424 lines
47 KiB
XML
424 lines
47 KiB
XML
<document id="FBC0D0B2B2875A348935618B18BAD6C1" ID-DOI="10.11646/zootaxa.3721.2.3" ID-GBIF-Dataset="f3d01271-9143-4021-b5ec-45791021361d" ID-ISSN="1175-5326" ID-Zenodo-Dep="218326" ID-ZooBank="71B4B001-FB10-4B99-ACF9-720131457534" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460688744557" checkinUser="plazi" docAuthor="Salvador, Rodrigo Brincalepe" docDate="2013" docId="5C371C6D271BFFC6FF208E43FD77FD74" docLanguage="en" docName="zt03721p171.pdf" docOrigin="Zootaxa 3721 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cepaea sylvestrina Schlotheim 1820" docType="treatment" docVersion="7" lastPageNumber="165" masterDocId="A00E6415271DFFCEFFB78C28FFCAFF84" masterDocTitle="The fossil pulmonate snails of Sandelzhausen (Early / Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea" masterLastPageNumber="171" masterPageNumber="157" pageNumber="163" updateTime="1698351515074" updateUser="plazi">
|
||
<mods:mods id="29BDA275DB60B2AF66211D32B2B8DBE7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="251E929510C35D4F32F506FD2FDE65A0">
|
||
<mods:title id="A6135943A552FE893606A81C22B4B00B">The fossil pulmonate snails of Sandelzhausen (Early / Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="097FCB0BB7FEEFABF3AC70AE1CD94D79" type="personal">
|
||
<mods:role id="9770A305C185F173AB983CE2759F0F60">
|
||
<mods:roleTerm id="55246118B53165959F7605A8585CD0F2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DCD9C9ECC12E2D92B7B560AC528A94B7">Salvador, Rodrigo Brincalepe</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F50104332C0F05720EB1855DDECD62F5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="87974568265FB127059EA73F17739ABF" type="host">
|
||
<mods:titleInfo id="29AB7E29A85A9BBCC9E397DED01DF3EC">
|
||
<mods:title id="9A627CC19D2197929CC4CCFED3854464">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="80028AA5AE094745C29C19F8E7AE3775">
|
||
<mods:date id="30C79FD2F9E45710B9F7A55987E38A24">2013</mods:date>
|
||
<mods:detail id="8491E565E54916B43E46E630F94D0BA4" type="volume">
|
||
<mods:number id="83BB66E6FA4F3445A514BEA17C109572">3721</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="469880037F47505957E51BAAE03A6ABF" type="issue">
|
||
<mods:number id="8069CAC40550E4878826DC39B2E4F319">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FFD61074B0DD7D2EBDDB23ED8D54908B" unit="page">
|
||
<mods:start id="E62D05CE679397E98CF8A832CD1865A3">157</mods:start>
|
||
<mods:end id="75FE691B60069030908BF88E21453C0F">171</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="20C96F55C942E40FC61C3D46E66960C5">journal article</mods:classification>
|
||
<mods:identifier id="EAE44B8A612BC2EF86FF69C03FD336E5" type="DOI">10.11646/zootaxa.3721.2.3</mods:identifier>
|
||
<mods:identifier id="53C70202DA1948C9A8676A3B3A4776CD" type="GBIF-Dataset">f3d01271-9143-4021-b5ec-45791021361d</mods:identifier>
|
||
<mods:identifier id="B05EA6E6E1440067BE7F66C909398B0B" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="4C1B9E7070D7476519DFCFA69D21411B" type="Zenodo-Dep">218326</mods:identifier>
|
||
<mods:identifier id="B4DA72E8D9CFF720DA158954669B1037" type="ZooBank">71B4B001-FB10-4B99-ACF9-720131457534</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5C371C6D271BFFC6FF208E43FD77FD74" ID-DOI="http://doi.org/10.5281/zenodo.3509193" ID-GBIF-Taxon="119577104" ID-Zenodo-Dep="3509193" LSID="urn:lsid:plazi:treatment:5C371C6D271BFFC6FF208E43FD77FD74" httpUri="http://treatment.plazi.org/id/5C371C6D271BFFC6FF208E43FD77FD74" lastPageId="8" lastPageNumber="165" pageId="6" pageNumber="163">
|
||
<subSubSection id="9C84FEF0271BFFC8FF208E43FDE2FD68" pageId="6" pageNumber="163" type="nomenclature">
|
||
<paragraph id="D421AD7B271BFFC8FF208E43FDB1FD02" blockId="6.[151,635,619,679]" box="[151,635,619,646]" pageId="6" pageNumber="163">
|
||
<heading id="8F691A17271BFFC8FF208E43FDB1FD02" bold="true" box="[151,635,619,646]" fontSize="11" level="1" pageId="6" pageNumber="163" reason="1">
|
||
<taxonomicName id="139ED6F8271BFFC8FF208E43FDB1FD02" ID-CoL="SBY6" authority="Schlotheim, 1820" authorityName="Schlotheim" authorityYear="1820" box="[151,635,619,646]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FF208E43FDB1FD02" bold="true" box="[151,635,619,646]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF208E43FF3BFD01" bold="true" box="[151,241,619,645]" italics="true" pageId="6" pageNumber="163">Cepaea</emphasis>
|
||
cf.
