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<document id="77FB0267A7F9EB66880160EF4E69211D" ID-DOI="10.5281/zenodo.197652" ID-GBIF-Dataset="46ecee1b-0146-4ad5-9579-6a2f245c6873" ID-ISSN="1175-5326" ID-Zenodo-Dep="197652" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460275562824" checkinUser="plazi" docAuthor="Dow, Rory A., Choong, Chee Yen &amp; Ng, Yong Foo" docDate="2010" docId="9A1C8796C735EF773F96FE4FFD66F4D3" docLanguage="en" docName="zt02605p055.pdf" docOrigin="Zootaxa 2605" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Amphicnemis hoisen Dow, Choong &amp; Ng, 2010, sp. nov." docType="treatment" docVersion="14" lastPageNumber="54" masterDocId="6625FFEEC733EF7E3F01F874FFA9F019" masterDocTitle="A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae)" masterLastPageNumber="55" masterPageNumber="45" pageNumber="51" updateTime="1698248054178" updateUser="plazi">
<mods:mods id="403D87A970517EC38331E43291CF0956" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="687251B919ABB5E24642F74006E35DE0">
<mods:title id="BDB92542201CF90B8996179078ABA640">A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="655566588CF05796B26442AEDE86F830" type="personal">
<mods:role id="20913BB089C459D086B9642A06567D9C">
<mods:roleTerm id="82C026231ECC5912F14685E996EBEC03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F76FB16D18D5D92528E9673BEDAE1E1D">Dow, Rory A.</mods:namePart>
</mods:name>
<mods:name id="5321725E09DB8C8901DE6234CF1931A2" type="personal">
<mods:role id="DA23FF1F0A757F8E33D742EB88C74121">
<mods:roleTerm id="C267CE59BD6CC060CD76B8AC73241032">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="366AF2481A38BC5DE2DC552CB807C67F">Choong, Chee Yen</mods:namePart>
</mods:name>
<mods:name id="C891751EAE3A7567C256A251E2C4246F" type="personal">
<mods:role id="178AB43257F3A4049F4CB3290578726D">
<mods:roleTerm id="1D9177E1FA8ED579B104B9102EA3CA0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="45BAAE264B77974037DF923081513E94">Ng, Yong Foo</mods:namePart>
</mods:name>
<mods:typeOfResource id="EB829E3CA6DBCB77B918FAD1815FD8B1">text</mods:typeOfResource>
<mods:relatedItem id="5A6523E6CB49542E063307260CAE4F45" type="host">
<mods:titleInfo id="BC31012277D05DB352A4CF8F093BE5A6">
<mods:title id="A6429D662B4FB242AF59C57D077FDDCB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1F0099A6AEDAABB5C12FFB6DF7887B19">
<mods:date id="063678D0A8678A206C7FC6AA96ABA8ED">2010</mods:date>
<mods:detail id="C4163C762E1B306C7D4685CF6AC4F60D" type="volume">
<mods:number id="04153DC735FB8A6008A50327F765BC8E">2605</mods:number>
</mods:detail>
<mods:extent id="09E5BB27C8D329BA61A0DE7E5866FBEF" unit="page">
<mods:start id="E0358AF01289FA2C0070A5F294B7A16B">45</mods:start>
<mods:end id="3A09243CC545466EA8D2C9CBCB67B8C6">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="60CCB6BDC818AAC6710A6391D441460C">journal article</mods:classification>
<mods:identifier id="F2705AF8D506FB9A08A2DCB30C765715" type="DOI">10.5281/zenodo.197652</mods:identifier>
<mods:identifier id="84327C7F7DA798FBC314DD9FCF2A31C7" type="GBIF-Dataset">46ecee1b-0146-4ad5-9579-6a2f245c6873</mods:identifier>
<mods:identifier id="77E67FEC08C3225077DE920E88E76253" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="72E34EBBBE90FF6A3EB63BDA6C47D2F8" type="Zenodo-Dep">197652</mods:identifier>
</mods:mods>
<treatment id="9A1C8796C735EF773F96FE4FFD66F4D3" ID-DOI="http://doi.org/10.5281/zenodo.5196158" ID-GBIF-Taxon="119395936" ID-Zenodo-Dep="5196158" LSID="urn:lsid:plazi:treatment:9A1C8796C735EF773F96FE4FFD66F4D3" httpUri="http://treatment.plazi.org/id/9A1C8796C735EF773F96FE4FFD66F4D3" lastPageId="9" lastPageNumber="54" pageId="6" pageNumber="51">
<subSubSection id="5AAF650BC735EF783F96FE4FFDB6F66E" pageId="6" pageNumber="51" type="nomenclature">
<paragraph id="120A3680C735EF783F96FE4FFE50F64D" blockId="6.[151,543,1594,1655]" box="[151,505,1594,1621]" pageId="6" pageNumber="51">
<heading id="494281ECC735EF783F96FE4FFE50F64D" bold="true" box="[151,505,1594,1621]" fontSize="11" level="1" pageId="6" pageNumber="51" reason="1">
<emphasis id="20C1EA92C735EF783F96FE4FFE50F64D" bold="true" box="[151,505,1594,1621]" pageId="6" pageNumber="51">
<taxonomicName id="D5B54D03C735EF783F96FE4FFE3DF64C" ID-CoL="CZSW" box="[151,404,1595,1621]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="hoisen" status="sp. nov.">
<emphasis id="20C1EA92C735EF783F96FE4FFE3DF64C" bold="true" box="[151,404,1595,1621]" italics="true" pageId="6" pageNumber="51">Amphicnemis hoisen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BF257E9C735EF783E9AFE4EFE50F64D" box="[411,505,1594,1620]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="120A3680C735EF783F96FE29FDB6F66E" blockId="6.[151,543,1594,1655]" box="[151,543,1629,1655]" pageId="6" pageNumber="51">
(
<figureCitation id="8A8E2A05C735EF783F9EFE29FF5AF66E" box="[159,243,1629,1655]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,279,1620,1644]" captionTargetBox="[151,1436,983,1595]" captionTargetId="figure@4.[151,1436,983,1595]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Female prothorax, dorsal view: (1) A. ecornuta (Sarawak); (2) A. hoisen paratype female (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197653/files/figure.png" pageId="6" pageNumber="51">Figs. 2</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783FFEFE29FEA6F66E" box="[255,271,1629,1655]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,273,798,822]" captionTargetBox="[177,1412,430,759]" captionTargetId="figure@5.[169,1418,421,773]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Female posterior pronotal lobe, lateral view: (3) A. bebar paratype female (PAH 09 _ COE 1); (4) A. ecornuta (Sarawak); (5) A. gracilis (Sg. Bebar); (6) A. hoisen paratype female (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197654/files/figure.png" pageId="6" pageNumber="51">6</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783E1DFE29FEC1F66E" box="[284,360,1629,1655]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,272,898,922]" captionTargetBox="[152,1434,449,862]" captionTargetId="figure@6.[151,1436,421,877]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pterostigma: (9) A. bebar holotype male Fw; (10) A. ecornuta male Fw (Sarawak); (11) A. gracilis male Fw (Sg. Bebar); (12) A. hoisen holotype male Fw; (13) A. hoisen holotype male Hw; (14) A. hoisen paratype female Fw (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197656/files/figure.png" pageId="6" pageNumber="51">12–14</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783E75FE29FE3AF66E" box="[372,403,1629,1655]" captionStart="FIGURES 15 – 17" captionStartId="6.[151,269,1490,1514]" captionTargetBox="[169,1419,1046,1453]" captionTargetId="figure@6.[158,1428,1036,1466]" captionTargetPageId="6" captionText="FIGURES 15 – 17. Genital valves: (15) A. bebar holotype; (16) A. ecornuta (Sarawak); (17) A. hoisen holotype." httpUri="https://zenodo.org/record/197657/files/figure.png" pageId="6" pageNumber="51">17</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783EA1FE29FE69F66E" box="[416,448,1629,1655]" captionStart="FIGURES 18 – 21" captionStartId="7.[151,271,1160,1184]" captionTargetBox="[154,1305,436,1125]" captionTargetId="figure@7.[151,1436,383,1139]" captionTargetPageId="7" captionText="FIGURES 18 – 21. Male caudal appendages, lateral view: (18) A. bebar holotype; (19) A. ecornuta (Sarawak); (20) A. gracilis (Sg. Bebar); (21) A. hoisen holotype." httpUri="https://zenodo.org/record/197658/files/figure.png" pageId="6" pageNumber="51">21</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783ECDFE29FE45F66E" box="[460,492,1629,1655]" captionStart="FIGURES 22 – 25" captionStartId="8.[151,271,1323,1347]" captionTargetBox="[197,1392,314,1271]" captionTargetId="figure@8.[187,1400,306,1302]" captionTargetPageId="8" captionText="FIGURES 22 – 25. Male cerci, dorsal views: (22) A. bebar holotype; (23) A. ecornuta (Sarawak); (24) A. gracilis (Sg. Bebar); (25) A. hoisen holotype." httpUri="https://zenodo.org/record/197659/files/figure.png" pageId="6" pageNumber="51">25</figureCitation>
,
<figureCitation id="8A8E2A05C735EF783EF9FE29FDBEF66E" box="[504,535,1629,1655]" captionStart="FIGURE 26. A" captionStartId="8.[151,255,1980,2004]" captionTargetBox="[506,1087,1464,1931]" captionTargetId="figure@8.[491,1096,1430,1959]" captionTargetPageId="8" captionText="FIGURE 26. A. hoisen holotype, ventral view lower branch cercus, right side. A part of the right paraproct is visible at the base of the cercus." httpUri="https://zenodo.org/record/197660/files/figure.png" pageId="6" pageNumber="51">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5AAF650BC735EF783F96FEDCFBB7F72B" pageId="6" pageNumber="51" type="materials_examined">
<paragraph id="120A3680C735EF783F96FEDCFBB7F72B" blockId="6.[151,1437,1704,2029]" pageId="6" pageNumber="51">
<emphasis id="20C1EA92C735EF783F96FEDCFE6CF6DB" bold="true" box="[151,453,1704,1730]" pageId="6" pageNumber="51">
<typeStatus id="CD0E8822C735EF783F96FEDCFF7DF6DB" box="[151,212,1704,1730]" pageId="6" pageNumber="51">Type</typeStatus>
material.
<typeStatus id="CD0E8822C735EF783E52FEDCFE6CF6DB" box="[339,453,1704,1730]" pageId="6" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
: ɗ (PAH09_COE6),
<collectingCountry id="6AA27610C735EF783DBCFEDCFC85F6DB" box="[701,812,1704,1730]" name="Malaysia" pageId="6" pageNumber="51">Malaysia</collectingCountry>
, Pahang, low pH swamp upstream on Sg
<taxonomicName id="D5B54D03C735EF783A0FFEDCFAFFF6DB" box="[1294,1366,1704,1730]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="bebar">Bebar</taxonomicName>
,
<date id="660B1040C735EF783A60FEDCFF7CF6FE" pageId="6" pageNumber="51" value="2009-09-22">22 ix 2009</date>
, leg. R.A. Dow,
<collectionCode id="74A4AE45C735EF783E9CFEB9FE53F6FE" LSID="urn:lsid:biocol.org:col:34992" box="[413,506,1741,1767]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="51">RMNH</collectionCode>
.
<typeStatus id="CD0E8822C735EF783D07FEB9FD2CF6FE" box="[518,645,1741,1767]" pageId="6" pageNumber="51" type="paratype">
<emphasis id="20C1EA92C735EF783D07FEB9FD2CF6FE" bold="true" box="[518,645,1741,1767]" pageId="6" pageNumber="51">Paratypes</emphasis>
</typeStatus>
: 2 ɗɗ (PAH09_COE7–8), Ψ (PAH09_COE9), data as
<typeStatus id="CD0E8822C735EF783A14FEB9FA28F6FE" box="[1301,1409,1741,1767]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
; 3 ɗɗ (PAH09_COE30–32), location as
<typeStatus id="CD0E8822C735EF783D57FE87FD16F714" box="[598,703,1779,1805]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
,
<date id="660B1040C735EF783DCDFE87FCE4F714" box="[716,845,1779,1805]" pageId="6" pageNumber="51" value="2009-09-23">23 ix 2009</date>
, leg. R.A. Dow; 3 ɗɗ, Ψ, location as
<typeStatus id="CD0E8822C735EF783A08FE87FADBF714" box="[1289,1394,1779,1805]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
,
<date id="660B1040C735EF783A7FFE87FF5BF72B" pageId="6" pageNumber="51" value="2009-09-23">23 ix 2009</date>
, leg. C.Y. Choong.
<typeStatus id="CD0E8822C735EF783ED6FF6CFDE0F72B" box="[471,585,1816,1842]" pageId="6" pageNumber="51" type="paratype">Paratypes</typeStatus>
in
<collectionCode id="74A4AE45C735EF783D6FFF6CFD60F72B" LSID="urn:lsid:biocol.org:col:34992" box="[622,713,1816,1842]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="51">RMNH</collectionCode>
, coll. Choong and coll. Dow.
</paragraph>
</subSubSection>
<subSubSection id="5AAF650BC735EF783FC4FF49FD3EF764" pageId="6" pageNumber="51" type="etymology">
<paragraph id="120A3680C735EF783FC4FF49FD3EF764" blockId="6.[151,1437,1704,2029]" pageId="6" pageNumber="51">
<emphasis id="20C1EA92C735EF783FC4FF49FEFCF74E" bold="true" box="[197,341,1853,1879]" pageId="6" pageNumber="51">Etymology:</emphasis>
<taxonomicName id="D5B54D03C735EF783E5DFF49FE1BF74E" box="[348,434,1853,1879]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="54" phylum="Arthropoda" rank="species" species="hoisen">Hoisen</taxonomicName>
, a noun in apposition. Named for Professor Emeritus Dr. Yong Hoi Sen in recognition of his contributions to zoology in
<collectingCountry id="6AA27610C735EF783D25FF17FD3AF764" box="[548,659,1891,1917]" name="Malaysia" pageId="6" pageNumber="51">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="5AAF650BC735EF773FC4FFFCFBB3F3DD" lastPageId="9" lastPageNumber="54" pageId="6" pageNumber="51" type="description">
<paragraph id="120A3680C735EF793FC4FFFCFE4AF15E" blockId="6.[151,1437,1704,2029]" lastBlockId="7.[151,1436,152,327]" lastPageId="7" lastPageNumber="52" pageId="6" pageNumber="51">
<emphasis id="20C1EA92C735EF783FC4FFFCFDE9F7BB" bold="true" box="[197,576,1928,1954]" pageId="6" pageNumber="51">
Description of
<typeStatus id="CD0E8822C735EF783E85FFFCFE5AF7BB" box="[388,499,1928,1954]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
male.
</emphasis>
Head: Labium pale. Mandible bases and genae yellow, labrum yellow except basally where dark brown. Anteclypeus pale along lower margin and more extensively centrally, brown above. Postclypeus very dark brown. Vertical face of frons shining black, with a pair of yellow transverse stripes, continuous with yellow on genae and nearly meeting centrally, yellow on genae extending narrowly beside eye margin to level of top of antennae bases, which are largely yellow. Horizontal surface of frons and vertex dark metallic green, ocelli whitish. Antennae dark brown except for yellow stripes along the anterior surface of the scape and pedicel and a yellow ring at top of scape. Occiput bronzy black except below genae where obscurely pale.
</paragraph>
<caption id="46CA6608C734EF793F96FCFCFDD0F4D9" httpUri="https://zenodo.org/record/197658/files/figure.png" pageId="7" pageNumber="52" targetBox="[154,1305,436,1125]" targetPageId="7">
<paragraph id="120A3680C734EF793F96FCFCFDD0F4D9" blockId="7.[151,1436,1160,1216]" pageId="7" pageNumber="52">
<emphasis id="20C1EA92C734EF793F96FCFCFEC9F4B9" bold="true" box="[151,352,1160,1184]" pageId="7" pageNumber="52">FIGURES 18–21.</emphasis>
Male caudal appendages, lateral view: (18)
<taxonomicName id="D5B54D03C734EF793C45FCFDFC0BF4B9" box="[836,930,1161,1184]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="bebar">
<emphasis id="20C1EA92C734EF793C45FCFDFC0BF4B9" box="[836,930,1161,1184]" italics="true" pageId="7" pageNumber="52">A. bebar</emphasis>
</taxonomicName>
holotype; (19)
<taxonomicName id="D5B54D03C734EF793B4DFCFDFAECF4B9" authority="(Sarawak)" baseAuthorityName="Sarawak" box="[1100,1349,1161,1184]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="ecornuta">
<emphasis id="20C1EA92C734EF793B4DFCFDFB62F4B9" box="[1100,1227,1161,1184]" italics="true" pageId="7" pageNumber="52">A. ecornuta</emphasis>
(Sarawak)
</taxonomicName>
; (20)
<taxonomicName id="D5B54D03C734EF793A84FCFDFEC7F4D9" authority="Sg. Bebar" authorityName="Sg. Bebar" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="20C1EA92C734EF793A84FCFDFF41F4D9" italics="true" pageId="7" pageNumber="52">A. gracilis</emphasis>
(Sg. Bebar)
</taxonomicName>
; (21)
<taxonomicName id="D5B54D03C734EF793EAAFCDDFDA7F4D9" box="[427,526,1193,1216]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="hoisen">
<emphasis id="20C1EA92C734EF793EAAFCDDFDA7F4D9" box="[427,526,1193,1216]" italics="true" pageId="7" pageNumber="52">A. hoisen</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<paragraph id="120A3680C734EF793FC4FC9AFBB8F70F" blockId="7.[151,1438,1262,2038]" pageId="7" pageNumber="52">
Thorax: Prothorax with pronotum almost entirely dark metallic green, becoming bronzy black to sides, with small obscurely pale areas laterally on anterior lobe. Propleuron bronzy black above, cream below. Synthorax with mesepisternum and mesepimeron dark metallic green, as is the upper part of the Mesinfraepisternum, the lower part is cream. Metallic green colouration extending onto upper part of metepisternum near mesinfraepisternum and more widely to rear, reaching to metepleural suture near wing bases, cream below as is entirety of metepimeron, metinfraepisternum and venter of synthorax. Legs with coxae cream coloured, trochanters cream above becoming yellow distally. Femur orange-yellow on all legs, except for small dark brown area immediately above tibia. Tibia pale yellow with a long dark streak on extensor surface of anterior tibia originating immediately below femur, smaller dark streaks below femur on flexor surface and laterally on extensor surface of other legs. Tarsi cream with brown spines and articulations, tarsal claws pale brown with darker tips and a tiny rudimentary tooth. Wings with 14 Px in Fw and 13 Px in Hw. Arculus at Ax 2 (left side) or slightly distal to it (right side). 1A arising distal of Ac, IR3 and R4 arising distal of subnodus. Fw Pt (
<figureCitation id="8A8E2A05C734EF793EE2FEC5FD97F6D2" box="[483,574,1713,1739]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,272,898,922]" captionTargetBox="[152,1434,449,862]" captionTargetId="figure@6.[151,1436,421,877]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pterostigma: (9) A. bebar holotype male Fw; (10) A. ecornuta male Fw (Sarawak); (11) A. gracilis male Fw (Sg. Bebar); (12) A. hoisen holotype male Fw; (13) A. hoisen holotype male Hw; (14) A. hoisen paratype female Fw (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197656/files/figure.png" pageId="7" pageNumber="52">Fig. 12</figureCitation>
) covering slightly more than one underlying cell, almost square, dark brown, with a yellow outline, this best developed on distal and posterior margins. Hw Pt (
<figureCitation id="8A8E2A05C734EF793BAEFEA2FAADF6E9" box="[1199,1284,1750,1776]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,272,898,922]" captionTargetBox="[152,1434,449,862]" captionTargetId="figure@6.[151,1436,421,877]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pterostigma: (9) A. bebar holotype male Fw; (10) A. ecornuta male Fw (Sarawak); (11) A. gracilis male Fw (Sg. Bebar); (12) A. hoisen holotype male Fw; (13) A. hoisen holotype male Hw; (14) A. hoisen paratype female Fw (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197656/files/figure.png" pageId="7" pageNumber="52">Fig. 13</figureCitation>
) deeper than long, bright orange-yellow, covering slightly more than one underlying cell.
</paragraph>
<paragraph id="120A3680C734EF763FC4FF55FC8EF0E5" blockId="7.[151,1438,1262,2038]" lastBlockId="8.[151,1436,152,252]" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
Abdomen: S1 very dark shining brown above, pale cream to sides. S2 bronzy brown dorsally, darker to rear, with a narrow pale basal annulus. S3–7 brown dorsally, becoming darker on S7, paler laterally (again S7 darker), this extended dorsally as a narrow basal annulus, becoming less clear on successive segments. S8–9 mostly bronzy brown to black, S8 only narrowly pale laterally and S9 mostly black laterally with an obscure paler area. S10 small, entirely pale cream coloured. Genital valves (
<figureCitation id="8A8E2A05C734EF793CCDFFC2FB8AF7C9" box="[972,1059,1974,2000]" captionStart="FIGURES 15 – 17" captionStartId="6.[151,269,1490,1514]" captionTargetBox="[169,1419,1046,1453]" captionTargetId="figure@6.[158,1428,1036,1466]" captionTargetPageId="6" captionText="FIGURES 15 – 17. Genital valves: (15) A. bebar holotype; (16) A. ecornuta (Sarawak); (17) A. hoisen holotype." httpUri="https://zenodo.org/record/197657/files/figure.png" pageId="7" pageNumber="52">Fig. 17</figureCitation>
) very large with long posterior spines laying flat against sternum of S10. Cerci (
<figureCitation id="8A8E2A05C734EF793DDAFFA8FC97F7EF" box="[731,830,2012,2038]" captionStart="FIGURES 18 – 21" captionStartId="7.[151,271,1160,1184]" captionTargetBox="[154,1305,436,1125]" captionTargetId="figure@7.[151,1436,383,1139]" captionTargetPageId="7" captionText="FIGURES 18 – 21. Male caudal appendages, lateral view: (18) A. bebar holotype; (19) A. ecornuta (Sarawak); (20) A. gracilis (Sg. Bebar); (21) A. hoisen holotype." httpUri="https://zenodo.org/record/197658/files/figure.png" pageId="7" pageNumber="52">Figs. 21</figureCitation>
,
<figureCitation id="8A8E2A05C734EF793C4AFFA8FCC3F7EF" box="[843,874,2012,2038]" captionStart="FIGURES 22 – 25" captionStartId="8.[151,271,1323,1347]" captionTargetBox="[197,1392,314,1271]" captionTargetId="figure@8.[187,1400,306,1302]" captionTargetPageId="8" captionText="FIGURES 22 – 25. Male cerci, dorsal views: (22) A. bebar holotype; (23) A. ecornuta (Sarawak); (24) A. gracilis (Sg. Bebar); (25) A. hoisen holotype." httpUri="https://zenodo.org/record/197659/files/figure.png" pageId="7" pageNumber="52">25</figureCitation>
) with upper branch with prominent inward and ventrally directed projection at ca mid length, after which the long tip turns abruptly downwards. Lower branch short, just reaching the level of the projection on the upper branch, of distinctive shape in ventral view (
<figureCitation id="8A8E2A05C73BEF763F9EF896FF5EF0E5" box="[159,247,226,252]" captionStart="FIGURE 26. A" captionStartId="8.[151,255,1980,2004]" captionTargetBox="[506,1087,1464,1931]" captionTargetId="figure@8.[491,1096,1430,1959]" captionTargetPageId="8" captionText="FIGURE 26. A. hoisen holotype, ventral view lower branch cercus, right side. A part of the right paraproct is visible at the base of the cercus." httpUri="https://zenodo.org/record/197660/files/figure.png" pageId="8" pageNumber="53">Fig. 26</figureCitation>
). Epiproct prominent, paraprocts barely visible.
</paragraph>
<caption id="46CA6608C73BEF763F96FD5FFE46F57A" httpUri="https://zenodo.org/record/197659/files/figure.png" pageId="8" pageNumber="53" targetBox="[197,1392,314,1271]" targetPageId="8">
<paragraph id="120A3680C73BEF763F96FD5FFE46F57A" blockId="8.[151,1436,1323,1379]" pageId="8" pageNumber="53">
<emphasis id="20C1EA92C73BEF763F96FD5FFEC8F55A" bold="true" box="[151,353,1323,1347]" pageId="8" pageNumber="53">FIGURES 22–25.</emphasis>
Male cerci, dorsal views: (22)
<taxonomicName id="D5B54D03C73BEF763DB7FD58FCBDF55A" box="[694,788,1324,1347]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="bebar">
<emphasis id="20C1EA92C73BEF763DB7FD58FCBDF55A" box="[694,788,1324,1347]" italics="true" pageId="8" pageNumber="53">A. bebar</emphasis>
</taxonomicName>
holotype; (23)
<taxonomicName id="D5B54D03C73BEF763CBCFD58FB1EF55A" authority="(Sarawak)" baseAuthorityName="Sarawak" box="[957,1207,1324,1347]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ecornuta">
<emphasis id="20C1EA92C73BEF763CBCFD58FB95F55A" box="[957,1084,1324,1347]" italics="true" pageId="8" pageNumber="53">A. ecornuta</emphasis>
(Sarawak)
</taxonomicName>
; (24)
<taxonomicName id="D5B54D03C73BEF763BF7FD58FF4AF57A" authority="Sg. Bebar" authorityName="Sg. Bebar" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="20C1EA92C73BEF763BF7FD58FAC1F55A" box="[1270,1384,1324,1347]" italics="true" pageId="8" pageNumber="53">A. gracilis</emphasis>
(Sg. Bebar)
</taxonomicName>
; (25)
<taxonomicName id="D5B54D03C73BEF763E21FD38FE2AF57A" box="[288,387,1356,1379]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="hoisen">
<emphasis id="20C1EA92C73BEF763E21FD38FE2AF57A" box="[288,387,1356,1379]" italics="true" pageId="8" pageNumber="53">A. hoisen</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<caption id="46CA6608C73BEF763F96FFC8FE2DF7EA" httpUri="https://zenodo.org/record/197660/files/figure.png" pageId="8" pageNumber="53" targetBox="[506,1087,1464,1931]" targetPageId="8">
<paragraph id="120A3680C73BEF763F96FFC8FE2DF7EA" blockId="8.[151,1436,1980,2035]" pageId="8" pageNumber="53">
<emphasis id="20C1EA92C73BEF763F96FFC8FE8EF7CD" bold="true" box="[151,295,1980,2004]" pageId="8" pageNumber="53">FIGURE 26.</emphasis>
<taxonomicName id="D5B54D03C73BEF763E2FFFC8FE3AF7CA" box="[302,403,1980,2003]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="hoisen">
<emphasis id="20C1EA92C73BEF763E2FFFC8FE3AF7CA" box="[302,403,1980,2003]" italics="true" pageId="8" pageNumber="53">A. hoisen</emphasis>
</taxonomicName>
holotype, ventral view lower branch cercus, right side. A part of the right paraproct is visible at the base of the cercus.
</paragraph>
</caption>
<paragraph id="120A3680C73AEF773FC4F8E3FA87F0AB" blockId="9.[151,1436,151,1226]" box="[197,1326,151,178]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4F8E3FED4F0A8" bold="true" box="[197,381,151,177]" pageId="9" pageNumber="54">Measurements</emphasis>
(mm): Hw 19.5, abdomen without appendages 31.5, upper branch cercus ca 0.8.
</paragraph>
<paragraph id="120A3680C73AEF773FC4F8C9FE5BF13B" blockId="9.[151,1436,151,1226]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4F8C9FDE4F0CE" bold="true" box="[197,589,189,215]" pageId="9" pageNumber="54">
Description of female
<typeStatus id="CD0E8822C73AEF773EDDF8C9FDE4F0CE" box="[476,589,189,215]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
</emphasis>
(PAH09_COE9). As male except as noted. Head: Labrum only pale in anterior 1/3. Transverse stripes on vertical face of frons joined centrally. A narrow pale stripe along transverse occipital carina behind ocelli.
</paragraph>
<paragraph id="120A3680C73AEF773FC4F959FB10F26D" blockId="9.[151,1436,151,1226]" pageId="9" pageNumber="54">
Thorax: Prothorax largely pale greenish blue, except along the lower margin of the propleuron, where cream, and along the anterior margin of the anterior pronotal lobe and centrally on the posterior pronotal lobe, where black (
<figureCitation id="8A8E2A05C73AEF773E35F90CFED2F18B" box="[308,379,376,402]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,279,1620,1644]" captionTargetBox="[151,1436,983,1595]" captionTargetId="figure@4.[151,1436,983,1595]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Female prothorax, dorsal view: (1) A. ecornuta (Sarawak); (2) A. hoisen paratype female (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197653/files/figure.png" pageId="9" pageNumber="54">Fig. 2</figureCitation>
). Posterior pronotal lobe simple, shaped as in
<figureCitation id="8A8E2A05C73AEF773C92F90CFC4CF18B" box="[915,997,376,402]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,279,1620,1644]" captionTargetBox="[151,1436,983,1595]" captionTargetId="figure@4.[151,1436,983,1595]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Female prothorax, dorsal view: (1) A. ecornuta (Sarawak); (2) A. hoisen paratype female (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197653/files/figure.png" pageId="9" pageNumber="54">Figs. 2</figureCitation>
,
<figureCitation id="8A8E2A05C73AEF773CF1F90CFBABF18B" box="[1008,1026,376,402]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,273,798,822]" captionTargetBox="[177,1412,430,759]" captionTargetId="figure@5.[169,1418,421,773]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Female posterior pronotal lobe, lateral view: (3) A. bebar paratype female (PAH 09 _ COE 1); (4) A. ecornuta (Sarawak); (5) A. gracilis (Sg. Bebar); (6) A. hoisen paratype female (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197654/files/figure.png" pageId="9" pageNumber="54">6</figureCitation>
. Mesepisternum pale greenish blue with a strong yellow tint, antealar triangles black. Mesepimeron and mesinfraepisternum pale greenish blue, this extending onto the upper half of the metepisternum, rest of synthorax cream. Legs cream except a black stripe along almost entire length of flexor surface of femurs, a small black lateral mark on femur just above joint, and obscure black marks below joint on tibia. Wings with 13 Px in Fw, 12 Px in Hw, R4 arising at subnodus in all wings. Pt (
<figureCitation id="8A8E2A05C73AEF773ECCFA41FD8BF256" box="[461,546,565,591]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,272,898,922]" captionTargetBox="[152,1434,449,862]" captionTargetId="figure@6.[151,1436,421,877]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pterostigma: (9) A. bebar holotype male Fw; (10) A. ecornuta male Fw (Sarawak); (11) A. gracilis male Fw (Sg. Bebar); (12) A. hoisen holotype male Fw; (13) A. hoisen holotype male Hw; (14) A. hoisen paratype female Fw (PAH 09 _ COE 9)." httpUri="https://zenodo.org/record/197656/files/figure.png" pageId="9" pageNumber="54">Fig. 14</figureCitation>
) nearly square but deeper than long on costal side, covering one underlying cell, grey brown centrally with a wide cream border, especially pronounced on costal side.
</paragraph>
<paragraph id="120A3680C73AEF773FC4FAF4FCB2F313" blockId="9.[151,1436,151,1226]" pageId="9" pageNumber="54">Abdomen: Very similar to male, except more brown above, with pale basal annuli better developed on S2– 7. S8 dark brown above, obscurely pale laterally. S9 dark brown above, with obscure pale patches laterally, especially to rear. S10 entirely pale cream except ventrally. Cerci and paraprocts pale cream, as is the ovipositor, except the gonapophyses, which are brown.</paragraph>
<paragraph id="120A3680C73AEF773FC4FB61FB2DF336" blockId="9.[151,1436,151,1226]" box="[197,1156,789,815]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4FB61FED4F336" bold="true" box="[197,381,789,815]" pageId="9" pageNumber="54">Measurements</emphasis>
(mm): Hw 20.5, abdomen without appendages or ovipositor 31.5.
</paragraph>
<paragraph id="120A3680C73AEF773FC4FB4EFDC5F386" blockId="9.[151,1436,151,1226]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4FB4EFE4DF34D" bold="true" box="[197,484,826,852]" pageId="9" pageNumber="54">
Variation in
<typeStatus id="CD0E8822C73AEF773E60FB4EFE76F34D" box="[353,479,826,852]" pageId="9" pageNumber="54" type="paratype">paratypes</typeStatus>
.
</emphasis>
Teneral males have the Pt pale yellow in Fw and Hw, and the margins of the dark areas of the prothorax and adjacent parts of the synthorax, as well as the pale area laterally on the anterior lobe of the pronotum, are strongly red tinted.
</paragraph>
<paragraph id="120A3680C73AEF773FC4FBDEFBB3F3DD" blockId="9.[151,1436,151,1226]" box="[197,1050,938,964]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4FBDEFED5F3DD" bold="true" box="[197,380,938,964]" pageId="9" pageNumber="54">Measurements</emphasis>
(mm): Hw 19.5–20.5, abdomen without cerci 30.5–32.5.
</paragraph>
</subSubSection>
<subSubSection id="5AAF650BC73AEF773FC4FBA4FD66F4D3" pageId="9" pageNumber="54" type="diagnosis">
<paragraph id="120A3680C73AEF773FC4FBA4FD66F4D3" blockId="9.[151,1436,151,1226]" pageId="9" pageNumber="54">
<emphasis id="20C1EA92C73AEF773FC4FBA4FE94F3F3" bold="true" box="[197,317,976,1002]" pageId="9" pageNumber="54">Diagnosis</emphasis>
. Both sexes of
<taxonomicName id="D5B54D03C73AEF773EF1FBA4FDF5F3F0" box="[496,604,976,1001]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="hoisen">
<emphasis id="20C1EA92C73AEF773EF1FBA4FDF5F3F0" box="[496,604,976,1001]" italics="true" pageId="9" pageNumber="54">A. hoisen</emphasis>
</taxonomicName>
are clearly separated from all species known from Peninsular
<collectingCountry id="6AA27610C73AEF773A31FBA4FA35F3F3" box="[1328,1436,976,1002]" name="Malaysia" pageId="9" pageNumber="54">Malaysia</collectingCountry>
and
<collectingCountry id="6AA27610C73AEF773FCBFB81FEE8F416" box="[202,321,1013,1039]" name="Singapore" pageId="9" pageNumber="54">Singapore</collectingCountry>
by the characters given in the keys. The closest allies of
<taxonomicName id="D5B54D03C73AEF773CEBFB82FBF0F416" box="[1002,1113,1014,1039]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="hoisen">
<emphasis id="20C1EA92C73AEF773CEBFB82FBF0F416" box="[1002,1113,1014,1039]" italics="true" pageId="9" pageNumber="54">A. hoisen</emphasis>
</taxonomicName>
are the Bornean species
<taxonomicName id="D5B54D03C73AEF773A82FB82FECFF42D" authority="Ris, 1911" authorityName="Ris" authorityYear="1911" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="20C1EA92C73AEF773A82FB82FF47F42D" italics="true" pageId="9" pageNumber="54">A. martini</emphasis>
Ris, 1911
</taxonomicName>
and
<taxonomicName id="D5B54D03C73AEF773EA1FC6FFCB8F42D" authority="Lieftinck, 1953" authorityName="Lieftinck" authorityYear="1953" box="[416,785,1050,1076]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="dactylostyla">
<emphasis id="20C1EA92C73AEF773EA1FC6FFDE6F42D" box="[416,591,1051,1076]" italics="true" pageId="9" pageNumber="54">A. dactylostyla</emphasis>
Lieftinck, 1953
</taxonomicName>
, from which the male differs in the details of the cerci, especially the extremely long down turned terminal part of the upper branch, and the shape of the lower branch seen in ventral view. The male differs from all other species in the remarkable long posterior spine on the genital valves (
<figureCitation id="8A8E2A05C73AEF773E77FCFEFE64F4BD" box="[374,461,1162,1188]" captionStart="FIGURES 15 – 17" captionStartId="6.[151,269,1490,1514]" captionTargetBox="[169,1419,1046,1453]" captionTargetId="figure@6.[158,1428,1036,1466]" captionTargetPageId="6" captionText="FIGURES 15 – 17. Genital valves: (15) A. bebar holotype; (16) A. ecornuta (Sarawak); (17) A. hoisen holotype." httpUri="https://zenodo.org/record/197657/files/figure.png" pageId="9" pageNumber="54">Fig. 17</figureCitation>
); this character is so distinctive that it would allow identification of a specimen so badly damaged that only S9–10 remained intact.
</paragraph>
</subSubSection>
</treatment>
</document>
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