenodo.org/record/3238886/files/figure.png" pageId="9" pageNumber="388">47</figureCitation>
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and
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<bibRefCitation id="70324B0CFFA00E7DC5C0FA23FC7EFA1F" author="Enushchenko, I. & Semenov, V." box="[662,978,1424,1446]" pageId="9" pageNumber="388" pagination="301 - 337" refId="ref11134" refString="Enushchenko, I. & Semenov, V. (2016) A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories. Zootaxa, 4126 (3), 301 - 337. https: // doi. org / 10.11646 / zootaxa. 4126.3.1" type="journal article" year="2016">Enushchenko & Semenov (2016</bibRefCitation>
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:
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<figureCitation id="8C982A78FFA00E7DC48CFA23FB86FA1C" box="[986,1066,1424,1446]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1937,1962]" captionTargetBox="[198,1372,195,1775]" captionTargetId="figure@4.[188,1399,181,1917]" captionTargetPageId="4" captionText="FIGURES 7–12. Encephalus angusticollis J. Sahlberg: 7 (type of E. torosus Eppelsheim, male), 10 (type of E. torosus Eppelsheim, female)—habitus, dorsal view, 11—habitus, frontal view (female), 8—aedeagus, lateral view, 9—labels of type of E. torosus Eppelsheim, male, 12—labels of type of E. torosus Eppelsheim, female. Scale bars: 1 mm (7, 10, 11), 0.1 mm (8)." httpUri="https://zenodo.org/record/3238874/files/figure.png" pageId="9" pageNumber="388">Figs. 12</figureCitation>
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,
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<figureCitation id="8C982A78FFA00E7DC362FA23FBE7FA1F" box="[1076,1099,1424,1446]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[237,1365,205,1920]" captionTargetId="figure@6.[194,1393,181,1938]" captionTargetPageId="6" captionText="FIGURES 13–16. Gyrophaena kangasi Rutanen (Holotype): 13—habitus, dorsal view, 14—aedeagus, lateral view, 15—male abdominal tergite VIII, dorsal view, 16—labels of the holotype. Scale bars: 1 mm (13), 0.1 mm (14, 15)." httpUri="https://zenodo.org/record/3238878/files/figure.png" pageId="9" pageNumber="388">13</figureCitation>
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))......................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="AF572E58FFA00E7DC7C1FA1DFA30F9EF" pageId="9" pageNumber="388">
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<paragraph id="141C36FDFFA00E7DC7C1FA1DFA30FA44" blockId="9.[151,1437,1219,1622]" pageId="9" pageNumber="388">
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<keyLead id="AF5295C8FFA00E7DC7C1FA1DFA30FA44" pageId="9" pageNumber="388">
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2 Elytra with indistinct, sparse, and small punctation. Median teeth of male abdominal tergite VIII rather thin and small (