|
||
<emphasis id="E6EA7169271BFFC8FEAA8E43FE54FD01" bold="true" box="[285,414,619,645]" italics="true" pageId="6" pageNumber="163">sylvestrina</emphasis>
|
||
Schlotheim, 1820
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF208EA6FEE7FD23" blockId="6.[151,635,619,679]" box="[151,301,654,679]" pageId="6" pageNumber="163">
|
||
(
|
||
<figureCitation id="4CA5B1FE271BFFC8FF288EA6FEEFFD23" box="[159,293,654,679]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="6" pageNumber="163">Figs. 14–17</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF208EFEFDE2FD68" blockId="6.[151,945,726,902]" box="[151,552,726,748]" pageId="6" pageNumber="163">
|
||
<taxonomicName id="139ED6F8271BFFC8FF208EFEFDEEFD68" authority="Schlotheim, 1820: 99" authorityName="Schlotheim" authorityPageNumber="99" authorityYear="1820" box="[151,548,726,748]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FF208EFEFE88FD68" box="[151,322,726,748]" italics="true" pageId="6" pageNumber="163">Helix sylvestrina</emphasis>
|
||
Schlotheim, 1820: 99
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271BFFC8FF208EDEFD5FFC02" pageId="6" pageNumber="163" type="description">
|
||
<paragraph id="D421AD7B271BFFC8FF208EDEFDC4FCAE" blockId="6.[151,945,726,902]" pageId="6" pageNumber="163">
|
||
<taxonomicName id="139ED6F8271BFFC8FF208EDEFD59FC8F" authority="Wenz, 1923a: 690" authorityName="Wenz" authorityPageNumber="690" authorityYear="1923" box="[151,659,757,779]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="sylvestrina" subSpecies="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FF208EDEFE06FC8F" box="[151,460,757,779]" italics="true" pageId="6" pageNumber="163">Cepaea sylvestrina sylvestrina</emphasis>
|
||
: Wenz, 1923a: 690
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="139ED6F8271BFFC8FF208F3CFDC0FCAE" authority="Truc, 1971: 282" authorityName="Truc" authorityPageNumber="282" authorityYear="1971" box="[151,522,788,810]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FF208F3CFE92FCAE" box="[151,344,788,810]" italics="true" pageId="6" pageNumber="163">Cepaea sylvestrina</emphasis>
|
||
: Truc, 1971: 282
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF208F1BFD4AFCE3" blockId="6.[151,945,726,902]" pageId="6" pageNumber="163">
|
||
<taxonomicName id="139ED6F8271BFFC8FF208F1BFEBBFCCC" box="[151,369,818,840]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="eversa" subSpecies="larteti">
|
||
<emphasis id="E6EA7169271BFFC8FF208F1BFEBBFCCC" box="[151,369,818,840]" italics="true" pageId="6" pageNumber="163">Cepaea eversa larteti</emphasis>
|
||
</taxonomicName>
|
||
(in part): Gall, 1972: 10; Gall, 1973: 10 (pl.1, figs. 4–6).
|
||
<taxonomicName id="139ED6F8271BFFC8FF208F7AFE49FCE3" box="[151,387,849,871]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="silvana" subSpecies="silvana">
|
||
<emphasis id="E6EA7169271BFFC8FF208F7AFE49FCE3" box="[151,387,849,871]" italics="true" pageId="6" pageNumber="163">Cepaea silvana silvana</emphasis>
|
||
</taxonomicName>
|
||
(in part): Gall, 1972: 11.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF208F59FD5FFC02" blockId="6.[151,945,726,902]" box="[151,661,880,902]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF208F59FECAFC02" box="[151,256,881,902]" italics="true" pageId="6" pageNumber="163">
|
||
“
|
||
<taxonomicName id="139ED6F8271BFFC8FF288F59FF33FC02" box="[159,249,881,902]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
sp. 1. (in part): Moser
|
||
<emphasis id="E6EA7169271BFFC8FE5D8F59FDDCFC02" box="[490,534,880,902]" italics="true" pageId="6" pageNumber="163">et al</emphasis>
|
||
., 2009b: 50.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271BFFC8FF208F85FC4FFAA9" pageId="6" pageNumber="163" type="materials_examined">
|
||
<paragraph id="D421AD7B271BFFC8FF208F85FC69FB1A" blockId="6.[151,1436,941,2010]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF208F85FE4BFC42" bold="true" box="[151,385,941,966]" pageId="6" pageNumber="163">Material examined.</emphasis>
|
||
BSPG
|
||
<date id="A0208BBB271BFFC8FE608F85FDE0FC41" box="[471,554,941,966]" pageId="6" pageNumber="163" value="1959-02">1959 II</date>
|
||
459 (1 spcm.), 16156 (1 spcm.), 17447 (2 spcm.), 17448 (2 spcm.), 17447 (1 spcm.), 17449 (1 spcm.), 17450 (1 spcm.), 17451 (1 spcm.), 17452 (1 spcm.), 17453 (1 spcm.), 17454 (1 spcm.), 17455 (1 spcm.), 17456 (2 spcm.), 17457 (1 spcm.), 17458 (1 spcm.), 17459 (1 spcm.), 17460 (1 spcm.), 17461 (1 spcm.), 17462 (1 spcm.), 17463 (1 spcm.), 17464 (1 spcm.), 17465 (1 spcm.), 17466 (1 spcm.), 17467 (1 spcm.), 17468 (1 spcm.), 17766 (1 spcm.), 17767 (1 spcm.), 17768 (1 spcm.), 17769 (1 spcm.), 17770 (1 spcm.), 17771 (1 spcm.), 17772 (1 spcm.), 17773 (1 spcm.), 17774 (1 spcm.), 17775 (1 spcm.), 17776 (2 spcm.), 17777 (1 spcm.), 17778 (1 spcm.), 17779 (3 spcm.), 17780 (2 spcm.), 17781 (2 spcm.).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF708881FC4FFAA9" blockId="6.[151,1436,941,2010]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF708881FE32FB46" bold="true" box="[199,504,1193,1218]" pageId="6" pageNumber="163">Stratigraphic occurrence.</emphasis>
|
||
Layers B1 (1 spcm.), B2 (2 spcm.), C (undetermined; 1 spcm.), C1 (3 spcm.), C3 (1 spcm.), D1 (4 spcm.). Moreover,
|
||
<specimenCount id="C29866F2271BFFC8FD8688E5FD03FB62" box="[561,713,1229,1254]" pageId="6" pageNumber="163" type="generic">25 specimens</specimenCount>
|
||
