210 lines
25 KiB
XML
210 lines
25 KiB
XML
<document id="2701E85D44A349D8369CEEC7654B6143" ID-CLB-Dataset="298353" ID-DOI="10.5194/fr-23-71-2020" ID-GBIF-Dataset="27226970-29d0-439f-b25e-f97186d9dd20" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965742" IM.treatments_approvedBy="karina" checkinTime="1712928012780" checkinUser="felipe" docAuthor="Chen, Lu, Ren, Dong & BØthoux, Olivier" docDate="2020" docId="761087FB701C8477FFC9FE4EB4FDFA68" docLanguage="en" docName="FossRec.23.1.71-74.pdf" docOrigin="Fossil Record 23 (1)" docSource="http://dx.doi.org/10.5194/fr-23-71-2020" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Phtanomiamia gui Chen & Ren & BØthoux 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="73" masterDocId="8A29FF83701E8475FFB5FF8CB17DFF9A" masterDocTitle="A new, rare and small “ lobeattid ” species (Insecta: Archaeorthoptera) found at Xiaheyan (Pennsylvanian; Ningxia, China)" masterLastPageNumber="74" masterPageNumber="71" pageNumber="73" updateTime="1718752468478" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="B4FD4B8072ADDF3A032F1D139B248F71">
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<mods:title id="EE8EE88AE2E5E27081BC70DA283C2812">A new, rare and small “ lobeattid ” species (Insecta: Archaeorthoptera) found at Xiaheyan (Pennsylvanian; Ningxia, China)</mods:title>
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<mods:name id="978FE140EB8B3680E32D8C2AA6A9CC15" type="personal">
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<mods:roleTerm id="0D39D7784D3839D61F48A0798244C3CE">Author</mods:roleTerm>
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<mods:namePart id="A3F1354D89E94FB0E3DDC23BEA4F0CAC">Chen, Lu</mods:namePart>
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<mods:roleTerm id="06E0C19B3F0FF72E90AB8CD1057EBD8C">Author</mods:roleTerm>
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<mods:namePart id="FADDFEAC5EDCD9F8FABD2B8DCD7CD04F">Ren, Dong</mods:namePart>
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<mods:roleTerm id="6477FA0E1F969DA0569ED2CE09D0DB35">Author</mods:roleTerm>
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<mods:namePart id="A93714B009B9A5FC866DE45FF6E044D0">BØthoux, Olivier</mods:namePart>
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<mods:typeOfResource id="7251DD065A6F8AD455F93B7E1779D9D2">text</mods:typeOfResource>
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<mods:title id="3B27BAAF2A51B95BC89C090B906678FA">Fossil Record</mods:title>
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<mods:part id="2460EEC2A9B3A1D68782826410B03763">
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<mods:date id="7C16AD38DB74B5C51E9AEE3BEF2812E4">2020</mods:date>
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<mods:number id="55885DDB0D0B3D3A3C32EFDEC7AE4ACB">2020-04-08</mods:number>
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<mods:number id="ECE44CFC695A226B787AAA2877CC46FE">23</mods:number>
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<mods:number id="195C58C6E8A915D97DDA5B45E5EA9EAB">1</mods:number>
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<mods:start id="51CFA0F785AA4E3A8301D0045622F3A3">71</mods:start>
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<mods:url id="9F69E1C4570221044602B97330E28E1A">http://dx.doi.org/10.5194/fr-23-71-2020</mods:url>
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<treatment id="761087FB701C8477FFC9FE4EB4FDFA68" ID-DOI="http://doi.org/10.5281/zenodo.11558044" ID-GBIF-Taxon="232727872" ID-Zenodo-Dep="11558044" LSID="urn:lsid:plazi:treatment:761087FB701C8477FFC9FE4EB4FDFA68" httpUri="http://treatment.plazi.org/id/761087FB701C8477FFC9FE4EB4FDFA68" lastPageNumber="73" pageId="2" pageNumber="73">
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<subSubSection id="B6A36566701C8477FFC9FE4EB0E7FE43" box="[124,410,450,474]" pageId="2" pageNumber="73" type="nomenclature">
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<paragraph id="FE0636ED701C8477FFC9FE4EB0E7FE43" blockId="2.[124,410,450,474]" box="[124,410,450,474]" pageId="2" pageNumber="73">
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<heading id="A54E8181701C8477FFC9FE4EB0E7FE43" box="[124,410,450,474]" fontSize="10" level="2" pageId="2" pageNumber="73" reason="5">
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<taxonomicName id="39B94D6E701C8477FFC9FE4EB03CFE40" authority="Chen & Ren & BØthoux, 2020" authorityName="Chen & Ren & BØthoux" authorityYear="2020" box="[124,321,450,474]" class="Insecta" genus="Phtanomiamia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="gui" status="sp. nov.">