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<figureCitation id="8C982A78FFA00E7DC23AFA1EFE8CFA59" captionStart="FIGURES 39–46" captionStartId="15.[151,264,1884,1909]" captionTargetBox="[224,1372,204,1780]" captionTargetId="figure@15.[205,1382,181,1864]" captionTargetPageId="15" captionText="FIGURES 39–46. Gyrophaena aryanamensis, sp. nov. (Figs. 39, 40: KYRGYZSTAN, Dzhany-Bazar) and Gyrophaena hochhuthi Bernhauer (Figs. 41–46: KYRGYZSTAN, Dzhany-Bazar): 41—habitus, dorsal view, 42—aedeagus, lateral view, 43, 44—male abdominal tergite VIII, dorsal view, 39, 45—female abdominal tergite VIII, dorsal view, 40, 46—female abdominal sternite VIII, dorsal view. Scale bars: 1 mm (41), 0.1 mm (39, 40, 42–46)." httpUri="https://zenodo.org/record/3238882/files/figure.png" pageId="9" pageNumber="388">Figs. 43–44</figureCitation>
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,
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<figureCitation id="8C982A78FFA00E7DC67DFA78FEEAFA58" box="[299,326,1483,1505]" captionStart="FIGURES 47–50" captionStartId="16.[151,264,1927,1952]" captionTargetBox="[186,1362,200,1802]" captionTargetId="figure@16.[186,1400,181,1907]" captionTargetPageId="16" captionText="FIGURES 47–50. Gyrophaena hochhuthi Bernhauer: 47—aedeagus, lateral view, 48—male abdominal tergite VIII, dorsal view (Figs. 47, 48: G. hochhuthi, KYRGYZSTAN, Terskei-Alatau), 49—habitus of holotype, dorsal view, 50—labels of holotype. Scale bars: 1 mm (49), 0.1 mm (47, 48, 50)." httpUri="https://zenodo.org/record/3238886/files/figure.png" pageId="9" pageNumber="388">48</figureCitation>
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). Lower edge of ventral process of aedeagus somewhat flat; dorsal projection of internal sac irregularly elongate, slightly widening to apex (
|
||
<figureCitation id="8C982A78FFA00E7DC6B4FA5BFD9FFA44" box="[482,563,1512,1534]" captionStart="FIGURES 39–46" captionStartId="15.[151,264,1884,1909]" captionTargetBox="[224,1372,204,1780]" captionTargetId="figure@15.[205,1382,181,1864]" captionTargetPageId="15" captionText="FIGURES 39–46. Gyrophaena aryanamensis, sp. nov. (Figs. 39, 40: KYRGYZSTAN, Dzhany-Bazar) and Gyrophaena hochhuthi Bernhauer (Figs. 41–46: KYRGYZSTAN, Dzhany-Bazar): 41—habitus, dorsal view, 42—aedeagus, lateral view, 43, 44—male abdominal tergite VIII, dorsal view, 39, 45—female abdominal tergite VIII, dorsal view, 40, 46—female abdominal sternite VIII, dorsal view. Scale bars: 1 mm (41), 0.1 mm (39, 40, 42–46)." httpUri="https://zenodo.org/record/3238882/files/figure.png" pageId="9" pageNumber="388">Figs. 42</figureCitation>
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,
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<figureCitation id="8C982A78FFA00E7DC56BFA5BFDFAFA47" box="[573,598,1512,1534]" captionStart="FIGURES 47–50" captionStartId="16.[151,264,1927,1952]" captionTargetBox="[186,1362,200,1802]" captionTargetId="figure@16.[186,1400,181,1907]" captionTargetPageId="16" captionText="FIGURES 47–50. Gyrophaena hochhuthi Bernhauer: 47—aedeagus, lateral view, 48—male abdominal tergite VIII, dorsal view (Figs. 47, 48: G. hochhuthi, KYRGYZSTAN, Terskei-Alatau), 49—habitus of holotype, dorsal view, 50—labels of holotype. Scale bars: 1 mm (49), 0.1 mm (47, 48, 50)." httpUri="https://zenodo.org/record/3238886/files/figure.png" pageId="9" pageNumber="388">47</figureCitation>
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)...........................................................
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||
<taxonomicName id="D3A34D7EFFA00E7DC249FA5AFA30FA44" ID-CoL="3HZZZ" authorityName="Bernhauer" authorityYear="1908" box="[1311,1436,1512,1534]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="388" phylum="Arthropoda" rank="species" species="hochhuthi">
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<emphasis id="26D7EAEFFFA00E7DC249FA5AFA30FA44" box="[1311,1436,1512,1534]" italics="true" pageId="9" pageNumber="388">G. hochhuthi</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="141C36FDFFA00E7DC7C1F9B5FA30F9EF" blockId="9.[151,1437,1219,1622]" pageId="9" pageNumber="388">
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<keyLead id="AF5295C8FFA00E7DC7C1F9B5FA30F9EF" pageId="9" pageNumber="388">
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- Punctation of elytra markedly dense. Median teeth of male tergite VIII stout, wide, and obtuse (
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<bibRefCitation id="70324B0CFFA00E7DC3CAF9B6FEBAF980" author="Enushchenko, I. & Semenov, V." pageId="9" pageNumber="388" pagination="301 - 337" refId="ref11134" refString="Enushchenko, I. & Semenov, V. (2016) A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories. Zootaxa, 4126 (3), 301 - 337. https: // doi. org / 10.11646 / zootaxa. 4126.3.1" type="journal article" year="2016">Enushchenko & Semenov 2016</bibRefCitation>
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||
:
|
||
<figureCitation id="8C982A78FFA00E7DC649F990FEDCF980" box="[287,368,1571,1593]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[237,1365,205,1920]" captionTargetId="figure@6.[194,1393,181,1938]" captionTargetPageId="6" captionText="FIGURES 13–16. Gyrophaena kangasi Rutanen (Holotype): 13—habitus, dorsal view, 14—aedeagus, lateral view, 15—male abdominal tergite VIII, dorsal view, 16—labels of the holotype. Scale bars: 1 mm (13), 0.1 mm (14, 15)." httpUri="https://zenodo.org/record/3238878/files/figure.png" pageId="9" pageNumber="388">Figs. 14</figureCitation>
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); lower edge of the ventral plate slightly angular in outline; dorsal projection of internal sac regularly elongate (
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<bibRefCitation id="70324B0CFFA00E7DC7BEF9F3FDB1F9EF" author="Enushchenko, I. & Semenov, V." box="[232,541,1600,1622]" pageId="9" pageNumber="388" pagination="301 - 337" refId="ref11134" refString="Enushchenko, I. & Semenov, V. (2016) A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories. Zootaxa, 4126 (3), 301 - 337. https: // doi. org / 10.11646 / zootaxa. 4126.3.1" type="journal article" year="2016">Enushchenko & Semenov 2016</bibRefCitation>
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:
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<figureCitation id="8C982A78FFA00E7DC570F9F3FDD9F9EC" box="[550,629,1600,1622]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1937,1962]" captionTargetBox="[198,1372,195,1775]" captionTargetId="figure@4.[188,1399,181,1917]" captionTargetPageId="4" captionText="FIGURES 7–12. Encephalus angusticollis J. Sahlberg: 7 (type of E. torosus Eppelsheim, male), 10 (type of E. torosus Eppelsheim, female)—habitus, dorsal view, 11—habitus, frontal view (female), 8—aedeagus, lateral view, 9—labels of type of E. torosus Eppelsheim, male, 12—labels of type of E. torosus Eppelsheim, female. Scale bars: 1 mm (7, 10, 11), 0.1 mm (8)." httpUri="https://zenodo.org/record/3238874/files/figure.png" pageId="9" pageNumber="388">Figs. 12</figureCitation>
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,
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<figureCitation id="8C982A78FFA00E7DC529F9F3FD3BF9EF" box="[639,663,1600,1622]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1958,1983]" captionTargetBox="[237,1365,205,1920]" captionTargetId="figure@6.[194,1393,181,1938]" captionTargetPageId="6" captionText="FIGURES 13–16. Gyrophaena kangasi Rutanen (Holotype): 13—habitus, dorsal view, 14—aedeagus, lateral view, 15—male abdominal tergite VIII, dorsal view, 16—labels of the holotype. Scale bars: 1 mm (13), 0.1 mm (14, 15)." httpUri="https://zenodo.org/record/3238878/files/figure.png" pageId="9" pageNumber="388">13</figureCitation>
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)......................................................
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<taxonomicName id="D3A34D7EFFA00E7DC279F9F2FA30F9EF" ID-CoL="3HZXT" authorityName="Erichson" authorityYear="1837" box="[1327,1436,1601,1622]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="388" phylum="Arthropoda" rank="species" species="congrua">
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<emphasis id="26D7EAEFFFA00E7DC279F9F2FA30F9EF" box="[1327,1436,1601,1622]" italics="true" pageId="9" pageNumber="388">G. congrua</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document> |