are from excavation sites for which no profile is available, but, based on their height in the sediment and their preservation, it is possible to infer the layer of origin for some: 3 are likely from layer D, 4 from either B2 or C1, 9 from either C3 or D.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271BFFC8FF708911FA52F9CA" pageId="6" pageNumber="163" type="description">
|
||
<paragraph id="D421AD7B271BFFC8FF708911FF2AF9AE" blockId="6.[151,1436,941,2010]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF708911FE93FAD6" bold="true" box="[199,345,1337,1362]" pageId="6" pageNumber="163">Description.</emphasis>
|
||
Shell medium-sized, ~4 whorls, helicoid; whorls quickly but regularly growing; shell length ~2/3 its width. Protoconch flattened, smooth; transition to teleoconch unclear. Teleoconch smooth, except for growth lines. Suture well-marked. Body whorl slightly bent downwards; faint carina apparently present. Aperture prosocline, crescent-shaped; callus faint. Peristome markedly thickened and reflexed. Umbilicus imperforate. On a single specimen (BSPG
|
||
<date id="A0208BBB271BFFC8FE0789E1FDCFFA65" box="[432,517,1481,1506]" pageId="6" pageNumber="163" value="1959-02">1959 II</date>
|
||
16156) it is possible to observe, under UV light, vestiges of three colored fine parallel spiral bands, the topmost one right below the carina, the other two regularly spaced on basal portion of whorl.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271BFFC8FF708A1DFA52F9CA" blockId="6.[151,1436,941,2010]" box="[199,1432,1589,1614]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF708A1DFE29F9CA" bold="true" box="[199,483,1589,1614]" pageId="6" pageNumber="163">Measurements (in mm).</emphasis>
|
||
BSPG
|
||
<date id="A0208BBB271BFFC8FD8F8A1DFD40F9C9" box="[568,650,1589,1614]" pageId="6" pageNumber="163" value="1959-02">1959 II</date>
|
||
16156 (
|
||
<figureCitation id="4CA5B1FE271BFFC8FD518A1DFCA5F9CA" box="[742,879,1589,1614]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="6" pageNumber="163">Figs. 14–17</figureCitation>
|
||
): 4 whorls; H = 13.7; D = 19.9; h = 6.9; d = 10.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271BFFC8FF708A71FAF8F912" pageId="6" pageNumber="163" type="materials_examined">
|
||
<paragraph id="D421AD7B271BFFC8FF708A71FAF8F912" blockId="6.[151,1436,941,2010]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF708A71FD6AF9F6" bold="true" box="[199,672,1625,1650]" pageId="6" pageNumber="163">Previous identification of the material.</emphasis>
|
||
Gall (1972: Nr. 20 and 21, in part): respectively,
|
||
<taxonomicName id="139ED6F8271BFFC8FB5C8A72FE8AF912" authority="Klein" authorityName="Klein" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="silvana" subSpecies="silvana">
|
||
<emphasis id="E6EA7169271BFFC8FB5C8A72FF22F912" italics="true" pageId="6" pageNumber="163">Cepaea silvana silvana</emphasis>
|
||
(Klein)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="139ED6F8271BFFC8FEC18A56FD1AF912" authority="Boissy" authorityName="Boissy" box="[374,720,1661,1686]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="eversa" subSpecies="larteti">
|
||
<emphasis id="E6EA7169271BFFC8FEC18A56FDADF912" box="[374,615,1661,1686]" italics="true" pageId="6" pageNumber="163">Cepaea eversa larteti</emphasis>
|
||
(Boissy)
|
||
</taxonomicName>
|
||
. Moser
|
||
<emphasis id="E6EA7169271BFFC8FC9E8A56FC90F912" box="[809,858,1661,1686]" italics="true" pageId="6" pageNumber="163">et al</emphasis>
|
||
. (2009b: Nr. 64, in part):
|
||
<emphasis id="E6EA7169271BFFC8FBCC8A56FB25F911" box="[1147,1263,1662,1685]" italics="true" pageId="6" pageNumber="163">
|
||
“
|
||
<taxonomicName id="139ED6F8271BFFC8FB348A56FB2DF911" box="[1155,1255,1662,1685]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
sp. 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271BFFC9FF708A89FDCFFEC4" lastPageId="7" lastPageNumber="164" pageId="6" pageNumber="163" type="discussion">
|
||
<paragraph id="D421AD7B271BFFC8FF708A89FCF1F85D" blockId="6.[151,1436,941,2010]" pageId="6" pageNumber="163">
|
||
<emphasis id="E6EA7169271BFFC8FF708A89FE86F93E" bold="true" box="[199,332,1697,1722]" pageId="6" pageNumber="163">Discussion.</emphasis>
|
||
The specimens from Sandelzhausen compare well with
|
||
<taxonomicName id="139ED6F8271BFFC8FC098A8AFB9DF93E" box="[958,1111,1697,1722]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FC098A8AFB9DF93E" box="[958,1111,1697,1722]" italics="true" pageId="6" pageNumber="163">C. sylvestrina</emphasis>
|
||
</taxonomicName>
|
||
, but the typical specimens of the latter are taller and have a rounder profile, especially the body whorl. Moreover, typical
|
||
<taxonomicName id="139ED6F8271BFFC8FB2A8AEEFAFCF95A" box="[1181,1334,1733,1758]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FB2A8AEEFAFCF95A" box="[1181,1334,1733,1758]" italics="true" pageId="6" pageNumber="163">C. sylvestrina</emphasis>
|
||
</taxonomicName>
|
||
have the apertural region bent more markedly downwards. Finally, the present specimens have a more distinct carina and less convex whorls. However, it cannot be completely excluded that these are artifacts due to preservational biases (for more, see the Discussion chapter below). The vestiges of colored bands can be seen in a single specimen of
|
||
<taxonomicName id="139ED6F8271BFFC8FA358B1AFEFDF8EA" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FA358B1AFA56F8CD" box="[1410,1436,1842,1865]" italics="true" pageId="6" pageNumber="163">C.</emphasis>
|
||
cf.
|
||
<emphasis id="E6EA7169271BFFC8FF0A8B7DFEFDF8EA" box="[189,311,1877,1902]" italics="true" pageId="6" pageNumber="163">sylvestrina</emphasis>
|
||
</taxonomicName>
|
||
and two of
|
||
<taxonomicName id="139ED6F8271BFFC8FE708B7EFD90F8E9" box="[455,602,1877,1902]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
|
||
<emphasis id="E6EA7169271BFFC8FE708B7EFE2BF8E9" box="[455,481,1878,1901]" italics="true" pageId="6" pageNumber="163">C.</emphasis>
|
||
cf.
|
||
<emphasis id="E6EA7169271BFFC8FDA68B7EFD90F8E9" box="[529,602,1878,1901]" italics="true" pageId="6" pageNumber="163">eversa</emphasis>
|
||
</taxonomicName>
|
||
(see above). The pattern is trifasciate in both species, but the bands are much broader in
|
||
<taxonomicName id="139ED6F8271BFFC8FEE48B52FE15F815" box="[339,479,1913,1938]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
|
||
<emphasis id="E6EA7169271BFFC8FEE48B52FEACF815" box="[339,358,1914,1937]" italics="true" pageId="6" pageNumber="163">C</emphasis>
|
||
. cf.
|
||
<emphasis id="E6EA7169271BFFC8FE218B52FE15F815" box="[406,479,1914,1937]" italics="true" pageId="6" pageNumber="163">eversa</emphasis>
|
||
</taxonomicName>
|
||
. This trifasciate pattern occurs in recent
|
||
<taxonomicName id="139ED6F8271BFFC8FC138B52FC33F815" box="[932,1017,1914,1937]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E6EA7169271BFFC8FC138B52FC33F815" box="[932,1017,1914,1937]" italics="true" pageId="6" pageNumber="163">Cepaea</emphasis>
|
||
</taxonomicName>
|
||
species and was found to be the most common one in fossils of
|
||
<taxonomicName id="139ED6F8271BFFC8FE0E8BB6FCD0F831" authority="Wenz" authorityName="Wenz" box="[441,794,1949,1974]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="sylvestrina" subSpecies="gottschicki">
|
||
<emphasis id="E6EA7169271BFFC8FE0E8BB6FD19F832" box="[441,723,1949,1974]" italics="true" pageId="6" pageNumber="163">C. sylvestrina gottschicki</emphasis>
|
||
Wenz
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="AC89EDEB271BFFC8FCE98BB5FC66F832" box="[862,940,1949,1974]" name="Poland" pageId="6" pageNumber="163">Poland</collectingCountry>
|
||
(Górka 2008).
|
||
<taxonomicName id="139ED6F8271BFFC8FBEE8BB6FAE4F832" box="[1113,1326,1949,1974]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271BFFC8FBEE8BB6FAE4F832" box="[1113,1326,1949,1974]" italics="true" pageId="6" pageNumber="163">Cepaea sylvestrina</emphasis>
|
||
</taxonomicName>
|
||
is known throughout the Middle and Late Miocene of Central Europe.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF708CBFFC3BFF50" blockId="7.[151,1436,151,213]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF708CBFFEF1FF34" bold="true" box="[199,315,151,176]" pageId="7" pageNumber="164">Remarks.</emphasis>
|
||
Truc (1971) alludes to the possibility that
|
||
<taxonomicName id="139ED6F8271AFFC9FCA88CB0FC73FF34" box="[799,953,151,176]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="sylvestrina">
|
||
<emphasis id="E6EA7169271AFFC9FCA88CB0FC73FF34" box="[799,953,151,176]" italics="true" pageId="7" pageNumber="164">C. sylvestrina</emphasis>
|
||
</taxonomicName>
|
||
belongs in the genus
|
||
<emphasis id="E6EA7169271AFFC9FB058CBFFA98FF34" box="[1202,1362,151,176]" italics="true" pageId="7" pageNumber="164">Megalotachea</emphasis>
|
||
. Zilch (
|
||
<date id="A0208BBB271AFFC9FF288C94FEE8FF51" box="[159,290,188,213]" pageId="7" pageNumber="164" value="1959" valueMax="1960">1959–1960</date>
|
||
), however, considers
|
||
<emphasis id="E6EA7169271AFFC9FDA38C94FD7EFF51" box="[532,692,188,213]" italics="true" pageId="7" pageNumber="164">Megalotachea</emphasis>
|
||
a synonym of
|
||
<taxonomicName id="139ED6F8271AFFC9FCED8C95FC7AFF50" box="[858,944,189,212]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="genus" sensu="stricto">
|
||
<emphasis id="E6EA7169271AFFC9FCED8C95FC7AFF50" box="[858,944,189,212]" italics="true" pageId="7" pageNumber="164">Cepaea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9CC12271AFFC9FC018C95FC3BFF50" box="[950,1009,189,212]" pageId="7" pageNumber="164" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208D0FFDCFFEC4" blockId="7.[151,517,295,320]" box="[151,517,295,320]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208D0FFDCFFEC4" bold="true" box="[151,517,295,320]" pageId="7" pageNumber="164">
|
||
Genus
|
||
<emphasis id="E6EA7169271AFFC9FF5D8D0FFE62FEC4" bold="true" box="[234,424,295,320]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
Pilsbry
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271AFFC9FF208D47FC87FC82" pageId="7" pageNumber="164" type="description">
|
||
<paragraph id="D421AD7B271AFFC9FF208D47FEE7FE28" blockId="7.[151,676,367,429]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208D47FD6EFE0C" bold="true" box="[151,676,367,392]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208D47FE9FFE0C" bold="true" box="[151,341,367,392]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
<taxonomicName id="139ED6F8271AFFC9FEEB8D47FD6EFE0C" authority="Klein, 1853" authorityName="Klein" authorityYear="1853" box="[348,676,367,392]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassatus">
|
||
cf.
|
||
<emphasis id="E6EA7169271AFFC9FE378D47FDC9FE0C" bold="true" box="[384,515,367,392]" italics="true" pageId="7" pageNumber="164">incrassatus</emphasis>
|
||
(Klein, 1853)
|
||
</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="4CA5B1FE271AFFC9FF288DBCFEECFE28" box="[159,294,404,429]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="7" pageNumber="164">Figs. 18–21</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208DF4FD9CFE76" blockId="7.[151,1436,476,774]" box="[151,598,476,498]" pageId="7" pageNumber="164">
|
||
<taxonomicName id="139ED6F8271AFFC9FF208DF4FE77FE76" authority="Klein, 1847: 71" authorityName="Klein" authorityPageNumber="71" authorityYear="1847" box="[151,445,476,498]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="inflexa">
|
||
<emphasis id="E6EA7169271AFFC9FF208DF4FED3FE76" box="[151,281,476,498]" italics="true" pageId="7" pageNumber="164">Helix inflexa</emphasis>
|
||
Klein, 1847: 71
|
||
</taxonomicName>
|
||
(pl. 1, fig. 12).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208DD2FDB1FD94" blockId="7.[151,1436,476,774]" box="[151,635,506,528]" pageId="7" pageNumber="164">
|
||
<taxonomicName id="139ED6F8271AFFC9FF208DD2FE25FD94" authority="Klein, 1853: 208" authorityName="Klein" authorityPageNumber="208" authorityYear="1853" box="[151,495,506,528]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassata">
|
||
<emphasis id="E6EA7169271AFFC9FF208DD2FEF4FD94" box="[151,318,506,528]" italics="true" pageId="7" pageNumber="164">Helix incrassata</emphasis>
|
||
Klein, 1853: 208
|
||
</taxonomicName>
|
||
(pl. 5, fig. 6).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208E31FBC0FDAB" blockId="7.[151,1436,476,774]" box="[151,1034,537,559]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208E31FD98FDAB" box="[151,594,537,559]" italics="true" pageId="7" pageNumber="164">Tropidomphalus (Pseudochloritis) extinctus</emphasis>
|
||
: Gall, 1972: 9; Gall, 1973: 10 (pl.1, fig. 1).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208E10FF3AFDE8" blockId="7.[151,1436,476,774]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208E10FD15FDCA" box="[151,735,568,590]" italics="true" pageId="7" pageNumber="164">
|
||
Tropidomphalus (Pseudochloritis) incrasstus
|
||
<taxonomicName id="139ED6F8271AFFC9FDDB8E11FD15FDCA" authority="Wenz, 1923a: 510" authorityName="Wenz" authorityPageNumber="510" authorityYear="1923" box="[620,735,569,590]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassatus">incrassatus</taxonomicName>
|
||
</emphasis>
|
||
: Wenz, 1923a: 510; Gall, 1972: 9; Schlickum, 1976: 16 (pl. 4, fig. 56).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208E5DFD71FD0F" blockId="7.[151,1436,476,774]" box="[151,699,629,651]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208E5DFDE8FD0F" box="[151,546,629,651]" italics="true" pageId="7" pageNumber="164">
|
||
Tropidomphalus
|
||
<taxonomicName id="139ED6F8271AFFC9FEF58E5DFDE8FD0F" authority="Gall, 1972: 9" authorityName="Gall" authorityPageNumber="9" authorityYear="1972" box="[322,546,629,651]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="zelli" subGenus="Pseudochloritis">(Pseudochloritis) zelli</taxonomicName>
|
||
</emphasis>
|
||
: Gall, 1972: 9.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208EBCFC92FD2E" blockId="7.[151,1436,476,774]" box="[151,856,660,682]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208EBCFEF6FD2E" box="[151,316,660,682]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
? sp.: Gall, 1972: 10; Gall, 1973: 10 (pl.1, figs. 2–3).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208E9AFC3DFD4C" blockId="7.[151,1436,476,774]" box="[151,1015,690,712]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208E9AFDA1FD4C" box="[151,619,690,712]" italics="true" pageId="7" pageNumber="164">
|
||
Tropidomphalus
|
||
<taxonomicName id="139ED6F8271AFFC9FEF58E9AFDA1FD4C" authority="Kokay, 2006: 90" authorityName="Kokay" authorityPageNumber="90" authorityYear="2006" box="[322,619,690,712]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassatus" subGenus="Pseudochloritis">(Pseudochloritis) incrassatus</taxonomicName>
|
||
</emphasis>
|
||
: Kókay, 2006: 90 (pl. 34, figs. 12–14).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208EF9FC5EFD63" blockId="7.[151,1436,476,774]" box="[151,916,721,743]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208EF9FE6FFD63" box="[151,421,721,743]" italics="true" pageId="7" pageNumber="164">
|
||
Pseudochloritis
|
||
<taxonomicName id="139ED6F8271AFFC9FE8B8EFAFE6FFD63" authority="Binder, 2008: 172" authorityName="Binder" authorityPageNumber="172" authorityYear="2008" box="[316,421,722,743]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassata">incrassata</taxonomicName>
|
||
</emphasis>
|
||
: Binder, 2008: 172 (pl. 3, figs. 2–4, pl. 6, fig. 2).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF208ED8FC87FC82" blockId="7.[151,1436,476,774]" box="[151,845,752,774]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208ED8FE3FFC82" box="[151,501,752,774]" italics="true" pageId="7" pageNumber="164">Tropidomphalus (Pseudochlorites)</emphasis>
|
||
[sic] sp.: Moser
|
||
<emphasis id="E6EA7169271AFFC9FD168ED9FD04FC82" box="[673,718,752,774]" italics="true" pageId="7" pageNumber="164">et al</emphasis>
|
||
., 2009b: 49.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271AFFC9FF208F1AFA47FA57" pageId="7" pageNumber="164" type="materials_examined">
|
||
<paragraph id="D421AD7B271AFFC9FF208F1AFBA2FB7F" blockId="7.[151,1437,818,2031]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF208F1AFE4EFCCF" bold="true" box="[151,388,818,843]" pageId="7" pageNumber="164">Material examined.</emphasis>
|
||
BSPG
|
||
<date id="A0208BBB271AFFC9FE568F1AFDFDFCCE" box="[481,567,818,843]" pageId="7" pageNumber="164" value="1959-02">1959 II</date>
|
||
456 (1 spcm.), 457 (1 spcm.), 458 (1 spcm.), 16153 (1 spcm.), 16154 (1 spcm.), 16155 (4 spcm.), 16163 (1 spcm.), 17318 (1 spcm.), 17322 (1 spcm.), 17323 (1 spcm.), 17324 (1 spcm.), 17325 (1 spcm.), 17326 (1 spcm.), 17327 (2 spcm.), 17328 (1 spcm.), 17329 (1 spcm.), 17330 (1 spcm.), 17331 (1 spcm.), 17332 (1 spcm.), 17333 (1 spcm.), 17334 (1 spcm.), 17335 (1 spcm.), 17348 (1 spcm.), 17349 (1 spcm.), 17350 (1 spcm.), 17351 (1 spcm.), 17353 (1 spcm.), 17354 (1 spcm.), 17355 (1 spcm.), 17356 (1 spcm.), 17357 (1 spcm.), 17358 (2 spcm.), 17359 (4 spcm.), 17360 (1 spcm.), 17361 (2 spcm.), 17362 (1 spcm.), 17363 (1 spcm.), 17364 (4 spcm.), 17365 (1 spcm.), 17366 (2 spcm.), 17367 (1 spcm.), 17368 (1 spcm.), 17369 (1 spcm.), 17370 (1 spcm.), 17473 (1 spcm.), 17730 (1 spcm.), 17731 (1 spcm.), 17732 (1 spcm.), 17733 (2 spcm.), 17734 (2 spcm.), 17735 (1 spcm.), 17736 (1 spcm.), 17737 (1 spcm.), 17738 (4 spcm.), 17739 (1 spcm.), 17740 (1 spcm.), 17741 (2 spcm.), 17742 (1 spcm.), 17743 (1 spcm.), 17744 (1 spcm.), 17745 (1 spcm.), 17746 (2 spcm.), 17747 (1 spcm.), 17748 (1 spcm.), 17749 (1 spcm.), 17750 (1 spcm.), 17751 (3 spcm.), 17752 (1 spcm.), 17753 (1 spcm.), 17754 (1 spcm.), 17755 (1 spcm.), 17756 (2 spcm.), 17757 (4 spcm.), 17758 (1 spcm.), 17759 (1 spcm.), 17760 (1 spcm.), 17761 (1 spcm.), 17762 (1 spcm.), 17763 (4 spcm.), 17764 (3 spcm.), 17765 (1 spcm.).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF70892EFA47FA57" blockId="7.[151,1437,818,2031]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF70892EFE3DFA9B" bold="true" box="[199,503,1286,1311]" pageId="7" pageNumber="164">Stratigraphic occurrence.</emphasis>
|
||
Layers B (undetermined; 10 spcm.), B1 (3 spcm.), B2 (2 spcm.), C (undetermined; 5 spcm.), C1 (7 spcm.), C2 (2 spcm.), C3 (2 spcm.) and D1 (4 spcm.). Moreover,
|
||
<specimenCount id="C29866F2271AFFC9FBA88902FB7DFAC7" box="[1055,1207,1322,1347]" pageId="7" pageNumber="164" type="generic">68 specimens</specimenCount>
|
||
are from excavation sites for which no profile is available, but, based on their height in the sediment and their preservation, it is possible to infer the layer of origin for some: 1 is likely from layer B (undetermined), 1 from C (undetermined), 3 from C1, 2 from C3, 8 from D1, 13 from either B or C, 9 from either B2 or C1, and 10 from either C3 or D1. Finally,
|
||
<specimenCount id="C29866F2271AFFC9FA3989BFFEC6FA57" pageId="7" pageNumber="164" type="generic">4 specimens</specimenCount>
|
||
stem from the Grube Bergmaier site and
|
||
<specimenCount id="C29866F2271AFFC9FD6B8993FCAFFA57" box="[732,869,1466,1491]" pageId="7" pageNumber="164" type="generic">2 specimens</specimenCount>
|
||
completely lack locality and stratigraphical data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271AFFC9FF7089F6FAF5F977" pageId="7" pageNumber="164" type="description">
|
||
<paragraph id="D421AD7B271AFFC9FF7089F6FE9BF94B" blockId="7.[151,1437,818,2031]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF7089F6FE93FA73" bold="true" box="[199,345,1502,1527]" pageId="7" pageNumber="164">Description.</emphasis>
|
||
Shell large, 4½ whorls, helicoid to disc-shaped, with flattened spire; shell length ~1/2 its width. Protoconch (1¼ whorl) flattened, large in relation to following whorl; sculptured by fine striae dotted with weak papillae; transition to teleoconch unclear. Teleoconch sculptured by thickened growth lines and irregular weak furrows, with regularly arranged papillae. Suture deep, well-marked. Body whorl enlarged, slightly bent downwards, with conspicuous constriction right before the aperture (“extralabial depression”
|
||
<emphasis id="E6EA7169271AFFC9FB0E8A47FB3DF902" box="[1209,1271,1647,1670]" italics="true" pageId="7" pageNumber="164">sensu</emphasis>
|
||
Binder 2008). Aperture prosocline; no callus apparent. Peristome markedly thickened and reflexed, slightly covering umbilicus. Umbilicus wide.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B271AFFC9FF708AF2FAF5F977" blockId="7.[151,1437,818,2031]" box="[199,1343,1754,1779]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF708AF2FE29F977" bold="true" box="[199,483,1754,1779]" pageId="7" pageNumber="164">Measurements (in mm).</emphasis>
|
||
BSPG
|
||
<date id="A0208BBB271AFFC9FD8F8AF2FD40F976" box="[568,650,1754,1779]" pageId="7" pageNumber="164" value="1959-02">1959 II</date>
|
||
456 (incomplete specimen;
|
||
<figureCitation id="4CA5B1FE271AFFC9FC718AF2FB85F977" box="[966,1103,1754,1779]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="7" pageNumber="164">Figs. 18–19</figureCitation>
|
||
): H = 15.9; D = 28.8.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271AFFC9FF708AD6FE74F806" pageId="7" pageNumber="164" type="materials_examined">
|
||
<paragraph id="D421AD7B271AFFC9FF708AD6FE74F806" blockId="7.[151,1437,818,2031]" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF708AD6FD1CF893" bold="true" box="[199,726,1790,1815]" pageId="7" pageNumber="164">Previous identification of the material.</emphasis>
|
||
Gall (1972: Nr. 13–16): respectively,
|
||
<emphasis id="E6EA7169271AFFC9FB528AD6FE1FF8BF" italics="true" pageId="7" pageNumber="164">Tropidomphalus (Pseudochloritis) extinctus</emphasis>
|
||
(Rambur),
|
||
<emphasis id="E6EA7169271AFFC9FDCE8B0AFB5FF8BF" box="[633,1173,1826,1851]" italics="true" pageId="7" pageNumber="164">Tropidomphalus (Pseudochloritis) incrasstus</emphasis>
|
||
<taxonomicName id="139ED6F8271AFFC9FB1B8B0CFA5DF8BF" authority="(Klein)" authorityName="Klein" baseAuthorityName="Klein" box="[1196,1431,1826,1851]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassatus">
|
||
<emphasis id="E6EA7169271AFFC9FB1B8B0CFAE6F8BF" box="[1196,1324,1828,1851]" italics="true" pageId="7" pageNumber="164">incrassatus</emphasis>
|
||
(Klein)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E6EA7169271AFFC9FF208B6EFE84F8DB" box="[151,334,1862,1887]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
<taxonomicName id="139ED6F8271AFFC9FEE38B6EFD56F8DB" authority="(Kurr)" authorityName="Kurr" baseAuthorityName="Kurr" box="[340,668,1862,1887]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="zelli" subGenus="Pseudochloritis">
|
||
<emphasis id="E6EA7169271AFFC9FEE38B6EFD86F8DB" box="[340,588,1862,1887]" italics="true" pageId="7" pageNumber="164">(Pseudochloritis) zelli</emphasis>
|
||
(Kurr)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E6EA7169271AFFC9FD108B6EFC97F8DB" box="[679,861,1862,1887]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
? sp. Moser
|
||
<emphasis id="E6EA7169271AFFC9FC5B8B6FFBD6F8DB" box="[1004,1052,1862,1887]" italics="true" pageId="7" pageNumber="164">et al</emphasis>
|
||
. (2009b: Nr. 56):
|
||
<emphasis id="E6EA7169271AFFC9FB528B6EFE97F807" italics="true" pageId="7" pageNumber="164">Tropidomphalus (Pseudochlorites)</emphasis>
|
||
[sic] sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84FEF0271AFFC6FF708BA6FD77FD74" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164" type="discussion">
|
||
<paragraph id="D421AD7B271AFFC6FF708BA6FE0DFF7C" blockId="7.[151,1437,818,2031]" lastBlockId="8.[151,1437,151,752]" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164">
|
||
<emphasis id="E6EA7169271AFFC9FF708BA6FE86F823" bold="true" box="[199,332,1934,1959]" pageId="7" pageNumber="164">Discussion.</emphasis>
|
||
The specimens from Sandelzhausen, despite their poor preservation, seem to conform well to the great morphological variation shown by
|
||
<emphasis id="E6EA7169271AFFC9FDDF8B9AFCD5F84F" box="[616,799,1970,1995]" italics="true" pageId="7" pageNumber="164">Tropidomphalus</emphasis>
|
||
<taxonomicName id="139ED6F8271AFFC9FC908B9CFB99F84F" authority="(Binder 2008)" authorityName="Binder" authorityYear="2008" baseAuthorityName="Binder" baseAuthorityYear="2008" box="[807,1107,1970,1995]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="164" phylum="Mollusca" rank="species" species="incrassatus">
|
||
<emphasis id="E6EA7169271AFFC9FC908B9CFC6DF84F" box="[807,935,1972,1995]" italics="true" pageId="7" pageNumber="164">incrassatus</emphasis>
|
||
(Binder 2008)
|
||
</taxonomicName>
|
||
. This species is known from the Middle Miocene of
|
||
<collectingCountry id="AC89EDEB271AFFC9FE178BFEFE24F86B" box="[416,494,2006,2031]" name="Poland" pageId="7" pageNumber="164">Poland</collectingCountry>
|
||
(Górka 2008),
|
||
<collectingCountry id="AC89EDEB271AFFC9FD2B8BFEFD3BF86B" box="[668,753,2006,2031]" name="Austria" pageId="7" pageNumber="164">Austria</collectingCountry>
|
||
, many localities in
|
||
<collectingCountry id="AC89EDEB271AFFC9FC7C8BFFFBF9F86B" box="[971,1075,2007,2031]" name="Germany" pageId="7" pageNumber="164">Germany</collectingCountry>
|
||
(Binder 2008) and perhaps also
|
||
<collectingCountry id="AC89EDEB2715FFC6FF208CB0FF33FF34" box="[151,249,152,176]" name="Hungary" pageId="8" pageNumber="165">Hungary</collectingCountry>
|
||
(Kókay 2006). Some of the specimens are poorly preserved, with varying degrees of deformation, which alters the overall shell shape and its size and may influence classification as different species (for more, see the Discussion chapter below).
|
||
</paragraph>
|
||
<paragraph id="D421AD7B2715FFC6FF708D2CFE24FE0C" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="165">
|
||
Binder (2008) considers the shell of
|
||
<emphasis id="E6EA71692715FFC6FDEA8D2CFCDEFE99" box="[605,788,260,285]" italics="true" pageId="8" pageNumber="165">Tropidomphalus</emphasis>
|
||
, based especially on the angulation of the body whorl and aperture, an adaptation for ground-dwelling to reduce water loss in drier environments. Moser
|
||
<emphasis id="E6EA71692715FFC6FB0F8D01FB22FEC4" box="[1208,1256,295,320]" italics="true" pageId="8" pageNumber="165">et al</emphasis>
|
||
. (2009) suggest that the genus inhabited drier and more open habitats, having a “way of life like strong-shelled Balkanese representatives of
|
||
<taxonomicName id="139ED6F82715FFC6FED38D47FE2AFE0C" box="[356,480,367,392]" class="Gastropoda" family="Helicidae" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="165" phylum="Mollusca" rank="subFamily" subFamily="Ariantinae">Ariantinae</taxonomicName>
|
||
”.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B2715FFC6FF708DBCFCF4FD00" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="165">
|
||
<emphasis id="E6EA71692715FFC6FF708DBCFEF1FE29" bold="true" box="[199,315,404,429]" pageId="8" pageNumber="165">Remarks.</emphasis>
|
||
<taxonomicName id="139ED6F82715FFC6FEF38DBDFE2BFE28" box="[324,481,405,428]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="165" phylum="Mollusca" rank="species" species="incrassatus">
|
||
<emphasis id="E6EA71692715FFC6FEF38DBDFE2BFE28" box="[324,481,405,428]" italics="true" pageId="8" pageNumber="165">T. incrassatus</emphasis>
|
||
</taxonomicName>
|
||
is considered to belong to the subgenus
|
||
<emphasis id="E6EA71692715FFC6FC0F8DBCFBA2FE29" box="[952,1128,404,429]" italics="true" pageId="8" pageNumber="165">Pseudochloritis</emphasis>
|
||
Boettger, 1909. Nordsieck (1986) was the first to regard
|
||
<emphasis id="E6EA71692715FFC6FE548D9FFD59FE54" box="[483,659,439,464]" italics="true" pageId="8" pageNumber="165">Pseudochloritis</emphasis>
|
||
as a distinct genus, but did not offer any explanation for his decision. Binder (2008) revised the genus and separated it from
|
||
<emphasis id="E6EA71692715FFC6FC988DF4FC2FFE71" box="[815,997,476,501]" italics="true" pageId="8" pageNumber="165">Tropidomphalus</emphasis>
|
||
basically based on the width of the umbilicus (narrow in
|
||
<emphasis id="E6EA71692715FFC6FE278DD7FD8AFD9C" box="[400,576,511,536]" italics="true" pageId="8" pageNumber="165">Pseudochloritis</emphasis>
|
||
, wide in
|
||
<emphasis id="E6EA71692715FFC6FD188DD7FCAFFD9C" box="[687,869,511,536]" italics="true" pageId="8" pageNumber="165">Tropidomphalus</emphasis>
|
||
) and protoconch sculpture. However, protoconch sculpture does not seem to be constant in each subgenera and the width of the umbilicus greatly varies even intraspecifically for specimens of different localities.
|
||
<collectingCountry id="AC89EDEB2715FFC6FD458E60FCDBFDE4" box="[754,785,584,608]" name="American Samoa" pageId="8" pageNumber="165">As</collectingCountry>
|
||
such, here we maintain the species only as
|
||
<taxonomicName id="139ED6F82715FFC6FAB78E61FA51FDE4" box="[1280,1435,585,608]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="165" phylum="Mollusca" rank="species" species="incrassatus">
|
||
<emphasis id="E6EA71692715FFC6FAB78E61FA51FDE4" box="[1280,1435,585,608]" italics="true" pageId="8" pageNumber="165">T. incrassatus</emphasis>
|
||
</taxonomicName>
|
||
and question the usefulness of the subgenus
|
||
<emphasis id="E6EA71692715FFC6FD3F8E44FCF2FD01" box="[648,824,620,645]" italics="true" pageId="8" pageNumber="165">Pseudochloritis</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D421AD7B2715FFC6FF708EA7FD77FD74" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="165">
|
||
Finally, there is some uncertainty as to the position of the genus
|
||
<emphasis id="E6EA71692715FFC6FC018EA7FBA7FD2C" box="[950,1133,655,680]" italics="true" pageId="8" pageNumber="165">Tropidomphalus</emphasis>
|
||
Pilsbry inside Helicoidea, belonging either to the
|
||
<taxonomicName id="139ED6F82715FFC6FE2A8E9CFDCBFD49" box="[413,513,692,717]" class="Gastropoda" family="Elonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="165" phylum="Mollusca" rank="family">Elonidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="139ED6F82715FFC6FD908E9CFD58FD49" box="[551,658,692,717]" class="Gastropoda" family="Helicidae" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="165" phylum="Mollusca" rank="family">Helicidae</taxonomicName>
|
||
(Zilch
|
||
<date id="A0208BBB2715FFC6FD528E9CFCADFD49" box="[741,871,692,717]" pageId="8" pageNumber="165" value="1959" valueMax="1960">1959–1960</date>
|
||
; Binder 2008; Moser
|
||
<emphasis id="E6EA71692715FFC6FBEA8E9DFB44FD49" box="[1117,1166,692,717]" italics="true" pageId="8" pageNumber="165">et al</emphasis>
|
||
. 2009b). Here the latter was preferred, in accordance with Binder (2008).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |