320 lines
38 KiB
XML
320 lines
38 KiB
XML
<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="7990FB77E5105D75BB4FA97D31F18594" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) vanensis Guéorguiev & Wrase & Assmann & Muilwijk & Machard 2022, sp. nov." docType="treatment" docUuid="5DAEBFAF-5211-4ED1-83DE-822DD8402705" docUuidSource="ZooBank" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
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<mods:title id="CB884E12C4A8509504827E2B718FCC78">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart id="78DBF824BDF779BF3E1591CC94DBF576">Gueorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier id="F7A71C226E85EF34344B5EB678CD5E9E" type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
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<mods:affiliation id="757EA9BA97D024A49A7806602A89CB27">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier id="DE83A1985AE2E22BDCD8F2431A1297A8" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart id="C8E47BBB5C63CAB35F65C7B49B448963">Wrase, David W.</mods:namePart>
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<mods:nameIdentifier id="277F776349FE9D32D0C733C25B0CCF8B" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation id="4603972F38CDBDFCDB91E34FF8F2842A">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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<mods:roleTerm id="36E0F6D78795638DEE754A275CAF79F4">Author</mods:roleTerm>
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<mods:namePart id="B618FB1B9E03F7886BAA4623F8D254AE">Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier id="D2D553C34F8435DE95DAF3FB72F6ED6E" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation id="08DC14E542F774D5017338FD93F94200">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:namePart id="9178E5EBCDA5F8554F03C5C7E9751F5F">Muilwijk, Jan</mods:namePart>
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<mods:affiliation id="7EC64E6F888A6D819F339A7B7E3E461C">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="E372B04B48B592B3DA827A6C44F828D3">Machard, Patrice</mods:namePart>
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<mods:affiliation id="19D55D8814DE37C27D50697604452A40">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title id="B8197B0B0A309DEA43B6DEA0B1BBF8F0">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="4FE5CA5A1088B3DF71E6A1D493E64A44">2022</mods:date>
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<mods:number id="B83ABAD11DD6D99B9DE36B22A4B49F51">2022-06-13</mods:number>
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<mods:number id="E329EC1EBA8718243DEEADFFE4848D51">69</mods:number>
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<mods:start id="80202D60EE045179327E3FC569BC6C3E">71</mods:start>
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<mods:url id="69EB37F0A0600BCCF82BD6E3493CD55A">http://dx.doi.org/10.3897/dez.69.83840</mods:url>
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<mods:classification id="C0564E0AC14FC9C7EF925E218E81BFDD">journal article</mods:classification>
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<mods:identifier id="F6105EF634838004EE3781A74598A397" type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
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<mods:identifier id="096C0E4D40C76CE54A1BB799066C0B2B" type="Pensoft-Pub">1860-1324-1-71</mods:identifier>
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<treatment id="7990FB77E5105D75BB4FA97D31F18594" ID-GBIF-Taxon="195868753" LSID="urn:lsid:zoobank.org:act:5DAEBFAF-5211-4ED1-83DE-822DD8402705" httpUri="http://treatment.plazi.org/id/7990FB77E5105D75BB4FA97D31F18594" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
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<subSubSection id="7527D7820B8855B9E414134D4600D376" pageId="0" pageNumber="71" type="nomenclature">
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<paragraph id="249E7790A4CEB1817D1579782B3B725C" pageId="0" pageNumber="71">
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5.
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<taxonomicName id="D5CBE350565AB7B8D8BAB7ADB869CFAD" LSID="http://zoobank.org/5DAEBFAF-5211-4ED1-83DE-822DD8402705" authority="Guéorguiev & Wrase & Assmann & Muilwijk & Machard, 2022" authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) vanensis</taxonomicName>
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<taxonomicNameLabel id="E5E287F04D34219AF89206977C67FF61" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="74236E33697054094DDE08F69D6CAD64" pageId="0" pageNumber="71" type="description">
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<paragraph id="98F54E730E50287AAF49C93448EFC787" pageId="0" pageNumber="71">
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<figureCitation id="127286FE560F73DACCC617DA00B73245" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1E</figureCitation>
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<figureCitation id="2696A128378861D55EACBF3D0302354C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8E</figureCitation>
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<figureCitation id="1FD0D4AA7A5A0FE059E9210E83EBF619" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10E</figureCitation>
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<figureCitation id="D96C5ADAACC7C896E97B52FBC58C8BBC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13D, E</figureCitation>
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<figureCitation id="182AB8E312FC5A4E9394A9838522FE28" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="36D590911FC607E2811B515CCD78BE47" pageId="0" pageNumber="71" type="type locality">
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<paragraph id="6BE33782730967AE9179E876AEF56B74" pageId="0" pageNumber="71">Type locality.</paragraph>
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<paragraph id="CF0049431E8F2789975AE03CDC7A471B" pageId="0" pageNumber="71">
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Turkey, Van Province,
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<normalizedToken id="C7A173F288F701054F144B716CEB757A" originalValue="Gevaş">Gevas</normalizedToken>
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Town environs.
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</paragraph>
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</subSubSection>
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<subSubSection id="6ACBC50DB8F702C3542352DD278231B9" pageId="0" pageNumber="71" type="note on type locality">
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<paragraph id="343BEBFB35C91559E9529DB1F9248E91" pageId="0" pageNumber="71">Note on type locality.</paragraph>
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<paragraph id="B665010B91021DF8800E830630E5F5C4" pageId="0" pageNumber="71">
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<normalizedToken id="84D4E459C4BAD6C26BB6E2E5F672138E" originalValue="Gevaş">Gevas</normalizedToken>
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Town is situated on the southern shore of Van Lake. The holotype was perhaps collected in vicinities of the town.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EA240491B982E7AD889CDF045842E9E" pageId="0" pageNumber="71" type="type material">
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<paragraph id="1A47DED39DC8C0F24C1072EC1F298644" pageId="0" pageNumber="71">Type material.</paragraph>
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<paragraph id="246EA9B9933225798DE2F060E5A5392A" pageId="0" pageNumber="71">
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<emphasis id="778CE5DE7C16C34B201C4257E8A74FE8" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
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: 1♂, 'TURCIA or. Van / Gevas 29.6.1993 / lgt. J.
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<normalizedToken id="B7DEE05417173E282B4B277CE9070337" originalValue="Růžička">Ruzicka</normalizedToken>
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//
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<taxonomicName id="B05B72A796EBE56AA14E01C0D5E56DF5" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
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/ cf. / punctiger / (REICHE & SAULCY) / WRASE det. 2008' (cWR). Paratypes: 1♀, 'TURKEY OR. / GEVAS env., 2100-2600m / (CADIR DAGI) / 1993-06-29,
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<normalizedToken id="7D7966B3320891D7DBFFE15F978D9F9D" originalValue="Klíma">Klima</normalizedToken>
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lgt. //
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<taxonomicName id="C07D210440AF1889285B3AF615DA3832" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
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/ cf. / punctiger / (REICHE & SAULCY) / WRASE det. 2008' (NMNHS); 1♂, 1♀, 'Resadiye 1900m / TR: Bitlis 6.91 / coll. B.
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<normalizedToken id="9E5820034AB3ED0F3E27A76C0F06B752" originalValue="Lassalle’">Lassalle'</normalizedToken>
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(cLAS); 1♀, 'CE TURKEY, prov. Bitlis /
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<geoCoordinate id="C18DD8C8F30EAB7C1AFE2F1BCA3E8A60" degrees="38" direction="north" minutes="37" orientation="latitude" precision="925" value="38.616665">38°37'N</geoCoordinate>
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,
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<geoCoordinate id="141D7D4BE8F51ECD36A6092E474FD494" degrees="42" direction="east" minutes="16" orientation="longitude" precision="925" value="42.266666">42°16'E</geoCoordinate>
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2290m /
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<normalizedToken id="AB841C27BF7FBD232324345243699CCA" originalValue="Şentepe">Sentepe</normalizedToken>
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env. / Nemrut Mt. - crater / 24.VI.08 lgt. E.Hajdaj // COLL. / E. &
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<taxonomicName id="CF97137CA8C53AB84BD27507050166D8" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
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HAJDAJ /
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<normalizedToken id="8A96541BF28EB614356EB046AAB71535" originalValue="JEŽOV">JEZOV</normalizedToken>
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/ Czech
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<normalizedToken id="B7AC6866719447D88639EADF115C8094" originalValue="Republic’">Republic'</normalizedToken>
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(cHAJ); 5♂♂, 2♀♀, 'TR, Bitlis, Yelkenli / Van lake, ca. 1800 m / ~
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<geoCoordinate id="0F437F0ED2D63C5A55DF4A758B8D78ED" degrees="38" direction="north" minutes="28" orientation="latitude" precision="925" value="38.466667">38°28'N</geoCoordinate>
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,
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<geoCoordinate id="C9234CE53B81BEE090F177D6537C57F5" degrees="42" direction="east" minutes="32" orientation="longitude" precision="925" value="42.533333">42°32'E</geoCoordinate>
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/ 21.IV.-20.V.2014 / pitfall, leg. C.
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<normalizedToken id="3C9901E4C2094B75106AF0A61A002663" originalValue="Reuter’">Reuter'</normalizedToken>
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(NMNHS, cREU, cWR); 1♀, 'col de Buglan 1500m / TR Mus 6.88 / col. B.
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<normalizedToken id="B48EB3CDEEF8E741E45181184F9C1041" originalValue="Lassalle’">Lassalle'</normalizedToken>
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(cLAS); 1♂, 'E Turkey, Mus / 8km SE Solhan, Buglan / Gec., 1700-1800m /
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<geoCoordinate id="5859EBB69E968AEEE70B794AA8BDCCD9" degrees="38" direction="north" minutes="56" orientation="latitude" precision="925" value="38.933334">38°56'N</geoCoordinate>
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,
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<geoCoordinate id="F240B25A6115688CD68B1228E3A8A691" degrees="41" direction="east" minutes="08" orientation="longitude" precision="925" value="41.133335">41°08'E</geoCoordinate>
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18. - / 20.VI.05 lgt. E.&
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<taxonomicName id="34C808B38C122DE93FF3DCE95639775E" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
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<normalizedToken id="EA19548702AFCE1C87429E83C7600898" originalValue="Hajdaj’">Hajdaj'</normalizedToken>
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(cHAJ); 3♂♂, 5♀♀, 'TR,
|
||
<normalizedToken id="1FF4FB702B0D8F368FBB3FD5C2A080A0" originalValue="Bingöl">Bingoel</normalizedToken>
|
||
/
|
||
<normalizedToken id="B60D34C811B30903066373C4EFB12661" originalValue="Muş">Mus</normalizedToken>
|
||
/
|
||
<normalizedToken id="3E07581BEE1AC5F4162AB420B47DA42F" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="A56B0D9CC34605102B7659B503E07B0F" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
, 1640 m / 21.IV. - 11.V.2014 / pitfall, leg. C.
|
||
<normalizedToken id="7070940EF0D441F2250A34E8490643F1" originalValue="Reuter’">Reuter'</normalizedToken>
|
||
(NMNHS, cREU, cWR).
|
||
</paragraph>
|
||
<paragraph id="02ACD96074917EB4DA57E5C04D6C43F0" pageId="0" pageNumber="71">TME: 22 specimens. TGE: 4♂♂, 2♀♀.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F6AC246A3B76B025BFADA7B0B42550EA" pageId="0" pageNumber="71" type="etymology">
|
||
<paragraph id="F9AF94069CA219E8D19D8A9A19D5B35A" pageId="0" pageNumber="71">Etymology.</paragraph>
|
||
<paragraph id="07A877074DFEC5C38018817D8D52ED60" pageId="0" pageNumber="71">An adjective derived from the geographical name Van.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EF3BDF653056509EBA3B988AF4B85A" pageId="0" pageNumber="71" type="diagnosis">
|
||
<paragraph id="E3550F2BB026552C000D3EA1CC87B7F1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
|
||
<paragraph id="0C3EEC7F9AB49F94E81EE2CC6D10B809" pageId="0" pageNumber="71">
|
||
The new species is most closely related to
|
||
<taxonomicName id="36C57071B27C7949CAC9DB40294131F7" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
|
||
<emphasis id="EBA1C897BD9B78FAD4B4D8EBB753B346" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., but easily differs from it by the apical lamella of the median lobe (dorsal and ventral view) that is less symmetrical and shorter (Fig.
|
||
<figureCitation id="C7033418CCA1423BA4285BB4722E4182" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
|
||
), conical shape of bursa copulatrix (Fig.
|
||
<figureCitation id="0CB3E936202603E47CE10AF39D1C7140" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
|
||
), darker coloration of the integument, elytral striae and striole more coarsely punctate and smaller value PW/
|
||
<normalizedToken id="74AABAECBCF5313982097F0717A345F4" originalValue="PА">PA</normalizedToken>
|
||
(1.32-1.41, vs. 1.41-1.42).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4A62B98FA6013458501C4C09F08FDD54" pageId="0" pageNumber="71" type="description">
|
||
<paragraph id="C6FC41D474926E33683C40209C8BB9D1" pageId="0" pageNumber="71">Description.</paragraph>
|
||
<paragraph id="68F326A994623064FD67D719762D2B9C" pageId="0" pageNumber="71">
|
||
<emphasis id="F990C4B510E1376384AA9858A64F848F" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="351163CB6C30DF27E117C6A932D48439" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Specimens of middle size for
|
||
<taxonomicName id="205920FA9C8D8FF869DF5C51FC75DCBF" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EE238B2A21F5CA2A6AE22A46853B1BA6" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
|
||
</taxonomicName>
|
||
species (BL: 6.85-8.40 mm; BW: 2.25-2.70 mm), with subelongate, slender body (Fig.
|
||
<figureCitation id="D06EE6F12E903BE2896DB0A49E412744" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1E</figureCitation>
|
||
).
|
||
<emphasis id="EB5465FCB19B99DC622586976D3F7FE9" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="F0D5E1C159BDC7E0381160145C2F3E94" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
|
||
.
|
||
</emphasis>
|
||
See Table
|
||
<tableCitation id="47465F20630E3C12989811E410748FCE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
|
||
.
|
||
<emphasis id="6EE4CDF262B197426FB47E9DB00180C5" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="EEF569E729996B7CB8DDCE514848FEE6" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body dorsally and ventrally mostly reddish-brown, appendages (antennae, palpi and legs) lighter colored than body, some specimens from
|
||
<normalizedToken id="E5A6680B452A782AA553E031B2C7F6FD" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="FA817D20B664B659AAC2F3848983678B" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
with elytra almost black, and head, pronotum and ventral surface reddish-brown. Integument rather shiny.
|
||
<emphasis id="CCBFB129B8FDBF331B2C0B3384B2792C" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="EF55B152A4FBAFC6C94338DC0FC3AFFC" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pronotum and elytral intervals 1-5 (rarely also 6-7) without distinct sculpticells or with very faint ones, intervals 6-9 mostly (all specimens from Yelkenli) with distinct isodiametric sculpticells. Ventral surface mostly without visible sculpticells, sometimes with scarcely-visible isodiametric meshes. Pronotum coarsely and densely punctate in basal third and adjacent lateral areas, punctures at sides often reaching medial third. Abdominal ventrites 1-4 punctate at sides.
|
||
<emphasis id="78734751893C6C68B6DB5BF575623AC1" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="27E2133D22A6209ED70A1060FFDF4C9E" italics="true" pageId="0" pageNumber="71">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
About two thirds as wide as pronotum (PW/HW= 1.30-1.43). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes long, subconvex. Labrum subrectangular, with anterior margin slightly concave. Frontal furrows shallow, subfoveolate. Paraorbital sulci moderately deep, ending slightly before level of posterior supraorbital pore.
|
||
<emphasis id="541C4D4638574BEF9FA83C26D4AE4BDB" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="E0335D30BD84726C7F5E25CE88FAE1F2" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pronotum wider than long (PW/PL= 1.13-1.24), widest point at anterior third. Anterior transverse impression barely distinct, posterior one distinct medially. Sides sinuate, convex anteriorly, concave posteriorly; anterior bead reduced in medial 1/10. Metepisternum slightly longer than wide, MA/MI= 0.87-0.93.
|
||
<emphasis id="DC25120E7CFE08234D48B4E1817F4457" bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
|
||
. Elongate, about one and two thirds as long as wide (EL/EW= 1.61-1.73), two and a half times as long as pronotum (EL/PL= 2.52-2.74), and one third wider than pronotum (EW/PW= 1.22-1.36), with widest point at medial fifth. Parascutellar striole and striae deeply impressed (specimens from
|
||
<normalizedToken id="98AF4DD97431CE438437934DC3D514D5" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="FD667B8C461D38393575DF4D1D6D1EAF" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
with striae 7-8 less impressed than others); parascutellar striole short, not connected with stria 1. Interval 3 with three discal setiferous punctures. Umbilicate setiferous punctures consisting of 16-17 (rarely 18) each side.
|
||
<emphasis id="9899F7860C5A2AC4F4DEB99FCE527F20" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="5D9C48A7E22F0851C3E11DD152280760" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with three or four setiferous punctures (three specimens with two on one side) on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
|
||
<emphasis id="C68ACBA10EEDF53350EE3176FFEB801D" bold="true" pageId="0" pageNumber="71">
|
||
<emphasis id="3AC1571A2AB17EBE42B51A6DBA77E599" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Median lobe of aedeagus in lateral view, with elongate basal bulb, long and broad shaft moderately constricted proximally, and short, straight apex; median lobe in ventral view straight, about 3.4 times longer than wide; apical lamella (dorsal view) short, symmetrical, rounded at tip, with sides straight or slightly convex. Internal sac in lateral view (Fig.
|
||
<figureCitation id="66882323D2A8FFD08BA4161F287FEF81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8E</figureCitation>
|
||
) with ventral sclerite distinctly broadened and rounded distally; same in ventral view (Fig.
|
||
<figureCitation id="7FF977329F3C3DCC052EC55383D817A3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
|
||
), with dorsal sclerite forming a slightly sclerotized and short left-sided protuberance and a convex and long right-sided curve, and ventral sclerite narrow and nearly straight (only just scarcely bent to left).
|
||
<emphasis id="1D331EE56F37B185886CA1C264584B83" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="92D74496517DE06409E5C484BCE6B173" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
|
||
). Apical gonocoxite with semi-pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="879195933E73673E23EE74F042821DE2" pageId="0" pageNumber="71" type="habitat">
|
||
<paragraph id="3F28540BE3482C7B71225AD937465F39" pageId="0" pageNumber="71">Habitat.</paragraph>
|
||
<paragraph id="18D8C89258206EF6F6C91EB135AF181C" pageId="0" pageNumber="71">
|
||
Christoph Reuter (CR) and Bernard Lassalle (BL) kindly provided us with information on habitat preferences. The specimens found by CR near Yelkenli were collected in a stunted oak forest. Underground is karst rock, in the middle of the tiny valley flows a small stream that still carries water, at least in spring. The series found by CR at
|
||
<normalizedToken id="546CF792B98690FE0F405F79974968C4" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="884521DFA56783F4BB46DDADFC4AC10C" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
, 1640 m, was collected in a semi-open landscape with a low, sparse oak forest of rather small, thin trees, a lot of scrubland, in between meadows, at least in some places, again karst rock, weathered rocks, and a lot of foliage. BL declare that he found a female specimen at
|
||
<normalizedToken id="5CF3757224EFA6CCC3B79136D02BBFA8" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="13C8F8966CE3DAFC029EFF3B0389B677" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
, 1500 m, in "oak bushes"; same stated that, in Resadiye one male and one female of the new species was collected in small oak grove in a little valley.
|
||
</paragraph>
|
||
<caption id="54CE467E5AE8CA15F4E55F3E84EEF034" doi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71" start="Figure 6" startId="F6">
|
||
<paragraph id="CE094207C439E5C2133EDD9BD33CCE37" pageId="0" pageNumber="71">
|
||
<emphasis id="8A69C19048D98E2D624D84E77FA8888D" bold="true" pageId="0" pageNumber="71">Figure 6.</emphasis>
|
||
Metatarsus, dorsal view.
|
||
<emphasis id="1471D5FAA5A2C5896F9DC5179F25E3A5" bold="true" pageId="0" pageNumber="71">A.</emphasis>
|
||
<taxonomicName id="7C3504C134ED48E6669416B7496019C1" authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus</taxonomicName>
|
||
Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya;
|
||
<emphasis id="B6BFCBB7C02F9BF003468A2B1D48A1E2" bold="true" pageId="0" pageNumber="71">B.</emphasis>
|
||
<taxonomicName id="9362B3BD06036C3361E7578FC9DFFFAC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
|
||
Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel;
|
||
<emphasis id="C790453B8D456A698F7D0C231D297E46" bold="true" pageId="0" pageNumber="71">C.</emphasis>
|
||
<taxonomicName id="EA67C2F2C009FD47E47A58EED937844A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
|
||
, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype;
|
||
<emphasis id="F3CE1103C8C44F009A98CF6D0A495E03" bold="true" pageId="0" pageNumber="71">D.</emphasis>
|
||
<taxonomicName id="E84BE2615700CD7E35B2F872FBA1A809" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
|
||
, sp. nov., right metatarsus, holotype;
|
||
<emphasis id="5ED1E47FEE85005B242F720D4952759D" bold="true" pageId="0" pageNumber="71">E.</emphasis>
|
||
<taxonomicName id="A6B321BDE20056B57ED23DDAF0E4703F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
|
||
(Reiche & Saulcy, 1855), right metatarsus, male specimen, Naẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="06D8F3EB8E8D378363CE20D1B40B19FF" pageId="0" pageNumber="71" type="distribution">
|
||
<paragraph id="B613CC6318CD13892EBB91C9672FE2BB" pageId="0" pageNumber="71">Distribution.</paragraph>
|
||
<paragraph id="C7B6A2546CBB89CB03BFE014222D51CF" pageId="0" pageNumber="71">
|
||
Turkey (provinces of
|
||
<normalizedToken id="ED8A35CFCECF8247B894B7229DE544EA" originalValue="Bingöl">Bingoel</normalizedToken>
|
||
and
|
||
<normalizedToken id="A3CCE0F493FBC90B165EE02AFC6394CF" originalValue="Muş">Mus</normalizedToken>
|
||
:
|
||
<normalizedToken id="A93E430FFACEBA6FFFA3E986A74A5638" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="95F05DE3A091F0D52320F7E1FEA2FAB5" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
; Bitlis Province; Van Province; Fig.
|
||
<figureCitation id="1774B6143AF8EA5CF70F9F8BA833CCF7" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
|
||
). In Bitlis Province, the species inhabits the Nemrut volcanic massif west of Bitlis Town and the mountain region south of the Yelkenli (=
|
||
<normalizedToken id="464BDB39868F44C7938E1FF06347D49E" originalValue="Reşadiye">Resadiye</normalizedToken>
|
||
)
|
||
<normalizedToken id="D69AFFC58562B3FD4ED18A81E9450881" originalValue="Bucaği">Bucagi</normalizedToken>
|
||
. In Van Province,
|
||
<taxonomicName id="3646B7DE8D7CE447CAFA8B40936ADFBA" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
|
||
<emphasis id="DDB1B0123369BA233472A833D672CA1C" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. lives around
|
||
<normalizedToken id="56162E8F15B95DC3FA18E448197F4502" originalValue="Gevaş">Gevas</normalizedToken>
|
||
Town and in the
|
||
<normalizedToken id="8C97556E83064F6EA383470955BEA615" originalValue="Çadir">Cadir</normalizedToken>
|
||
<normalizedToken id="D30F7734E3F6B1F9132A29EBEDAA22AB" originalValue="Daği">Dagi</normalizedToken>
|
||
, a massif situated south of the town aforementioned. In the border area of provinces of
|
||
<normalizedToken id="0C45AC8923A94802D6AC0F00963DE63A" originalValue="Bingöl">Bingoel</normalizedToken>
|
||
and
|
||
<normalizedToken id="40A1609CFE0211A5BB879463C0AC01C4" originalValue="Muş">Mus</normalizedToken>
|
||
, it lives along the pass
|
||
<normalizedToken id="3CE54328108E5A04DDEB476D40E8F233" originalValue="Buğlan">Buglan</normalizedToken>
|
||
<normalizedToken id="D929657808DC6868CAD718E1CF24943F" originalValue="Geçidi">Gecidi</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |