163 lines
21 KiB
XML
163 lines
21 KiB
XML
<document id="469710D2F448B8040C9EDC518F9A25D3" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629613271721" checkinUser="felipe" docAuthor="Paquin, Pierre & Dupérré, Nadine" docDate="2009" docId="03C55737FFCAFFE2FF50FBFAD022FEB3" docLanguage="en" docName="zt02002p067.pdf" docOrigin="Zootaxa 2002" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cicurina (Cicurella) maya Gertsch 1977" docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FFFC2F4FFFD6FFFFFFC7FFA4D12FFFC0" masterDocTitle="of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution" masterLastPageNumber="67" masterPageNumber="1" pageNumber="29" updateTime="1699083252512" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C9BD3A7D753E58F77E90ED299074E768">of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution</mods:title>
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<mods:name id="DECAE6E296CEE50160117C2F5EC0A6CA" type="personal">
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<mods:namePart id="4E98378A7A9004C44A7BEA87E43200E4">Paquin, Pierre</mods:namePart>
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<mods:affiliation id="21641ACB006114657F9B85409815FB0B">Cave and Endangered Invertebrate Research Laboratory, SWCA Environmental Consultants, 4407 Monterey Oaks Boulevard, Building 1, Suite 110, Austin, Texas, 78749, USA. E-mail: ppaquin @ swca. com & This is publication no. 15 of the Karst Biosciences and Environmental Geophysics Research Laboratories, SWCA Environmental Consultants</mods:affiliation>
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<mods:namePart id="0E80836C20B9E24625B9B7D1656936D8">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="252AFF3A3B41EE9E1B89BC613905CAD7">American Museum of Natural History, Division of Invertebrate Zoology, Central Park West at 79 Street, New York, New York, 10024 USA. E-mail: nduperre @ amnh. org</mods:affiliation>
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<mods:title id="8AA98675B1999CDE4AA46142E508A8E1">Zootaxa</mods:title>
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<mods:date id="A2E79E1F7D21F98B48826EA71C5207EF">2009</mods:date>
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<mods:number id="EE0CC52266BFB40F501F30F389443242">2009-02-04</mods:number>
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<mods:number id="E55CBEA4B59E3B683E1DF59DD4E6F0BC">2002</mods:number>
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<treatment id="03C55737FFCAFFE2FF50FBFAD022FEB3" ID-DOI="http://doi.org/10.5281/zenodo.5243037" ID-Zenodo-Dep="5243037" LSID="urn:lsid:plazi:treatment:03C55737FFCAFFE2FF50FBFAD022FEB3" httpUri="http://treatment.plazi.org/id/03C55737FFCAFFE2FF50FBFAD022FEB3" lastPageId="29" lastPageNumber="29" pageId="28" pageNumber="29">
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<subSubSection id="C376B5AAFFCAFFE3FF50FBFAD3ACFBB8" box="[151,643,1118,1144]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="8BD3E621FFCAFFE3FF50FBFAD3ACFBB8" blockId="28.[151,643,1118,1179]" box="[151,643,1118,1144]" pageId="28" pageNumber="29">
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<taxonomicName id="4C6C9DA2FFCAFFE3FF50FBFAD3ACFBB8" ID-CoL="5Z673" authority="Gertsch 1977" authorityName="Gertsch" authorityYear="1977" box="[151,643,1118,1144]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="maya" subGenus="Cicurella">
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<emphasis id="B9183A33FFCAFFE3FF50FBFAD3ACFBB8" bold="true" box="[151,643,1118,1144]" pageId="28" pageNumber="29">
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<emphasis id="B9183A33FFCAFFE3FF50FBFAD0F9FBB8" bold="true" box="[151,470,1118,1144]" italics="true" pageId="28" pageNumber="29">Cicurina (Cicurella) maya</emphasis>
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<bibRefCitation id="EFFD9BD0FFCAFFE3FE1AFBFAD3ACFBB8" author="Gertsch, W. J." box="[477,643,1118,1144]" pageId="28" pageNumber="29" pagination="103 - 131" refId="ref39983" refString="Gertsch, W. J. (1977) Report on cavernicole and epigean spiders from the Yucatan Peninsula. Association for Mexican Cave Studies, Bulletin, 6, 103 - 131." type="journal article" year="1977">Gertsch 1977</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFCAFFE3FF50FB25D047FB5B" box="[151,360,1153,1179]" pageId="28" pageNumber="29" type="description">
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<paragraph id="8BD3E621FFCAFFE3FF50FB25D047FB5B" blockId="28.[151,643,1118,1179]" box="[151,360,1153,1179]" pageId="28" pageNumber="29">
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(
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<figureCitation id="1357FAA4FFCAFFE3FF58FB25D009FB5B" box="[159,294,1153,1179]" captionStart="FIGURES 48–67" captionStartId="30.[151,269,1797,1821]" captionTargetBox="[272,1310,209,1785]" captionTargetId="figure-135@30.[272,1315,194,1788]" captionTargetPageId="30" captionText="FIGURES 48–67. Genitalic structure of troglobitic Cicurina (Cicurella) holotypes. For each female holotype, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. For the male holotype, a ventral and a lateral view of the left palp are presented, respectively. 48–49 Cicurina machete Gertsch 1992. 50–51 Cicurina madla Gertsch 1992. 52–53 Cicurina maya Gertsch 1977. 54–55" pageId="28" pageNumber="29">Figs 52–53</figureCitation>
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,
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<figureCitation id="1357FAA4FFCAFFE3FEF5FB25D070FB5B" box="[306,351,1153,1179]" captionStart="FIGURE 128" captionStartId="49.[151,256,1654,1678]" captionTargetBox="[162,1430,467,1637]" captionTargetId="figure-211@49.[158,1443,467,1654]" captionTargetPageId="49" captionText="FIGURE 128. Distribution of troglobitic Cicurina (subgenus Cicurella) species over the total range. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). See figures 129 to 138 for more detailed coverage of certain areas." pageId="28" pageNumber="29">128</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFCAFFE3FF50FB6AD0FDFAC5" pageId="28" pageNumber="29" type="reference_group">
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<paragraph id="8BD3E621FFCAFFE3FF50FB6AD0FDFAC5" blockId="28.[151,1436,1230,1285]" pageId="28" pageNumber="29">
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<treatmentCitationGroup id="AB7CC10FFFCAFFE3FF50FB6AD0FDFAC5" pageId="28" pageNumber="29">
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<treatmentCitation id="0ACDC030FFCAFFE3FF50FB6AD32CFB25" author="Gertsch, W. J." box="[151,515,1230,1253]" page="127" pageId="28" pageNumber="29" year="1977">
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<taxonomicName id="4C6C9DA2FFCAFFE3FF50FB6AD32CFB25" ID-CoL="5Z673" authority="Gertsch 1977: 127" authorityName="Gertsch" authorityPageNumber="127" authorityYear="1977" box="[151,515,1230,1253]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="maya">
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<emphasis id="B9183A33FFCAFFE3FF50FB6AD019FB25" box="[151,310,1230,1253]" italics="true" pageId="28" pageNumber="29">Cicurina maya</emphasis>
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<bibRefCitation id="EFFD9BD0FFCAFFE3FEFBFB6AD32CFB25" author="Gertsch, W. J." box="[316,515,1230,1253]" pageId="28" pageNumber="29" pagination="103 - 131" refId="ref39983" refString="Gertsch, W. J. (1977) Report on cavernicole and epigean spiders from the Yucatan Peninsula. Association for Mexican Cave Studies, Bulletin, 6, 103 - 131." type="journal article" year="1977">Gertsch 1977: 127</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 86 (description of the female).
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<emphasis id="B9183A33FFCAFFE3FC44FB6AD2B1FB26" bold="true" box="[899,926,1230,1254]" pageId="28" pageNumber="29">—</emphasis>
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<treatmentCitation id="0ACDC030FFCAFFE3FC63FB6AD544FB25" author="Reddell, J. R." box="[932,1131,1230,1253]" page="232" pageId="28" pageNumber="29" year="1977">
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<bibRefCitation id="EFFD9BD0FFCAFFE3FC63FB6AD544FB25" author="Reddell, J. R." box="[932,1131,1230,1253]" pageId="28" pageNumber="29" pagination="215 - 296" refId="ref40749" refString="Reddell, J. R. (1977) A preliminary survey of the caves of the Yucatan Peninsula. Bulletin of Association for Mexican Cave Studies, 6, 215 - 296." type="journal article" year="1977">Reddell 1977: 232</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0ACDC030FFCAFFE3FBB1FB6AD411FB25" author="Reddell, J. R." box="[1142,1342,1230,1253]" page="136" pageId="28" pageNumber="29" year="1981">
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||
<bibRefCitation id="EFFD9BD0FFCAFFE3FBB1FB6AD411FB25" author="Reddell, J. R." box="[1142,1342,1230,1253]" pageId="28" pageNumber="29" pagination="1 - 327" refId="ref40783" refString="Reddell, J. R. (1981) A review of the cavernicole fauna of Mexico, Guatemala, and Belize. Bulletin of the Texas Memorial Museum, 27, 1 - 327." type="journal article" year="1981">Reddell 1981: 136</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0ACDC030FFCAFFE3FA8EFB6AD01CFAC5" author="Gertsch, W. J." page="119" pageId="28" pageNumber="29" year="1992">
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<bibRefCitation id="EFFD9BD0FFCAFFE3FA8EFB6AD01CFAC5" author="Gertsch, W. J." pageId="28" pageNumber="29" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992: 119</bibRefCitation>
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</treatmentCitation>
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, figs 161–162.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFCAFFE3FF50FA91D023FAB6" pageId="28" pageNumber="29" type="materials_examined">
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||
<paragraph id="8BD3E621FFCAFFE3FF50FA91D023FAB6" blockId="28.[151,1437,1333,1939]" pageId="28" pageNumber="29">
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||
<materialsCitation id="3B04EC7CFFCAFFE3FF50FA91D023FAB6" collectingDate="1973-03-27" collectionCode="AMNH" collectorName="J. Reddell" country="Mexico" location="Actun Tucil" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Yucatan" typeStatus="holotype">
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<emphasis id="B9183A33FFCAFFE3FF50FA91D08DFA8F" bold="true" box="[151,418,1333,1359]" pageId="28" pageNumber="29">
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||
<typeStatus id="54D75883FFCAFFE3FF50FA91D012FA8F" box="[151,317,1333,1359]" pageId="28" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
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female:
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</emphasis>
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“
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<collectingCountry id="F37BA6B1FFCAFFE3FE72FA91D339FA8F" box="[437,534,1333,1359]" name="Mexico" pageId="28" pageNumber="29">Mexico</collectingCountry>
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||
:
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||
<collectingRegion id="49A828C3FFCAFFE3FDE6FA91D350FA8F" box="[545,639,1333,1359]" country="Mexico" name="Yucatan" pageId="28" pageNumber="29">Yucatan</collectingRegion>
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||
<location id="8EB3B0FAFFCAFFE3FD4FFA91D237FA8F" LSID="urn:lsid:plazi:treatment:03C55737FFCAFFE2FF50FBFAD022FEB3:8EB3B0FAFFCAFFE3FD4FFA91D237FA8F" box="[648,792,1333,1359]" country="Mexico" name="Actun Tucil" pageId="28" pageNumber="29" stateProvince="Yucatan">Actun Tucil</location>
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,
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<locationDeviation id="9A61FF03FFCAFFE3FCE3FA91D2E6FA8F" box="[804,969,1333,1359]" location="Muna" pageId="28" pageNumber="29">
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<quantity id="4C944BC4FFCAFFE3FCE3FA91D24FFA8F" box="[804,864,1333,1359]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="km" value="2.0">2 km</quantity>
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S Muna
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</locationDeviation>
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,
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<date id="FFD2C0E1FFCAFFE3FC12FA91D5A6FA8F" box="[981,1161,1333,1359]" pageId="28" pageNumber="29" value="1973-03-27">
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<collectingDate id="EF963909FFCAFFE3FC12FA91D5A6FA8F" box="[981,1161,1333,1359]" pageId="28" pageNumber="29" value="1973-03-27">27 March 1973</collectingDate>
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</date>
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<collectorName id="269983F7FFCAFFE3FB55FA91D420FA8F" box="[1170,1295,1333,1359]" pageId="28" pageNumber="29">J. Reddell</collectorName>
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”. Examined (
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<collectionCode id="ED7D7EE4FFCAFFE3FF58FAF8D1D0FAB6" box="[159,255,1372,1398]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="28" pageNumber="29">AMNH</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFCAFFE3FF00FA26D24BF886" pageId="28" pageNumber="29" type="description">
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||
<paragraph id="8BD3E621FFCAFFE3FF00FA26D305FA5C" blockId="28.[151,1437,1333,1939]" box="[199,554,1410,1436]" pageId="28" pageNumber="29">
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||
<emphasis id="B9183A33FFCAFFE3FF00FA26D070FA5C" bold="true" box="[199,351,1410,1436]" pageId="28" pageNumber="29">Description.</emphasis>
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<emphasis id="B9183A33FFCAFFE3FEA1FA27D305FA5C" box="[358,554,1411,1436]" italics="true" pageId="28" pageNumber="29">
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Female
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<typeStatus id="54D75883FFCAFFE3FE02FA27D305FA5C" box="[453,554,1411,1436]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
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||
</emphasis>
|
||
</paragraph>
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||
<paragraph id="8BD3E621FFCAFFE3FF00FA0DD24BF886" blockId="28.[151,1437,1333,1939]" pageId="28" pageNumber="29">
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||
Total length: 3.13; carapace length: 1.43; carapace width: 0.95; opisthosoma length: 1.70. Four eyes highly reduced to pale remnant spots. Cheliceral promargin right side with 3 teeth (2 large, 1 small), left side likewise; retromargin right side with 9 denticles, left side likewise. Leg I: total length: 6.64; femur: 1.43; patella: 0.45; tibia: 0.90; metatarsus: 0.75; tarsus: 0.49; leg IV: total length: 4.15; femur: 1.15; patella: 0.45; tibia: 0.95; metatarsus: 1.05; tarsus:0.55. Tarsal claw IV: 0.12. Epigynum:
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||
<quantity id="4C944BC4FFCAFFE3FBC9F9E0D5A5F99E" box="[1038,1162,1604,1630]" metricMagnitude="-4" metricUnit="m" metricValue="2.94" pageId="28" pageNumber="29" unit="mm" value="0.294">0.294 mm</quantity>
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. Epigynal ventral plate with short transverse slit, indentation absent; atrium rectangular, half intruded by slightly rounded dorsal plate, atrial aperture moderate; bursa enlarged, not fused; copulatory ducts separated at midpoint, reaching top of primary lobe apex; copulatory ducts constriction located basally; primary pores not visible; stalk, short, procurved, joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore rounded; primary lobe hat-shaped; secondary lobe absent; fertilization canal long, internally positioned from the dictynoid pore; fertilization duct internally attached to the primary lobe base.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFCAFFE3FF00F8F6D5DFF8AC" box="[199,1264,1874,1900]" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8BD3E621FFCAFFE3FF00F8F6D5DFF8AC" blockId="28.[151,1437,1333,1939]" box="[199,1264,1874,1900]" pageId="28" pageNumber="29">
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||
<emphasis id="B9183A33FFCAFFE3FF00F8F6D04FF8AC" bold="true" box="[199,352,1874,1900]" pageId="28" pageNumber="29">Distribution</emphasis>
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||
. Only known from the
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||
<typeStatus id="54D75883FFCAFFE3FDB4F8F6D38AF8AC" box="[627,677,1874,1900]" pageId="28" pageNumber="29">type</typeStatus>
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||
locality Actun Tucil,
|
||
<collectingRegion id="49A828C3FFCAFFE3FC61F8F6D528F8AC" box="[934,1031,1874,1900]" country="Mexico" name="Yucatan" pageId="28" pageNumber="29">Yucatan</collectingRegion>
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||
,
|
||
<collectingCountry id="F37BA6B1FFCAFFE3FBD4F8F6D543F8AC" box="[1043,1132,1874,1900]" name="Mexico" pageId="28" pageNumber="29">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="1357FAA4FFCAFFE3FBBAF8F6D5CCF8AC" box="[1149,1251,1874,1900]" captionStart="FIGURE 128" captionStartId="49.[151,256,1654,1678]" captionTargetBox="[162,1430,467,1637]" captionTargetId="figure-211@49.[158,1443,467,1654]" captionTargetPageId="49" captionText="FIGURE 128. Distribution of troglobitic Cicurina (subgenus Cicurella) species over the total range. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). See figures 129 to 138 for more detailed coverage of certain areas." pageId="28" pageNumber="29">Fig. 128</figureCitation>
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||
).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376B5AAFFCAFFE2FF00F8DCD022FEB3" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8BD3E621FFCAFFE2FF00F8DCD022FEB3" blockId="28.[151,1437,1333,1939]" lastBlockId="29.[151,1436,152,371]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
<emphasis id="B9183A33FFCAFFE3FF00F8DCD06FF852" bold="true" box="[199,320,1912,1938]" pageId="28" pageNumber="29">Remarks.</emphasis>
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Male unknown. Gertsch mentioned that this species is an eyeless troglobite, but the female known specimen of the species. However, it is not clear that figure 161 is a ventral view as the spermathecal stalks are not illustrated (compare to
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<figureCitation id="1357FAA4FFCBFFE2FD8FFF1AD3B1FF18" box="[584,670,190,216]" captionStart="FIGURES 48–67" captionStartId="30.[151,269,1797,1821]" captionTargetBox="[272,1310,209,1785]" captionTargetId="figure-135@30.[272,1315,194,1788]" captionTargetPageId="30" captionText="FIGURES 48–67. Genitalic structure of troglobitic Cicurina (Cicurella) holotypes. For each female holotype, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. For the male holotype, a ventral and a lateral view of the left palp are presented, respectively. 48–49 Cicurina machete Gertsch 1992. 50–51 Cicurina madla Gertsch 1992. 52–53 Cicurina maya Gertsch 1977. 54–55" pageId="29" pageNumber="30">Fig. 52</figureCitation>
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, present paper), and looks as the figure could be of a dorsal view. However, the obvious atrium and the ventral position of the connecting ducts in relation to the spermathecae, suggest that this is simply an inaccurate illustration, likely inspired from
|
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<bibRefCitation id="EFFD9BD0FFCBFFE2FC21FEA8D5B5FEE6" author="Gertsch, W. J." box="[998,1178,268,294]" pageId="29" pageNumber="30" pagination="103 - 131" refId="ref39983" refString="Gertsch, W. J. (1977) Report on cavernicole and epigean spiders from the Yucatan Peninsula. Association for Mexican Cave Studies, Bulletin, 6, 103 - 131." type="journal article" year="1977">Gertsch (1977)</bibRefCitation>
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: fig. 86 of the original description.
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<bibRefCitation id="EFFD9BD0FFCBFFE2FEEEFE96D0F0FE8C" author="Gertsch, W. J." box="[297,479,306,332]" pageId="29" pageNumber="30" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch (1992)</bibRefCitation>
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: fig. 162 matches
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<figureCitation id="1357FAA4FFCBFFE2FD7DFE96D221FE8C" box="[698,782,306,332]" captionStart="FIGURES 48–67" captionStartId="30.[151,269,1797,1821]" captionTargetBox="[272,1310,209,1785]" captionTargetId="figure-135@30.[272,1315,194,1788]" captionTargetPageId="30" captionText="FIGURES 48–67. Genitalic structure of troglobitic Cicurina (Cicurella) holotypes. For each female holotype, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. For the male holotype, a ventral and a lateral view of the left palp are presented, respectively. 48–49 Cicurina machete Gertsch 1992. 50–51 Cicurina madla Gertsch 1992. 52–53 Cicurina maya Gertsch 1977. 54–55" pageId="29" pageNumber="30">Fig. 53</figureCitation>
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(present paper), and allows the recognition of the
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||
<typeStatus id="54D75883FFCBFFE2FAAEFE96D4B4FE8C" box="[1385,1435,306,332]" pageId="29" pageNumber="30">type</typeStatus>
|
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specimen.
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||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |