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<document id="B34E4B3EDEB5BBBD35A27AAF18DBFF44" ID-CLB-Dataset="264199" ID-DOI="10.11646/zootaxa.5332.1.1" ID-GBIF-Dataset="66e0ec51-d494-43d4-965e-a2cd1462ef54" ID-ISSN="1175-5326" ID-Zenodo-Dep="8261502" ID-ZooBank="424F7439-4095-46A5-93E3-C4130E3B6D9A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692350092877" checkinUser="plazi" docAuthor="POORANI, J." docDate="2023" docId="03C74162141E4670BDDF32C4FBD8FEDA" docLanguage="en" docName="zootaxa.5332.1.1.pdf" docOrigin="Zootaxa 5332 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5332.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cheilomenes Chevrolat" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="FFFE391A14584637BD48367CFFCDFF9B" masterDocTitle="An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini" masterLastPageNumber="307" masterPageNumber="1" pageNumber="71" updateTime="1699593964221" updateUser="ExternalLinkService">
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<mods:title id="01890F37D4B640B84A30543911240DB4">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<treatment id="03C74162141E4670BDDF32C4FBD8FEDA" ID-DOI="http://doi.org/10.5281/zenodo.8273729" ID-GBIF-Taxon="212450606" ID-Zenodo-Dep="8273729" LSID="urn:lsid:plazi:treatment:03C74162141E4670BDDF32C4FBD8FEDA" httpUri="http://treatment.plazi.org/id/03C74162141E4670BDDF32C4FBD8FEDA" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<subSubSection id="C374A3FF141E4671BDDF32C4FDC2FB49" box="[151,527,1208,1235]" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="8BD1F074141E4671BDDF32C4FDC2FB49" blockId="70.[151,527,1208,1235]" box="[151,527,1208,1235]" pageId="70" pageNumber="71">
<emphasis id="B91A2C66141E4671BDDF32C4FDC2FB49" bold="true" box="[151,527,1208,1235]" pageId="70" pageNumber="71">
Genus
<taxonomicName id="4C6E8BF7141E4671BDA632C4FDC2FB49" ID-CoL="3MHC" ID-ENA="158621" authority="Chevrolat" authorityName="Chevrolat" box="[238,527,1208,1235]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BDA632C4FE47FB48" bold="true" box="[238,394,1208,1235]" italics="true" pageId="70" pageNumber="71">Cheilomenes</emphasis>
Chevrolat
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4671BDDF3281FDB8FA8E" box="[151,629,1277,1301]" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="8BD1F074141E4671BDDF3281FDB8FA8E" blockId="70.[151,1437,1277,1393]" box="[151,629,1277,1301]" pageId="70" pageNumber="71">
<taxonomicName id="4C6E8BF7141E4671BDDF3281FE4AFA8E" ID-CoL="3MHC" authority="Chevrolat" authorityName="Chevrolat" box="[151,391,1277,1301]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BDDF3281FED7FA8E" box="[151,282,1277,1301]" italics="true" pageId="70" pageNumber="71">Cheilomenes</emphasis>
Chevrolat
</taxonomicName>
in Dejean, 1836: 435.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4671BF353281FF39FAAF" pageId="70" pageNumber="71" type="type_taxon">
<paragraph id="8BD1F074141E4671BF353281FF39FAAF" blockId="70.[151,1437,1277,1393]" pageId="70" pageNumber="71">
<typeStatus id="54D54ED6141E4671BF353281FD62FA8E" box="[637,687,1277,1301]" pageId="70" pageNumber="71">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF7141E4671BE5A3281FBA2FA8E" authority="Fabricius 1775" authorityName="Fabricius" authorityYear="1775" box="[786,1135,1277,1301]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="lunata">
<emphasis id="B91A2C66141E4671BE5A3281FC07FA8E" box="[786,970,1277,1301]" italics="true" pageId="70" pageNumber="71">Coccinella lunata</emphasis>
Fabricius 1775
</taxonomicName>
, designated by Crotch 1874: 179.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4671BDDF3346FB8CFAEA" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="8BD1F074141E4671BDDF3346FD1DFAC9" blockId="70.[151,1437,1277,1393]" box="[151,720,1338,1362]" pageId="70" pageNumber="71">
<taxonomicName id="4C6E8BF7141E4671BDDF3346FE1CFAC9" authority=": Crotch 1874: 179" authorityName=": Crotch" authorityPageNumber="179" authorityYear="1874" box="[151,465,1338,1362]" class="Insecta" family="Coccinellidae" genus="Chilomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BDDF3346FEC2FAC9" box="[151,271,1338,1362]" italics="true" pageId="70" pageNumber="71">Chilomenes</emphasis>
: Crotch 1874: 179
</taxonomicName>
. Unjustified emendation.
</paragraph>
<paragraph id="8BD1F074141E4671BDDF3325FB8CFAEA" blockId="70.[151,1437,1277,1393]" box="[151,1089,1369,1393]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="AB7ED75A141E4671BDDF3325FB8CFAEA" box="[151,1089,1369,1393]" pageId="70" pageNumber="71">
<treatmentCitation id="0ACFD665141E4671BDDF3325FE3DFAEA" author="Timberlake, P. H." box="[151,496,1369,1393]" page="40" pageId="70" pageNumber="71" year="1943">
<taxonomicName id="4C6E8BF7141E4671BDDF3325FE3DFAEA" authority="Timberlake, 1943: 40" authorityName="Timberlake" authorityPageNumber="40" authorityYear="1943" box="[151,496,1369,1393]" class="Insecta" family="Coccinellidae" genus="Menochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BDDF3325FEC0FAEA" box="[151,269,1369,1393]" italics="true" pageId="70" pageNumber="71">Menochilus</emphasis>
<bibRefCitation id="EFFF8D85141E4671BC5C3325FE3DFAEA" author="Timberlake, P. H." box="[276,496,1369,1393]" pageId="70" pageNumber="71" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake, 1943: 40</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Replacement name for
<taxonomicName id="4C6E8BF7141E4671BFAE3325FBF0FAEA" ID-CoL="3MHC" authority="sensu Mulsant 1850" authorityName="sensu Mulsant" authorityYear="1850" box="[742,1085,1369,1393]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BFAE3325FCA4FAEA" box="[742,873,1369,1393]" italics="true" pageId="70" pageNumber="71">Cheilomenes</emphasis>
sensu
<bibRefCitation id="EFFF8D85141E4671BEE63325FBF0FAEA" author="Mulsant, E." box="[942,1085,1369,1393]" pageId="70" pageNumber="71" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant 1850</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4671BDDF33E3FCC3F8BA" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="8BD1F074141E4671BDDF33E3FCC3F8BA" blockId="70.[151,1437,1439,2042]" pageId="70" pageNumber="71">
<emphasis id="B91A2C66141E4671BDDF33E3FEDFFA22" bold="true" box="[151,274,1439,1465]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
Form broad oval to slightly elongate. Anterior clypeal margin semicircularly arcuate between lateral projections (
<figureCitation id="1355ECF1141E4671BC6A33BFFE4EFA46" box="[290,387,1475,1501]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46a</figureCitation>
). Antenna (
<figureCitation id="1355ECF1141E4671BF4933BFFDADFA46" box="[513,608,1475,1501]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46b</figureCitation>
) 11-segmented, shorter than interocular distance, scape broader than long, with inner side strongly rounded and produced and outwardly carinate. Inner ocular margins straight. Terminal maxillary palpomere securiform (
<figureCitation id="1355ECF1141E4671BF503077FDB1F9BE" box="[536,636,1547,1573]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46c</figureCitation>
). Pronotum with anterior margin trapezoidally excavated and slightly convex, anterior angles narrowly rounded; sides gently arched with lateral borders narrowly upturned. Elytra narrowly expanded on the sides, not marginated. Prothoracic hypomeron lacking a fovea, prosternal process with a pair of short, abbreviated carinae reaching up to middle of prosternum (
<figureCitation id="1355ECF1141E4671BE88300BFBEEF90A" box="[960,1059,1655,1681]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46d</figureCitation>
); anterior margin of mesoventrite weakly emarginate. Elytral epipleura very slightly foveate to receive meso- and metatibial apices. Abdominal postcoxal line incomplete, with a short, oblique lateral line (
<figureCitation id="1355ECF1141E4671BE0C30C3FC6BF942" box="[836,934,1727,1753]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46e</figureCitation>
). Meso- and metatibial apices with a pair of spurs. Male genitalia (
<figureCitation id="1355ECF1141E4671BCDE309FFDC4F966" box="[406,521,1763,1789]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46fj</figureCitation>
) with penis apex greatly attenuated, long and thread-like. Female genitalia (
<figureCitation id="1355ECF1141E4671B827309FFF0EF8BA" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46k</figureCitation>
) with sperm duct very long and coiled (
<figureCitation id="1355ECF1141E4671BFCA317BFCCFF8BA" box="[642,770,1799,1825]" captionStart="FIGURE 46" captionStartId="72.[151,250,1943,1967]" captionTargetBox="[211,1377,187,1914]" captionTargetId="figure-20@72.[203,1385,181,1920]" captionTargetPageId="72" captionText="FIGURE 46. Cheilomenes sexmaculata (F.): a. head; b. antenna; c. maxilla; d. prosternal process; e. abdominal postcoxal line; fj. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h, i. penis, variants; j. penis apex; k. female genitalia; l, m. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261648" httpUri="https://zenodo.org/record/8261648/files/figure.png" pageId="70" pageNumber="71">Fig. 46l, m</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4671BD8F3157FED2F8F2" pageId="70" pageNumber="71" type="distribution">
<paragraph id="8BD1F074141E4671BD8F3157FED2F8F2" blockId="70.[151,1437,1439,2042]" pageId="70" pageNumber="71">
<emphasis id="B91A2C66141E4671BD8F3157FEACF8DE" bold="true" box="[199,353,1835,1861]" pageId="70" pageNumber="71">Distribution.</emphasis>
This genus is widely distributed in the
<collectingRegion id="49AA3E96141E4671BE533157FCB7F8DE" box="[795,890,1835,1861]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="70" pageNumber="71">Oriental</collectingRegion>
, Australasian and Afrotropical regions (
<bibRefCitation id="EFFF8D85141E4671B87F3151FEDEF8F2" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." pageId="70" pageNumber="71" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C66141E4671BDDF312CFF05F8F2" box="[151,200,1871,1897]" italics="true" pageId="70" pageNumber="71">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141E4670BD8F310FFE21FF62" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8BD1F074141E4671BD8F310FFEE5F862" blockId="70.[151,1437,1439,2042]" pageId="70" pageNumber="71">
<emphasis id="B91A2C66141E4671BD8F310FFEF0F816" bold="true" box="[199,317,1907,1933]" pageId="70" pageNumber="71">Affinities.</emphasis>
<bibRefCitation id="EFFF8D85141E4671BC0A310FFDA1F816" author="Tomaszewska, W. &amp; Escalona, H. E. &amp; Hartley, D. &amp; Li, J. &amp; Wang, X. &amp; Li, H. S. &amp; Pang, H. &amp; Slipinski, A. &amp; Zwick, A." box="[322,620,1907,1934]" pageId="70" pageNumber="71" pagination="611 - 631" refId="ref123461" refString="Tomaszewska, W., Escalona, H. E., Hartley, D., Li, J., Wang, X., Li, H. S., Pang, H., Slipinski, A. &amp; Zwick, A. (2021) Phylogeny of true ladybird beetles (Coccinellidae: Coccinellini) reveals pervasive convergent evolution and a rapid Cenozoic radiation. Systematic Entomology, 46, 611 - 631. https: // doi. org / 10.1111 / syen. 12479" type="journal article" year="2021">
Tomaszewska
<emphasis id="B91A2C66141E4671BCAD3108FDD8F816" box="[485,533,1907,1933]" italics="true" pageId="70" pageNumber="71">et al</emphasis>
. (2021)
</bibRefCitation>
included it in the
<emphasis id="B91A2C66141E4671BE7B310FFC7BF816" box="[819,950,1907,1933]" italics="true" pageId="70" pageNumber="71">Coccinella-</emphasis>
group of genera based on their phylogenetic analyses and recovered it as a distinct clade including
<taxonomicName id="4C6E8BF7141E4671BFA631EBFC44F82A" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[750,905,1943,1969]" class="Insecta" family="Coccinellidae" genus="Elpis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BFA631EBFCE5F82A" box="[750,808,1943,1969]" italics="true" pageId="70" pageNumber="71">Elpis</emphasis>
Mulsant
</taxonomicName>
and
<taxonomicName id="4C6E8BF7141E4671BEF631EBFB03F82A" authority="Timberlake." authorityName="Timberlake." box="[958,1230,1943,1969]" class="Insecta" family="Coccinellidae" genus="Menochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BEF631EBFB8CF82A" box="[958,1089,1943,1969]" italics="true" pageId="70" pageNumber="71">Menochilus</emphasis>
Timberlake.
</taxonomicName>
The generic limits of
<taxonomicName id="4C6E8BF7141E4671BDFE31C7FE8AF84E" authorityName="sensu Mulsant" authorityYear="1850" box="[182,327,1979,2005]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BDFE31C7FE8AF84E" box="[182,327,1979,2005]" italics="true" pageId="70" pageNumber="71">Cheilomenes</emphasis>
</taxonomicName>
and its related genera,
<taxonomicName id="4C6E8BF7141E4671BF0531C7FD4AF84E" authorityName="Mulsant" authorityYear="1850" box="[589,647,1979,2005]" class="Insecta" family="Coccinellidae" genus="Elpis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141E4671BF0531C7FD4AF84E" box="[589,647,1979,2005]" italics="true" pageId="70" pageNumber="71">Elpis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E8BF7141E4671BFF631C7FC4EF84E" authority="Mulsant" authorityName="Mulsant" box="[702,899,1979,2005]" class="Magnoliopsida" family="Rosaceae" genus="Cydonia" kingdom="Plantae" order="Rosales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="B91A2C66141E4671BFF631C7FCD0F84E" box="[702,797,1979,2005]" italics="true" pageId="70" pageNumber="71">Cydonia</emphasis>
Mulsant
</taxonomicName>
, were discussed by Fürsch (1989) and
<bibRefCitation id="EFFF8D85141E4671B87131C1FEE9F862" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." pageId="70" pageNumber="71" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C66141E4671BDDF319CFF05F862" box="[151,200,2015,2041]" italics="true" pageId="70" pageNumber="71">et al</emphasis>
. (2020)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BD1F074141F4670BD8F36E4FE21FF62" blockId="71.[151,1437,151,321]" pageId="71" pageNumber="72">
<bibRefCitation id="EFFF8D85141F4670BD8F36E4FEB0FF2A" author="Slipinski, A." box="[199,381,151,179]" pageId="71" pageNumber="72" refId="ref122747" refString="Slipinski, A. (2007) Australian ladybird beetles (Coleoptera: Coccinellidae). Their biology and classification. Australian Biological Resources Study, Canberra, xviii + 286 pp." type="book" year="2007">Ślipiński (2007)</bibRefCitation>
used the name
<taxonomicName id="4C6E8BF7141F4670BF6536EBFD7DFF2A" box="[557,688,151,177]" class="Insecta" family="Coccinellidae" genus="Menochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141F4670BF6536EBFD7DFF2A" box="[557,688,151,177]" italics="true" pageId="71" pageNumber="72">Menochilus</emphasis>
</taxonomicName>
for this genus in his review of Australian genera of
<taxonomicName id="4C6E8BF7141F4670B9B336EBFA55FF2A" authorityName="Latreille" authorityYear="1807" box="[1275,1432,151,177]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="family">Coccinellidae</taxonomicName>
, but
<bibRefCitation id="EFFF8D85141F4670BD8936C0FE79FF4E" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[193,436,187,215]" pageId="71" pageNumber="72" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C66141F4670BC6336C7FE91FF4E" box="[299,348,187,213]" italics="true" pageId="71" pageNumber="72">et al</emphasis>
. (2020)
</bibRefCitation>
used the name
<taxonomicName id="4C6E8BF7141F4670BF2836C7FD3CFF4E" authorityName="sensu Mulsant" authorityYear="1850" box="[608,753,187,213]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66141F4670BF2836C7FD3CFF4E" box="[608,753,187,213]" italics="true" pageId="71" pageNumber="72">Cheilomenes</emphasis>
</taxonomicName>
in their conspectus of
<taxonomicName id="4C6E8BF7141F4670BEA336C7FBB9FF4E" baseAuthorityName="Booth" baseAuthorityYear="1996" box="[1003,1140,187,213]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="tribe" tribe="Coccinellini">Coccinellini</taxonomicName>
of Australo-Pacific region and the same is followed here.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF141F4670BD8F377FFBD8FEDA" pageId="71" pageNumber="72" type="taxon_list">
<paragraph id="8BD1F074141F4670BD8F377FFBD8FEDA" blockId="71.[151,1437,151,321]" pageId="71" pageNumber="72">
<emphasis id="B91A2C66141F4670BD8F377FFE5FFE87" bold="true" box="[199,402,258,285]" pageId="71" pageNumber="72">Included species.</emphasis>
In South Asia, this genus is represented by
<taxonomicName id="4C6E8BF7141F4670BECB377FFAE0FE86" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[899,1325,259,285]" class="Insecta" family="Coccinellidae" genus="Cheilomenes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B91A2C66141F4670BECB377FFB6AFE86" box="[899,1191,259,285]" italics="true" pageId="71" pageNumber="72">Cheilomenes sexmaculata</emphasis>
(Fabricius)
</taxonomicName>
, probably the most widespread and well-studied species of the
<collectingRegion id="49AA3E96141F4670BFAD375BFC8CFEDA" box="[741,833,295,321]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="71" pageNumber="72">Oriental</collectingRegion>
and Indian region.
</paragraph>
</subSubSection>
</treatment>
</document>