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<emphasis id="CCCDEAFF701C8477FFC9FE4EB03CFE40" box="[124,321,450,474]" italics="true" pageId="2" pageNumber="73">Phtanomiamia gui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D7FE5784701C8477FEFDFE4EB0E7FE43" box="[328,410,450,473]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B6A36566701C8477FFC9FE73B3BDFDCE" box="[124,704,511,596]" pageId="2" pageNumber="73" type="materials_examined">
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<paragraph id="FE0636ED701C8477FFC9FE73B1A2FD8D" blockId="2.[124,223,511,535]" box="[124,223,511,535]" pageId="2" pageNumber="73">
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<emphasis id="CCCDEAFF701C8477FFC9FE73B1A2FD8D" bold="true" box="[124,223,511,535]" pageId="2" pageNumber="73">Material</emphasis>
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</paragraph>
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<paragraph id="FE0636ED701C8477FFC9FDB1B3BDFDCE" blockId="2.[124,704,573,596]" box="[124,704,573,596]" pageId="2" pageNumber="73">
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<materialsCitation id="4ED13CB0701C8477FFC9FDB1B3BDFDCE" ID-GBIF-Occurrence="4875704301" box="[124,704,573,596]" collectionCode="CNU" pageId="2" pageNumber="73" specimenCount="1" typeStatus="holotype">
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<typeStatus id="2102884F701C8477FFC9FDB1B1A2FDCE" box="[124,223,573,596]" pageId="2" pageNumber="73" type="holotype">Holotype</typeStatus>
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only, CNU-NX1-324, deposited at the
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<collectionCode id="98A8AE28701C8477FD35FDB1B3C1FDCE" box="[640,700,573,596]" country="China" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Capital Normal University, College of Life Sciences" pageId="2" pageNumber="73">CNU</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6A36566701C8477FFC9FDF6B18AFC94" pageId="2" pageNumber="73" type="etymology">
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<paragraph id="FE0636ED701C8477FFC9FDF6B18AFD08" blockId="2.[124,247,634,658]" box="[124,247,634,658]" pageId="2" pageNumber="73">
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<emphasis id="CCCDEAFF701C8477FFC9FDF6B18AFD08" bold="true" box="[124,247,634,658]" pageId="2" pageNumber="73">Etymology</emphasis>
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</paragraph>
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<paragraph id="FE0636ED701C8477FFC9FD34B18AFC94" blockId="2.[124,767,696,782]" pageId="2" pageNumber="73">The species name is dedicated to Junjie Gu, for his contributions to fieldwork at Xiaheyan and to the systematics of Orthoptera.</paragraph>
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</subSubSection>
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<subSubSection id="B6A36566701C8477FFC9FCB8B0F5FBD3" pageId="2" pageNumber="73" type="diagnosis">
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<paragraph id="FE0636ED701C8477FFC9FCB8B194FCD6" blockId="2.[124,233,820,844]" box="[124,233,820,844]" pageId="2" pageNumber="73">
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<emphasis id="CCCDEAFF701C8477FFC9FCB8B194FCD6" bold="true" box="[124,233,820,844]" pageId="2" pageNumber="73">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="FE0636ED701C8477FFC9FCFEB0F5FBD3" blockId="2.[124,767,882,1097]" pageId="2" pageNumber="73">
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Forewing: ScP–
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<collectionCode id="98A8AE28701C8477FE9FFCFEB046FC13" box="[298,315,882,905]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="73">R</collectionCode>
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/RA area broad (at its broadest, broader than the area between the anterior wing margin and ScP); first fork of M in a distal position (distal to the second fifth of wing length, basal to wing mid-length); CuA + CuPa branched shortly after the fusion of CuA and CuPa; CuA with a distinct, free portion between its divergence from M + CuA and its fusion with CuPa.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6A36566701C8477FFC9FBE2B47FFEFB" pageId="2" pageNumber="73" type="description">
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<paragraph id="FE0636ED701C8477FFC9FBE2B182FB1C" blockId="2.[124,255,1134,1158]" box="[124,255,1134,1158]" pageId="2" pageNumber="73">
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<emphasis id="CCCDEAFF701C8477FFC9FBE2B182FB1C" bold="true" box="[124,255,1134,1158]" pageId="2" pageNumber="73">Description</emphasis>
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</paragraph>
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<paragraph id="FE0636ED701C8477FFC9FB21B0E7F81B" blockId="2.[124,767,1197,1921]" pageId="2" pageNumber="73">
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<typeStatus id="2102884F701C8477FFC9FB21B1A2FB5E" box="[124,223,1197,1220]" pageId="2" pageNumber="73" type="holotype">Holotype</typeStatus>
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specimen (CNU-NX1-324): negative imprint of an isolated, left forewing, apex missing; preserved length about
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<quantity id="39419B08701C8477FFC9FB60B1A6FA99" box="[124,219,1260,1283]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="73" unit="mm" value="12.2">12.2 mm</quantity>
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, estimated length about
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||
<quantity id="39419B08701C8477FE56FB60B33FFA99" box="[483,578,1260,1283]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="73" unit="mm" value="13.8">13.8 mm</quantity>
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, maximum width
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<quantity id="39419B08701C8477FFC9FA80B1B2FAB9" box="[124,207,1292,1315]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="73" unit="mm" value="4.2">4.2 mm</quantity>
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; ScA present, short; ScP–
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<collectionCode id="98A8AE28701C8477FE46FA80B379FAB9" box="[499,516,1292,1315]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="73">R</collectionCode>
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/RA area broader than area between anterior wing margin and ScP; area between the anterior wing margin and ScP/ScP + RA with strong cross-veins, mostly oblique and simple, reticulated near the wing base; ScP reaching RA slightly basal to the second third of wing length; RA/RP fork slightly basal to the first third of wing length; RA simple; RA and RP close and parallel until wing mid-length, where both veins diverge; RP simple for
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<quantity id="39419B08701C8477FF13F987B185F9B8" box="[166,248,1547,1570]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="73" unit="mm" value="4.7">4.7 mm</quantity>
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, posteriorly pectinate (as preserved), with five branches preserved, the anterior-most one reaching RA; M weak, simple for
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<quantity id="39419B08701C8477FE88F9C7B0F6F9F8" box="[317,395,1611,1634]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="73" unit="mm" value="3.7">3.7 mm</quantity>
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before it forks into MA and
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<collectionCode id="98A8AE28701C8477FD64F9C7B386F9F8" box="[721,763,1611,1634]" country="0" name="Mohonk Preserve, Inc." pageId="2" pageNumber="73">MP</collectionCode>
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; MA simple (as preserved),
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<collectionCode id="98A8AE28701C8477FE2AF9E7B0B8F918" box="[415,453,1643,1666]" country="0" name="Mohonk Preserve, Inc." pageId="2" pageNumber="73">MP</collectionCode>
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with two preserved branches (it probably possessed 3); CuA
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<quantity id="39419B08701C8477FE7CF907B36AF938" box="[457,535,1675,1698]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="73" unit="mm" value="0.5">0.5 mm</quantity>
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long before its fusion with CuPa; CuA + CuPa forked immediately after the fusion of CuA and CuPa; CuA + CuPa with a total of five branches (2, 3); CuPb simple and straight; area between CuPb and AA1 narrow; AA1 strong and simple, some cross-vein between AA1 and AA2 have cross-veins between them; AA2 with six branches; cross-veins occasionally reticulated; no colouration pattern visible.
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</paragraph>
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||
<paragraph id="FE0636ED701C8477FC86FF23B562FF5D" blockId="2.[819,1055,175,199]" box="[819,1055,175,199]" pageId="2" pageNumber="73">
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<emphasis id="CCCDEAFF701C8477FC86FF23B562FF5D" bold="true" box="[819,1055,175,199]" pageId="2" pageNumber="73">Locality and horizon</emphasis>
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</paragraph>
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<paragraph id="FE0636ED701C8477FC86FF67B47FFEFB" blockId="2.[819,1462,235,353]" pageId="2" pageNumber="73">
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Xiaheyan Village, Zhongwei City, Yanghugou Formation (
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<collectingRegion id="3C7DF80F701C8477FC8EFE87B2BDFEB8" box="[827,960,267,290]" country="China" name="Ningxia" pageId="2" pageNumber="73">Ningxia Hui</collectingRegion>
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Autonomous Region,
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<collectingCountry id="86AE767D701C8477FB1AFE87B58EFEB8" box="[1199,1267,267,290]" name="China" pageId="2" pageNumber="73">China</collectingCountry>
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); latest Bashkirian (latest Duckmantian) to middle Moscovian (Bolsovian), early Pennsylvanian (
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<bibRefCitation id="9A284B1C701C8477FBA2FEC6B58AFEFB" author="Trumper, S. & Schneider, J. W. & Nemyrovska, T. & Korn, D. & Linnemann, U. G. & Ren, D. & BOthoux, O." box="[1047,1271,330,353]" pageId="2" pageNumber="73" pagination="109444" refId="ref2720" refString="Trumper, S., Schneider, J. W., Nemyrovska, T., Korn, D., Linnemann, U. G., Ren, D., and BOthoux, O.: Age and depositional environment of the Xiaheyan insect fauna, embedded in marine black shales (Early Pennsylvanian, China), Palaeogeogr. Palaeocl., 538, 109444, https: // doi. org / 10.1016 / j. palaeo. 2019.109444, 2020." type="journal article" year="2020">Trümper et al., 2020</bibRefCitation>
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||
).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="B6A36566701C8477FC86FE08B4FDFA68" pageId="2" pageNumber="73" type="discussion">
|
||
<paragraph id="FE0636ED701C8477FC86FE08B2D6FE06" blockId="2.[819,939,388,412]" box="[819,939,388,412]" pageId="2" pageNumber="73">
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||
<emphasis id="CCCDEAFF701C8477FC86FE08B2D6FE06" bold="true" box="[819,939,388,412]" pageId="2" pageNumber="73">Discussion</emphasis>
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||
</paragraph>
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<paragraph id="FE0636ED701C8477FC86FE33B4FDFA68" blockId="2.[819,1463,447,1523]" pageId="2" pageNumber="73">
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The new specimen can be confidently assigned to Archaeorthoptera as it exhibits the character “CuA (diverging from M + CuA) fused with the anterior branch of CuP (CuPa)”, which is the defining character state of this taxon (
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<bibRefCitation id="9A284B1C701C8477FAE6FD93B210FDCC" author="BOthoux, O." pageId="2" pageNumber="73" pagination="135 - 156" refId="ref2002" refString="BOthoux, O.: Cladotypic taxonomy applied: titanopterans are orthopterans, Arth. Syst. Phyl., 65, 135 - 156, 2007." type="journal article" year="2007">BØthoux, 2007</bibRefCitation>
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). Conversely, assignment to the Panorthoptera (including crown Orthoptera and stem groups closely related to them) can be excluded, as CuPa is not branched before its fusion with CuA. Non-panorthopteran Archaeorthoptera include “lobeattid” insects, the order
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<taxonomicName id="39B94D6E701C8477FB0BFD32B2BFFD6F" authority="Handlirsch, 1937" authorityName="Handlirsch" authorityYear="1937" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">
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Cnemidolestodea
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<bibRefCitation id="9A284B1C701C8477FA34FD32B2BFFD6F" author="Handlirsch, A." pageId="2" pageNumber="73" pagination="1 - 140" refId="ref2587" refString="Handlirsch, A.: Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. I Teil, Ann. Naturhist. Mus. Wien, 48, 1 - 140, 1937." type="journal article" year="1937">Handlirsch, 1937</bibRefCitation>
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||
</taxonomicName>
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(which is probably derived from a subset of lobeattid insects) and a number of unplaced species. The new specimen displays an RA–RP area narrow for a long distance, a trait occurring in the former, represented at Xiaheyan by
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<taxonomicName id="39B94D6E701C8477FC1EFCD1B4CFFCEF" authority="Prokop and Ren, 2007" authorityName="Prokop and Ren" authorityYear="2007" box="[939,1458,861,885]" class="Insecta" family="Spanioderidae" genus="Sinopteron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="huangheense">
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<emphasis id="CCCDEAFF701C8477FC1EFCD1B5CFFCEF" box="[939,1202,861,885]" italics="true" pageId="2" pageNumber="73">Sinopteron huangheense</emphasis>
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<bibRefCitation id="9A284B1C701C8477FB0EFCD2B4CFFCEF" author="Prokop, J. & Ren, D." box="[1211,1458,862,885]" pageId="2" pageNumber="73" pagination="267 - 275" refId="ref2674" refString="Prokop, J. and Ren, D.: New significant fossil insects from the Upper Carboniferous of Ningxia in northern China (Insecta: Palaeodictyoptera, Archaeorthoptera), Eur. J. Entomol., 104, 267 - 275, 2007." type="journal article" year="2007">Prokop and Ren, 2007</bibRefCitation>
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||
</taxonomicName>
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,
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<taxonomicName id="39B94D6E701C8477FC86FCF1B470FC0F" authority="Liu, Ren and Prokop, 2009" authorityName="Liu, Ren and Prokop" authorityYear="2009" box="[819,1293,893,917]" class="Insecta" genus="Chenxiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="liuae">
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<emphasis id="CCCDEAFF701C8477FC86FCF1B29FFC0F" box="[819,994,893,917]" italics="true" pageId="2" pageNumber="73">Chenxiella liuae</emphasis>
|
||
<bibRefCitation id="9A284B1C701C8477FC5FFCF2B470FC0F" author="Liu, Y. & Ren, D. & Prokop, J." box="[1002,1293,894,917]" pageId="2" pageNumber="73" pagination="63 - 68" refId="ref2632" refString="Liu, Y., Ren, D., and Prokop, J.: Discovery of a new Namurian archaeorthopterid from Ningxia, China (Insecta: Archaeorthoptera), Zootaxa, 2032, 63 - 68, 2009." type="journal article" year="2009">Liu, Ren and Prokop, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39B94D6E701C8477FAADFCF1B4F6FC0F" authorityName="Gu, BOthoux and Ren" authorityYear="2011" box="[1304,1419,893,917]" class="Insecta" family="Spanioderidae" genus="Longzhua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="loculata">
|
||
<emphasis id="CCCDEAFF701C8477FAADFCF1B4F6FC0F" box="[1304,1419,893,917]" italics="true" pageId="2" pageNumber="73">L. loculata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39B94D6E701C8477FA2CFCF1B2FEFC2F" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="maimai">
|
||
<emphasis id="CCCDEAFF701C8477FA2CFCF1B2FEFC2F" italics="true" pageId="2" pageNumber="73">M. maimai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="39B94D6E701C8477FC0CFC11B567FC2F" authorityName="Du, BOthoux, Gu and Ren" authorityYear="2017" box="[953,1050,925,949]" class="Insecta" genus="Protomiamia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="yangi">
|
||
<emphasis id="CCCDEAFF701C8477FC0CFC11B567FC2F" box="[953,1050,925,949]" italics="true" pageId="2" pageNumber="73">Pr. yangi</emphasis>
|
||
</taxonomicName>
|
||
. Among lobeattids, the new specimen shares with
|
||
<taxonomicName id="39B94D6E701C8477FC06FC31B544FC4E" box="[947,1081,957,981]" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="CCCDEAFF701C8477FC06FC31B57BFC4F" box="[947,1030,957,981]" italics="true" pageId="2" pageNumber="73">Miamia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(here considered to include
|
||
<taxonomicName id="39B94D6E701C8477FADCFC31B4CCFC6E" authority="Coty, HAEva, Prokop, Roques and Nel, 2014" authorityName="Coty" authorityYear="2014" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Titanoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="duquesnei">
|
||
<emphasis id="CCCDEAFF701C8477FADCFC31B2B0FC6F" italics="true" pageId="2" pageNumber="73">Aviologus duquesnei</emphasis>
|
||
<bibRefCitation id="9A284B1C701C8477FC63FC51B4CCFC6E" author="Coty, D. & HAEva, J. & Prokop, J. & Roques, P. & Nel, A." box="[982,1457,989,1012]" pageId="2" pageNumber="73" pagination="462 - 470" refId="ref2299" refString="Coty, D., HAEva, J., Prokop, J., Roques, P., and Nel, A.: New archaeorthopteran insects from the Late Carboniferous of the Nord and Pas-de-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera), Zootaxa, 3878, 462 - 470, https: // doi. org / 10.11646 / zootaxa. 3878.5.4, 2014." type="journal article" year="2014">Coty, HAEva, Prokop, Roques and Nel, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
; and see BØthoux, 2008;
|
||
<bibRefCitation id="9A284B1C701C8477FB8DFC71B45FFB8E" author="BOthoux, O. & Gu, J. & Yue, Y. & Ren, D." box="[1080,1314,1021,1044]" pageId="2" pageNumber="73" pagination="103 - 113" refId="ref2227" refString="BOthoux, O., Gu, J., Yue, Y., and Ren, D.: Miamia maimai n. sp., a new Pennsylvanian stem-orthopteran insect, and a case study on the application of cladotypic nomenclature, Foss. Rec., 15, 103 - 113, https: // doi. org / 10.1002 / mmng. 201200008, 2012 b." type="journal article" year="2012">BØthoux et al., 2012b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A284B1C701C8477FA9BFC71B573FBAE" author="BOthoux, O. & Jarzembowski, E. A." pageId="2" pageNumber="73" pagination="87 - 96" refId="ref2075" refString="BOthoux, O. and Jarzembowski, E. A.: New basal neopterans from Writhlington (UK, Pennsylvanian), Alavesia, 3, 87 - 96, 2010." type="journal article" year="2010">BØthoux and Jarzembowski, 2010</bibRefCitation>
|
||
) a comparatively broad ScP–
|
||
<collectionCode id="98A8AE28701C8477FA8AFB91B42DFBAE" box="[1343,1360,1053,1076]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="73">R</collectionCode>
|
||
/RA area. Another peculiar trait shared with the corresponding species (but not unique to
|
||
<taxonomicName id="39B94D6E701C8477FC47FBD0B50AFBEE" box="[1010,1143,1116,1140]" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="CCCDEAFF701C8477FC47FBD0B538FBEE" box="[1010,1093,1116,1140]" italics="true" pageId="2" pageNumber="73">Miamia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
) is the very early branching of CuA + CuPa. However, the new specimen lacks the defining character state of
|
||
<taxonomicName id="39B94D6E701C8477FC5BFB10B53CFB2E" authorityName="Smith" authorityYear="1871" box="[1006,1089,1180,1204]" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CCCDEAFF701C8477FC5BFB10B53CFB2E" box="[1006,1089,1180,1204]" italics="true" pageId="2" pageNumber="73">Miamia</emphasis>
|
||
</taxonomicName>
|
||
, namely “in forewings, CuPa fuses with M + CuA” (BØthoux, 2008), a condition implying that CuA does not possess a free portion between its divergence from M + CuA and its fusion with CuPa. This free portion of CuA is present in the new specimen, which represents a plesiomorphy within Archaeorthoptera. In summary, the new specimen belongs to a species closely related to
|
||
<taxonomicName id="39B94D6E701C8477FA80FAD7B4F5FAE9" authorityName="Smith" authorityYear="1871" box="[1333,1416,1371,1395]" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CCCDEAFF701C8477FA80FAD7B4F5FAE9" box="[1333,1416,1371,1395]" italics="true" pageId="2" pageNumber="73">Miamia</emphasis>
|
||
</taxonomicName>
|
||
, yet it can be confidently excluded from this genus. It follows that it belongs to a new species to be accommodated in its own genus.
|
||
<taxonomicName id="39B94D6E701C8477FCCAFA37B539FA49" authorityName="Chen & Ren & BØthoux" authorityYear="2020" box="[895,1092,1467,1491]" class="Insecta" genus="Phtanomiamia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="gui">
|
||
<emphasis id="CCCDEAFF701C8477FCCAFA37B539FA49" box="[895,1092,1467,1491]" italics="true" pageId="2" pageNumber="73">Phtanomiamia gui</emphasis>
|
||
</taxonomicName>
|
||
is smaller than any known species of
|
||
<taxonomicName id="39B94D6E701C8477FCE5FA57B2DEFA69" authorityName="Smith" authorityYear="1871" box="[848,931,1499,1523]" class="Insecta" family="Sylvaphlebiidae" genus="Miamia" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CCCDEAFF701C8477FCE5FA57B2DEFA69" box="[848,931,1499,1523]" italics="true" pageId="2" pageNumber="73">Miamia</emphasis>
|
||
</taxonomicName>
|
||
, and among the smallest “lobeattid” species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |