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<mods:title id="FBA0563512299E46A25EFF1E3DDC47E2">Revision of the Neotropical genus Cheiromyia Dyte (Diptera: Dolichopodidae)</mods:title>
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<mods:namePart id="F99F7D95FC8FA2DA9C5F4C1349CBCC1C">Cumming, Jeffrey M.</mods:namePart>
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<treatment id="48372857FFAFFFBBDAD5D4A5789CDDCC" ID-DOI="http://doi.org/10.5281/zenodo.6202521" ID-GBIF-Taxon="124955422" ID-Zenodo-Dep="6202521" LSID="urn:lsid:plazi:treatment:48372857FFAFFFBBDAD5D4A5789CDDCC" httpUri="http://treatment.plazi.org/id/48372857FFAFFFBBDAD5D4A5789CDDCC" lastPageId="4" lastPageNumber="45" pageId="1" pageNumber="42">
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<paragraph id="C0219941FFAFFFBEDAD5D4A578ABDC93" blockId="1.[151,445,1698,1724]" box="[151,445,1698,1724]" pageId="1" pageNumber="42">
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<emphasis id="F2EA4553FFAFFFBEDAD5D4A578ABDC93" bold="true" box="[151,445,1698,1724]" pageId="1" pageNumber="42">
Genus
<taxonomicName id="079EE2C2FFAFFFBEDAACD4A578ABDC93" ID-CoL="3MKS" authority="Dyte" authorityName="Dyte" box="[238,445,1698,1724]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAFFFBEDAACD4A5786ADC93" bold="true" box="[238,380,1698,1724]" italics="true" pageId="1" pageNumber="42">Cheiromyia</emphasis>
Dyte
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8884CACAFFAFFFBDDAD5D4EC7C96DAF7" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42" type="reference_group">
<paragraph id="C0219941FFAFFFBEDAD5D4EC7B86DD0D" blockId="1.[151,1436,1771,1890]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="E08EBE6FFFAFFFBEDAD5D4EC7B86DD0D" pageId="1" pageNumber="42">
<taxonomicName id="079EE2C2FFAFFFBEDAD5D4EC78C5DD2D" ID-CoL="3MJY" authority="Parent, 1930: 13" authorityName="Parent" authorityPageNumber="13" authorityYear="1930" box="[151,467,1771,1794]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="F2EA4553FFAFFFBEDAD5D4EC780CDD2D" box="[151,282,1771,1794]" italics="true" pageId="1" pageNumber="42">Cheirocerus</emphasis>
<treatmentCitation id="413FBF50FFAFFFBEDB62D4EC78C5DD2D" author="Parent" box="[288,467,1771,1794]" page="13" pageId="1" pageNumber="42" year="1930">
<bibRefCitation id="A40FE4B0FFAFFFBEDB62D4EC78BDDD2D" author="Parent" box="[288,427,1771,1794]" pageId="1" pageNumber="58" refString="Parent, O. (1930) Ergebnisse einer zoologischen Sammelreise nach Brasilien, insbesondere in das Amazonasgebiet, ausgefuhrt von Dr. H. Zerny III. Teil. Diptera: Dolichopodidae. Annalen des Naturhistorischen Museums in Wien, 44, 5 - 26." type="journal article" year="1930">Parent, 1930</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="1F2527E3FFAFFFBEDB9FD4EC7B04DD2D" box="[477,530,1771,1794]" pageId="1" pageNumber="42">Type</typeStatus>
species:
<taxonomicName id="079EE2C2FFAFFFBED836D4EC7ACADD2D" authority="Parent" authorityName="Parent" box="[628,988,1771,1794]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="42" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFAFFFBED836D4EC7A87DD2D" box="[628,913,1771,1794]" italics="true" pageId="1" pageNumber="42">Cheirocerus palmaticornis</emphasis>
Parent
</taxonomicName>
[Neotropical], by monotypy. Preoccupied by
<taxonomicName id="079EE2C2FFAFFFBEDAAAD50C7B3BDD0D" ID-CoL="3MJY" authority="Eigenmann, 1917" authorityName="Eigenmann" authorityYear="1917" box="[232,557,1803,1826]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="F2EA4553FFAFFFBEDAAAD50C787DDD0D" box="[232,363,1803,1826]" italics="true" pageId="1" pageNumber="42">Cheirocerus</emphasis>
<bibRefCitation id="A40FE4B0FFAFFFBEDB30D50C7B3BDD0D" author="Eigenmann" box="[370,557,1803,1826]" pageId="1" pageNumber="58" refString="Eigenmann, C. H. (1917) New and rare species of South American Siluridae in the Carnegie Museum. Annals of the Carnegie Museum, 11, 398 - 404." type="journal article" year="1917">Eigenmann, 1917</bibRefCitation>
</taxonomicName>
[Pisces].
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</paragraph>
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<taxonomicName id="079EE2C2FFAFFFBEDAD5D52C78C9DD6D" ID-CoL="3MKS" authority="Dyte, 1980: 223" authorityName="Dyte" authorityPageNumber="223" authorityYear="1980" box="[151,479,1835,1858]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAFFFBEDAD5D52C7802DD6D" box="[151,276,1835,1858]" italics="true" pageId="1" pageNumber="42">Cheiromyia</emphasis>
<treatmentCitation id="413FBF50FFAFFFBEDB63D52C78C9DD6D" author="Dyte" box="[289,479,1835,1858]" page="223" pageId="1" pageNumber="42" year="1980">
<bibRefCitation id="A40FE4B0FFAFFFBEDB63D52C78B4DD6D" author="Dyte" box="[289,418,1835,1858]" pageId="1" pageNumber="58" refString="Dyte, C. E. (1980) Some replacement names in the Dolichopodidae (Diptera). Entomologica Scandinavica, 11, 223 - 224." type="journal article" year="1980">Dyte, 1980</bibRefCitation>
: 223
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</taxonomicName>
.
<typeStatus id="1F2527E3FFAFFFBEDBB2D52C7B33DD6D" box="[496,549,1835,1858]" pageId="1" pageNumber="42">Type</typeStatus>
species:
<taxonomicName id="079EE2C2FFAFFFBED8D6D52C7D1BDD6D" authority="Parent" authorityName="Parent" box="[660,1037,1835,1858]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="42" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFAFFFBED8D6D52C7AAFDD6D" box="[660,953,1835,1858]" italics="true" pageId="1" pageNumber="42">Cheirocerus palmaticornis</emphasis>
Parent
</taxonomicName>
, automatic. Replacement name for
<taxonomicName id="079EE2C2FFAFFFBEDA85D54C78CDDD4D" ID-CoL="3MJY" authority="Parent, 1930" authorityName="Parent" authorityYear="1930" box="[199,475,1867,1890]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="F2EA4553FFAFFFBEDA85D54C785CDD4D" box="[199,330,1867,1890]" italics="true" pageId="1" pageNumber="42">Cheirocerus</emphasis>
<bibRefCitation id="A40FE4B0FFAFFFBEDB12D54C78CDDD4D" author="Parent" box="[336,475,1867,1890]" pageId="1" pageNumber="58" refString="Parent, O. (1930) Ergebnisse einer zoologischen Sammelreise nach Brasilien, insbesondere in das Amazonasgebiet, ausgefuhrt von Dr. H. Zerny III. Teil. Diptera: Dolichopodidae. Annalen des Naturhistorischen Museums in Wien, 44, 5 - 26." type="journal article" year="1930">Parent, 1930</bibRefCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="C0219941FFAFFFBDDAD5D5957C96DAF7" blockId="1.[151,1436,1938,2003]" lastBlockId="2.[151,1438,152,2012]" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42">
<emphasis id="F2EA4553FFAFFFBEDAD5D595782FDD83" bold="true" box="[151,313,1938,1964]" pageId="1" pageNumber="42">Recognition.</emphasis>
Males can be easily recognized by their distinctive antenna bearing one to several elongate projections on the outer surface of the postpedicel (
<figureCitation id="58A585C4FFAFFFBED8ADD5BE7A25DDFC" box="[751,819,1977,2003]" captionStart="FIGURE 1" captionStartId="3.[151,255,1910,1934]" captionTargetBox="[182,1422,218,1856]" captionTargetId="figure@3.[151,1436,194,1887]" captionTargetPageId="3" captionText="FIGURE 1. Male antenna: (A) Cheiromyia bicornis Brooks sp. nov. left antenna (outer surface); (B) C. palmaticornis (Parent) right antenna (medial surface); (C) C. pennaticornis (Parent) right antenna (medial surface) of holotype; (D) C. pennaticornis (Parent) slide preparation of left antenna (outer surface). Abbreviations: pp apex: postpedicel apex; pp proj: postpedicel projection." httpUri="https://zenodo.org/record/193136/files/figure.png" pageId="1" pageNumber="42">Fig. 1</figureCitation>
), and by the large ventrally projecting apv epandrial lobe of the male hypopygium, with a pair of long fine setae. Females of the genus are problematic to recognize if not collected with males, and cannot readily be distinguished from females of
<taxonomicName id="079EE2C2FFACFFBDDEFCD2B87C96DAF7" ID-CoL="63J56" authority="Loew." authorityName="Loew." box="[1214,1408,190,216]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFACFFBDDEFCD2B87C26DAF7" box="[1214,1328,191,216]" italics="true" pageId="2" pageNumber="43">Paraclius</emphasis>
Loew.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8884CACAFFACFFBDDA87D2E278F0DD46" pageId="2" pageNumber="43" type="description">
<paragraph id="C0219941FFACFFBDDA87D2E278F0DD46" blockId="2.[151,1438,152,2012]" pageId="2" pageNumber="43">
<emphasis id="F2EA4553FFACFFBDDA87D2E278DFDAD0" bold="true" box="[197,457,229,255]" pageId="2" pageNumber="43">Redescription. Head</emphasis>
: Slightly to distinctly broader than high. Vertex slightly to distinctly excavated; 1 pair of strong, inclinate vertical bristles; 1 pair of strong, divergent ocellar bristles; 1 pair of paravertical bristles, subequal to distinctly stronger than uppermost postoculars; postocular setae uniseriate. Face height much greater than height of clypeus. Clypeus subquadrate to subrectangular, lower margin truncate, ending well above lower eye margin. Antenna: scape somewhat enlarged in males, setose dorsally; pedicel somewhat flattened and distorted in males, with marginal setae; male postpedicel modified and elongate with pointed apex and 1 to several elongate projections on the outer surface (
<figureCitation id="58A585C4FFACFFBDD9C3D3CA7AD0DBC8" box="[897,966,461,487]" captionStart="FIGURE 1" captionStartId="3.[151,255,1910,1934]" captionTargetBox="[182,1422,218,1856]" captionTargetId="figure@3.[151,1436,194,1887]" captionTargetPageId="3" captionText="FIGURE 1. Male antenna: (A) Cheiromyia bicornis Brooks sp. nov. left antenna (outer surface); (B) C. palmaticornis (Parent) right antenna (medial surface); (C) C. pennaticornis (Parent) right antenna (medial surface) of holotype; (D) C. pennaticornis (Parent) slide preparation of left antenna (outer surface). Abbreviations: pp apex: postpedicel apex; pp proj: postpedicel projection." httpUri="https://zenodo.org/record/193136/files/figure.png" pageId="2" pageNumber="43">Fig. 1</figureCitation>
); female postpedicel unmodified, short, subtriangular, lacking projections; stylus 2-articled, arista-like.
<emphasis id="F2EA4553FFACFFBDD9CED3F47AFFD822" bold="true" box="[908,1001,499,525]" pageId="2" pageNumber="43">Thorax</emphasis>
: Scutum: acr biserial; 6 dc (anterior bristle usually weak), 1 presutural ial, 1 su, 1 strong outer presutural spal, 1 weaker inner presutural spal, 2 postsutural spal, 1 pal and setula, and 2 npl on each side. Scutellum: 1 strong medial bristle and 1 short lateral bristle on each side. Postpronotum with 1 medioclinate bristle and 1-2 weak setae below. Upper proepisternum in front of anterior spiracle with patch of fine hairs, lower proepisternum with 1 strong medioclinate proepisternal bristle and patch of fine hairs medially; pleural surface in front of posterior spiracle bare; metepisternum with a row of 23 fine hairs.
<emphasis id="F2EA4553FFACFFBDD912D0DC7A9DD8DA" bold="true" box="[848,907,731,757]" pageId="2" pageNumber="43">Legs</emphasis>
: I: CxI: anterior surface with dark setulae, 3 strong setae on lateral margin and several long setulae on apical margin; FI with series of 24 apical pv setae increasing in size distally, distalmost seta usually distinctly stronger in
<taxonomicName id="079EE2C2FFACFFBDD991D12E7D81D96D" box="[979,1175,809,834]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="43" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFACFFBDD991D12E7D81D96D" box="[979,1175,809,834]" italics="true" pageId="2" pageNumber="43">C. palmaticornis</emphasis>
</taxonomicName>
; TI with 2 dorsal, 12 ad and 2 apical (1 dorsal, 1 posterior) bristles, with comb-like row of ad setae (weaker in males); tarsus I longer than TI, It15 usually decreasing in length distally (It2 shorter than It
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males of
<taxonomicName id="079EE2C2FFACFFBDDEE4D1707C50D9BF" box="[1190,1350,887,912]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="F2EA4553FFACFFBDDEE4D1707C50D9BF" box="[1190,1350,887,912]" italics="true" pageId="2" pageNumber="43">C. brevitarsis</emphasis>
</taxonomicName>
and
<taxonomicName id="079EE2C2FFACFFBDDFC3D17079FCD996" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="F2EA4553FFACFFBDDFC3D17079FCD996" italics="true" pageId="2" pageNumber="43">C. laselva</emphasis>
</taxonomicName>
) (
<figureCitation id="58A585C4FFACFFBDDB41D1A7785CD995" box="[259,330,928,954]" captionStart="FIGURE 4" captionStartId="7.[151,255,1873,1897]" captionTargetBox="[164,1420,509,1835]" captionTargetId="figure@7.[151,1436,502,1849]" captionTargetPageId="7" captionText="FIGURE 4. Cheiromyia brevitarsis Brooks sp. nov. male foreleg and terminalia: (A) tibia and tarsus of right foreleg; (B) hypopygium (left lateral view); (C) hypandrium and phallus (ventral view); (D) sperm pump and phallus (left lateral view). Abbreviations: apv lb: apicoventral epandrial lobe; cerc: cercus; dsur: dorsal arm of surstylus; ej apod: ejaculatory apodeme; hypd: hypandrium; ph: phallus; spm pump: sperm pump; t: tarsomere; vsur: ventral arm of surstylus." httpUri="https://zenodo.org/record/193138/files/figure.png" pageId="2" pageNumber="43">Fig. 4</figureCitation>
A). II: CxII: anterior surface with dark setulae and setae, 1 stronger seta near outer margin; FII with 1 strong ad preapical bristle, with series of apical pv setae increasing in size distally, distalmost seta sometimes distinctly stronger; TII with 4 ad, 34 pd, 2 ventral and 5 apical bristles; tarsus II slightly longer than TII, IIt15 decreasing in length distally. III: CxIII with strong lateral seta near apex; FIII with 1 strong ad preapical bristle; TIII usually with 4 ad (
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<taxonomicName id="079EE2C2FFACFFBDD8EAD6397A78DE78" box="[680,878,1086,1111]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="43" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFACFFBDD8EAD6397A78DE78" box="[680,878,1086,1111]" italics="true" pageId="2" pageNumber="43">C. palmaticornis</emphasis>
</taxonomicName>
), 3 pd, 1 preapical dorsal, 1 weak ventral and 2 apical bristles; tarsus III subequal to or longer than TIII, IIIt1 shorter than IIIt2 with distinct basiventral seta, IIIt25 decreasing in length distally.
<emphasis id="F2EA4553FFACFFBDD803D68A7B90DE88" bold="true" box="[577,646,1165,1191]" pageId="2" pageNumber="43">Wing</emphasis>
(Figs. 2D,E): Hyaline or with brownish tint; R2+3 weakly sinuous basally, straight in distal section; R4+5 straight, sometimes with a weak posterior curve in distal section; distal section of M beyond dm-cu with strong anterior bend near middle ending above wing apex; crossvein dm-cu subequal to, or longer than distal section of CuA1; calypter with black setae; halter pale.
<emphasis id="F2EA4553FFACFFBDDF50D7007C82DF0E" bold="true" box="[1298,1428,1287,1313]" pageId="2" pageNumber="43">Abdomen</emphasis>
: Subconical; tergites 15 setose. Male: tergite 6 bare; sternite 2 unmodifed; sternite 3 unmodified to emarginate and membranous posteromedially; sternite 4 strongly emarginate and membranous posteromedially, sometimes divided; sternite 5 mainly membranous, apparently with paired tubular eversible sacs (observed in some specimens); sternite 6 mainly membranous, weakly sclerotized along anterior margin; segment 7 forming tubular peduncle; sternite 8 ovoid to teardrop-shaped, setose.
<emphasis id="F2EA4553FFACFFBDDE0CD7CB7DF2DFCA" box="[1102,1252,1484,1509]" italics="true" pageId="2" pageNumber="43">Hypopygium</emphasis>
(
<figureCitation id="58A585C4FFACFFBDDEB6D7CB7C50DFC9" box="[1268,1350,1484,1510]" captionStart="FIGURE 3" captionStartId="6.[151,255,1885,1909]" captionTargetBox="[152,1430,541,1884]" captionTargetId="figure@6.[151,1436,525,1895]" captionTargetPageId="6" captionText="FIGURE 3. Cheiromyia bicornis Brooks sp. nov. male terminalia: (A) hypopygium (left lateral view); (B) hypandrium and phallus (ventral view); (C) sperm pump and phallus (left lateral view). Abbreviations: apv lb: apicoventral epandrial lobe; bv lb: basiventral epandrial lobe; cerc: cercus; ej dt: ejaculatory duct; ej apod: ejaculatory apodeme; hypd: hypandrium; ph: phallus; spm pump: sperm pump." httpUri="https://zenodo.org/record/193137/files/figure.png" pageId="2" pageNumber="43">Figs. 3</figureCitation>
,
<figureCitation id="58A585C4FFACFFBDDF11D7CB7C74DFC9" box="[1363,1378,1484,1510]" captionStart="FIGURE 4" captionStartId="7.[151,255,1873,1897]" captionTargetBox="[164,1420,509,1835]" captionTargetId="figure@7.[151,1436,502,1849]" captionTargetPageId="7" captionText="FIGURE 4. Cheiromyia brevitarsis Brooks sp. nov. male foreleg and terminalia: (A) tibia and tarsus of right foreleg; (B) hypopygium (left lateral view); (C) hypandrium and phallus (ventral view); (D) sperm pump and phallus (left lateral view). Abbreviations: apv lb: apicoventral epandrial lobe; cerc: cercus; dsur: dorsal arm of surstylus; ej apod: ejaculatory apodeme; hypd: hypandrium; ph: phallus; spm pump: sperm pump; t: tarsomere; vsur: ventral arm of surstylus." httpUri="https://zenodo.org/record/193138/files/figure.png" pageId="2" pageNumber="43">4</figureCitation>
B-D, 5,6,7): Epandrium subrectangular in lateral view, bv epandrial lobe present or absent (as indicated by bv seta), apv epandrial lobe well-developed, projecting ventrally with a pair of long fine setae, left and right bv and apv epandrial lobes with varying degrees of asymmetry; surstylus divided into digitiform dorsal and ventral arms; postgonite digitiform (upturned in
<taxonomicName id="079EE2C2FFACFFBDD86FD4607BE4DCAF" box="[557,754,1639,1664]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="43" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFACFFBDD86FD4607BE4DCAF" box="[557,754,1639,1664]" italics="true" pageId="2" pageNumber="43">C. palmaticornis</emphasis>
</taxonomicName>
); cercus large, shape various; hypandrium simple, troughlike; sperm pump folded back on itself, with simple single fold, opposing surfaces tightly appressed; phallus tubular, enlarged preapically with dorsal flap-like process, apex narrow; ejaculatory apodeme rather narrow and elongate. Female: Sternites 25 setose, unmodified; segments 68 bare, tergites and sternites of segments 6 and 7 undivided; tergite 8 and sternite 8 divided medially, tergite and sternite fused anterolaterally forming a narrow sclerite. Tergite10 divided medially into hemitergites each bearing 5 spines, spines rounded and somewhat flattened apically.
</paragraph>
</subSubSection>
<subSubSection id="8884CACAFFACFFBBDA87D5727AADDC83" lastPageId="4" lastPageNumber="45" pageId="2" pageNumber="43" type="distribution">
<paragraph id="C0219941FFACFFBDDA87D5727B5BDDF3" blockId="2.[151,1438,152,2012]" pageId="2" pageNumber="43">
<emphasis id="F2EA4553FFACFFBDDA87D5727870DDA0" bold="true" box="[197,358,1909,1935]" pageId="2" pageNumber="43">Distribution.</emphasis>
<taxonomicName id="079EE2C2FFACFFBDDB2CD57178EEDDA0" box="[366,504,1910,1935]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFACFFBDDB2CD57178EEDDA0" box="[366,504,1910,1935]" italics="true" pageId="2" pageNumber="43">Cheiromyia</emphasis>
</taxonomicName>
is now known from Central
<collectingCountry id="B889D9D1FFACFFBDD90DD5727AA2DDA0" box="[847,948,1909,1935]" name="United States of America" pageId="2" pageNumber="43">America</collectingCountry>
(
<collectingCountry id="B889D9D1FFACFFBDD986D5727D5DDDA0" box="[964,1099,1909,1935]" name="Costa Rica" pageId="2" pageNumber="43">Costa Rica</collectingCountry>
),
<collectingCountry id="B889D9D1FFACFFBDDE22D5727DD2DDA0" box="[1120,1220,1909,1935]" name="Ecuador" pageId="2" pageNumber="43">Ecuador</collectingCountry>
,
<collectingCountry id="B889D9D1FFACFFBDDE92D5727C51DDA0" box="[1232,1351,1909,1935]" name="Colombia" pageId="2" pageNumber="43">Colombia</collectingCountry>
,
<collectingCountry id="B889D9D1FFACFFBDDF16D5727C8ADDA0" box="[1364,1436,1909,1935]" name="Brazil" pageId="2" pageNumber="43">Brazil</collectingCountry>
(Acre, Amapá, Amazonas, Roraima, Pará, Maranhão, Sergipe, Pernambuco),
<collectingCountry id="B889D9D1FFACFFBDDE1FD59B7DD6DD99" box="[1117,1216,1948,1974]" name="Guyana" pageId="2" pageNumber="43">Guyana</collectingCountry>
,
<collectingCountry id="B889D9D1FFACFFBDDE8CD59B7C2CDD99" box="[1230,1338,1948,1974]" name="Suriname" pageId="2" pageNumber="43">Surinam</collectingCountry>
,
<collectingCountry id="B889D9D1FFACFFBDDF05D59B79FADDF3" name="French Guiana" pageId="2" pageNumber="43">French Guiana</collectingCountry>
and
<collectingCountry id="B889D9D1FFACFFBDDB66D5C5786CDDF3" box="[292,378,1986,2012]" name="Bolivia" pageId="2" pageNumber="43">Bolivia</collectingCountry>
(La Paz) (
<figureCitation id="58A585C4FFACFFBDDBB4D5C57B2BDDF3" box="[502,573,1986,2012]" captionStart="FIGURE 8" captionStartId="16.[151,256,1978,2002]" captionTargetBox="[190,1403,232,1977]" captionTargetId="figure@16.[190,1414,194,1994]" captionTargetPageId="16" captionText="FIGURE 8. Known distribution of Cheiromyia species: (A) C. bicornis Brooks sp. nov. and C. brevitarsis Brooks sp. nov.; (B) C. laselva Brooks sp. nov., C. palmaticornis (Parent), and C. pennaticornis (Parent)." httpUri="https://zenodo.org/record/193141/files/figure.png" pageId="2" pageNumber="43">Fig. 8</figureCitation>
).
</paragraph>
<caption id="94E1C9C9FFADFFBCDAD5D57178D2DDC2" httpUri="https://zenodo.org/record/193136/files/figure.png" pageId="3" pageNumber="44" targetBox="[182,1422,218,1856]" targetPageId="3">
<paragraph id="C0219941FFADFFBCDAD5D57178D2DDC2" blockId="3.[151,1436,1910,2029]" pageId="3" pageNumber="44">
<emphasis id="F2EA4553FFADFFBCDAD5D571780CDDA1" bold="true" box="[151,282,1910,1934]" pageId="3" pageNumber="44">FIGURE 1.</emphasis>
Male antenna: (A)
<taxonomicName id="079EE2C2FFADFFBCDBB2D5707A37DDA2" authority="Brooks" authorityName="Brooks" box="[496,801,1910,1934]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="bicornis" status="sp. nov.">
<emphasis id="F2EA4553FFADFFBCDBB2D5707BDBDDA1" box="[496,717,1911,1934]" italics="true" pageId="3" pageNumber="44">Cheiromyia bicornis</emphasis>
Brooks
</taxonomicName>
<emphasis id="F2EA4553FFADFFBCD968D5717A97DDA1" bold="true" box="[810,897,1910,1934]" pageId="3" pageNumber="44">
<taxonomicNameLabel id="E9D9F828FFADFFBCD968D5717A97DDA1" box="[810,897,1910,1934]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
left antenna (outer surface); (B)
<taxonomicName id="079EE2C2FFADFFBCDEA4D57079FADD82" authority="Parent" authorityName="Parent" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFADFFBCDEA4D5707C8ADDA1" box="[1254,1436,1911,1934]" italics="true" pageId="3" pageNumber="44">C. palmaticornis</emphasis>
(Parent)
</taxonomicName>
right antenna (medial surface); (C)
<taxonomicName id="079EE2C2FFADFFBCD82DD5907A68DD82" authority="Parent" authorityName="Parent" box="[623,894,1942,1966]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="pennaticornis">
<emphasis id="F2EA4553FFADFFBCD82DD5907A37DD81" box="[623,801,1943,1966]" italics="true" pageId="3" pageNumber="44">C. pennaticornis</emphasis>
(Parent)
</taxonomicName>
right antenna (medial surface) of holotype; (D)
<taxonomicName id="079EE2C2FFADFFBCDFC6D590789ADDE2" authority="Parent" authorityName="Parent" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="pennaticornis">
<emphasis id="F2EA4553FFADFFBCDFC6D590783BDDE1" italics="true" pageId="3" pageNumber="44">C. pennaticornis</emphasis>
(Parent)
</taxonomicName>
slide preparation of left antenna (outer surface). Abbreviations: pp apex: postpedicel apex; pp proj: postpedicel projection.
</paragraph>
</caption>
<paragraph id="C0219941FFAAFFBBDAD5D4257AADDC83" blockId="4.[151,1436,1570,1713]" pageId="4" pageNumber="45">
<emphasis id="F2EA4553FFAAFFBBDAD5D4257808DC15" bold="true" box="[151,286,1570,1594]" pageId="4" pageNumber="45">FIGURE 2.</emphasis>
Male face and wing: (A)
<taxonomicName id="079EE2C2FFAAFFBBD87CD4257AD4DC16" authority="Parent" authorityName="Parent" box="[574,962,1570,1593]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pennaticornis">
<emphasis id="F2EA4553FFAAFFBBD87CD4257A4BDC16" box="[574,861,1570,1593]" italics="true" pageId="4" pageNumber="45">Cheiromyia pennaticornis</emphasis>
(Parent)
</taxonomicName>
; (B)
<taxonomicName id="079EE2C2FFAAFFBBD9B8D4257DF0DC16" authority="Brooks" authorityName="Brooks" box="[1018,1254,1570,1593]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="brevitarsis" status="sp. nov.">
<emphasis id="F2EA4553FFAAFFBBD9B8D4257D99DC16" box="[1018,1167,1570,1593]" italics="true" pageId="4" pageNumber="45">C. brevitarsis</emphasis>
Brooks
</taxonomicName>
<emphasis id="F2EA4553FFAAFFBBDEACD4257C51DC15" bold="true" box="[1262,1351,1570,1594]" pageId="4" pageNumber="45">
<taxonomicNameLabel id="E9D9F828FFAAFFBBDEACD4257C51DC15" box="[1262,1351,1570,1594]" pageId="4" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (C)
<taxonomicName id="079EE2C2FFAAFFBBDFC1D425789CDC73" authority="Parent" authorityName="Parent" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="45" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFAAFFBBDFC1D425783ADC73" italics="true" pageId="4" pageNumber="45">C. palmaticornis</emphasis>
(Parent)
</taxonomicName>
; (D)
<taxonomicName id="079EE2C2FFAAFFBBDB83D4427BB2DC73" authority="Brooks" authorityName="Brooks" box="[449,676,1605,1628]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="brevitarsis" status="sp. nov.">
<emphasis id="F2EA4553FFAAFFBBDB83D4427B47DC73" box="[449,593,1605,1628]" italics="true" pageId="4" pageNumber="45">C. brevitarsis</emphasis>
Brooks
</taxonomicName>
<emphasis id="F2EA4553FFAAFFBBD8E9D4427A16DC72" bold="true" box="[683,768,1605,1629]" pageId="4" pageNumber="45">
<taxonomicNameLabel id="E9D9F828FFAAFFBBD8E9D4427A16DC72" box="[683,768,1605,1629]" pageId="4" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (E)
<taxonomicName id="079EE2C2FFAAFFBBD974D4427D51DC73" authority="Parent" authorityName="Parent" box="[822,1095,1605,1628]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pennaticornis">
<emphasis id="F2EA4553FFAAFFBBD974D4427AFEDC73" box="[822,1000,1605,1628]" italics="true" pageId="4" pageNumber="45">C. pennaticornis</emphasis>
(Parent)
</taxonomicName>
. Abbreviations: A1: anal vein of wing; CuA1: 1st anterior branch of cubital vein of wing; dm-cu: discal medial-cubital crossvein of wing; R1: 1st radial vein of wing; R2+3: 2nd + 3rd radial vein of wing; R4+5: 4th + 5th radial vein of wing.
</paragraph>
</subSubSection>
<subSubSection id="8884CACAFFAAFFBBDA87D4E6789CDDCC" pageId="4" pageNumber="45" type="discussion">
<paragraph id="C0219941FFAAFFBBDA87D4E6789CDDCC" blockId="4.[151,1437,1761,2019]" pageId="4" pageNumber="45">
<emphasis id="F2EA4553FFAAFFBBDA87D4E67853DCD4" bold="true" box="[197,325,1761,1787]" pageId="4" pageNumber="45">Remarks.</emphasis>
<bibRefCitation id="A40FE4B0FFAAFFBBDB0DD4E67A76DCD4" author="Capellari" box="[335,864,1761,1787]" pageId="4" pageNumber="58" refString="Capellari, R. S. &amp; Amorim, D. S. (2009) Four new species of Paraclius Loew (Diptera: Dolichopodidae) from the Brazilian Atlantic Forest, and notes on the systematic position of Cheiromyia Dyte. Zootaxa, 2274, 51 - 61." type="journal article" year="2009">Capellari and Amorim (2009, figs. 11,12)</bibRefCitation>
recently described and illustrated the female terminalia of
<taxonomicName id="079EE2C2FFAAFFBBDB7ED50F7BA7DD0D" authority="Brooks" authorityName="Brooks" box="[316,689,1800,1826]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="brevitarsis" status="sp. nov.">
<emphasis id="F2EA4553FFAAFFBBDB7ED50F7B47DD0E" box="[316,593,1800,1825]" italics="true" pageId="4" pageNumber="45">Cheiromyia brevitarsis</emphasis>
Brooks
</taxonomicName>
<emphasis id="F2EA4553FFAAFFBBD8FBD50F7A0CDD0D" bold="true" box="[697,794,1800,1826]" pageId="4" pageNumber="45">
<taxonomicNameLabel id="E9D9F828FFAAFFBBD8FBD50F7A0CDD0D" box="[697,794,1800,1826]" pageId="4" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described below), and noted the proximal fusion of sternite 8 and tergite 8 (character state
<quantity id="076634A4FFAAFFBBD8C2D5297BC1DD67" box="[640,727,1838,1864]" metricMagnitude="0" metricUnit="m" metricValue="1.7805399999999998" pageId="4" pageNumber="45" unit="in" value="70.1">70.1 in</quantity>
<bibRefCitation id="A40FE4B0FFAAFFBBD8A2D5297A93DD67" author="Brooks" box="[736,901,1838,1864]" pageId="4" pageNumber="58" refString="Brooks, S. E. (2005) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857, 1 - 158." type="journal article" year="2005">Brooks 2005</bibRefCitation>
). We have also observed this feature in the previously unknown female of the
<typeStatus id="1F2527E3FFAAFFBBD872D5527B74DD40" box="[560,610,1877,1903]" pageId="4" pageNumber="45">type</typeStatus>
species,
<taxonomicName id="079EE2C2FFAAFFBBD88ED5527A84DD41" box="[716,914,1877,1902]" class="Actinopterygii" family="Pimelodidae" genus="Cheirocerus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="45" phylum="Chordata" rank="species" species="palmaticornis">
<emphasis id="F2EA4553FFAAFFBBD88ED5527A84DD41" box="[716,914,1877,1902]" italics="true" pageId="4" pageNumber="45">C. palmaticornis</emphasis>
</taxonomicName>
. As stated by
<bibRefCitation id="A40FE4B0FFAAFFBBDE7AD5527C81DD40" author="Capellari" box="[1080,1431,1877,1903]" pageId="4" pageNumber="58" refString="Capellari, R. S. &amp; Amorim, D. S. (2009) Four new species of Paraclius Loew (Diptera: Dolichopodidae) from the Brazilian Atlantic Forest, and notes on the systematic position of Cheiromyia Dyte. Zootaxa, 2274, 51 - 61." type="journal article" year="2009">Capellari and Amorim (2009)</bibRefCitation>
, the presence of this character state supports the inclusion of
<taxonomicName id="079EE2C2FFAAFFBBD91CD57B7AF1DDBA" box="[862,999,1916,1941]" class="Insecta" family="Dolichopodidae" genus="Cheiromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAAFFBBD91CD57B7AF1DDBA" box="[862,999,1916,1941]" italics="true" pageId="4" pageNumber="45">Cheiromyia</emphasis>
</taxonomicName>
in a clade with
<taxonomicName id="079EE2C2FFAAFFBBDEE6D57B7C8EDDB9" authority="Loew" authorityName="Loew" box="[1188,1432,1916,1942]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAAFFBBDEE6D57B7C5ADDBA" box="[1188,1356,1916,1941]" italics="true" pageId="4" pageNumber="45">Pelastoneurus</emphasis>
Loew
</taxonomicName>
,
<taxonomicName id="079EE2C2FFAAFFBBDAD5D5A4789DDD93" authority="Becker" authorityName="Becker" box="[151,395,1954,1980]" class="Insecta" family="Dolichopodidae" genus="Stenopygium" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAAFFBBDAD5D5A47838DD93" box="[151,302,1955,1980]" italics="true" pageId="4" pageNumber="45">Stenopygium</emphasis>
Becker
</taxonomicName>
,
<taxonomicName id="079EE2C2FFAAFFBBDBDAD5A47B7CDD93" authority="Parent" authorityName="Parent" box="[408,618,1954,1980]" class="Insecta" family="Dolichopodidae" genus="Platyopsis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAAFFBBDBDAD5A47B05DD93" box="[408,531,1955,1980]" italics="true" pageId="4" pageNumber="45">Platyopsis</emphasis>
Parent
</taxonomicName>
, and
<taxonomicName id="079EE2C2FFAAFFBBD8EBD5A47A71DD93" authority="Loew" authorityName="Loew" box="[681,871,1954,1980]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F2EA4553FFAAFFBBD8EBD5A47A0ADD93" box="[681,796,1955,1980]" italics="true" pageId="4" pageNumber="45">Paraclius</emphasis>
Loew
</taxonomicName>
(in part), including the
<taxonomicName id="079EE2C2FFAAFFBBDEC1D5A47C11DD93" box="[1155,1287,1955,1980]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="F2EA4553FFAAFFBBDEC1D5A47C11DD93" box="[1155,1287,1955,1980]" italics="true" pageId="4" pageNumber="45">P. arcuatus</emphasis>
</taxonomicName>
lineage (see
<bibRefCitation id="A40FE4B0FFAAFFBBDAD5D5CE7826DDCC" author="Brooks" box="[151,304,1993,2019]" pageId="4" pageNumber="58" refString="Brooks, S. E. (2005) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857, 1 - 158." type="journal article" year="2005">Brooks 2005</bibRefCitation>
, fig. 2).
</paragraph>
</subSubSection>
</treatment>
</document>