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<document id="20D38AE5C6C46A4F539F454A571CC2FB" ID="10.11646/zootaxa.4147.1.4" ID-DOI="10.11646/zootaxa.4147.1.4" ID-GBIF-Dataset="3166e193-d052-4efa-953b-4e5ac1b6f568" ID-ISSN="1175-5326" ID-Zenodo-Dep="266251" ID-ZooBank="39ECCC38-BC63-40D2-9B04-F435841AA0D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470120521773" checkinUser="plazi" docAuthor="Daniel, Gimo M., Davis, Adrian Lv. &amp; Scholtz, Clarke H." docDate="2016" docId="03E087A1E553FFCE88BE9A9464A87F22" docLanguage="en" docName="zootaxa.4147.1.4.pdf" docOrigin="Zootaxa 4147 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sisyphus neobornemisszanus Daniel &amp; Davis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="73" masterDocId="FFD9FFD9E557FFC888299C0B66117C33" masterDocTitle="Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa" masterLastPageNumber="74" masterPageNumber="67" pageNumber="71" updateTime="1698366689639" updateUser="plazi">
<mods:mods id="C2B48F295C1FFA2707775D545A808147" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C2B4FD5634E3576C4ABA8086DE78BD45">
<mods:title id="B85ED1591DF509E418F44FBF4699EADC">Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="0E6B86B3B23163D7A01600C362B537C4" type="personal">
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<mods:namePart id="555452B9E754D00B399B7ABA436F022E">Daniel, Gimo M.</mods:namePart>
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<mods:namePart id="75629FEE2D52451988B50C18FFE03042">Davis, Adrian Lv.</mods:namePart>
</mods:name>
<mods:name id="F98F9CA3742A582585D9602CC4FD470E" type="personal">
<mods:role id="18A77586426181BB39F0663A55D3B593">
<mods:roleTerm id="0AB27D1E4FA3040917234DDF1A97E385">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36497C9433673E9EC182A6C2B4038C97">Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="ECA815ADCDC2882149A61D0BCCFD0574">text</mods:typeOfResource>
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<mods:title id="353DFDDB84A0430252147F1BBEE62846">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5137717FD7F1B7BCC5E05481A515ED1E">
<mods:date id="B872915B4F8C85EDA909309E75C96B47">2016</mods:date>
<mods:detail id="E8AE48FD2F17FEF8B6FF7E26EFF62CE0" type="volume">
<mods:number id="0F1F3F38CF6B9A8C652D68D534FDED2A">4147</mods:number>
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<mods:detail id="4AD9BBA7AB8B185B9F25FD6AD1E758CF" type="issue">
<mods:number id="80111CCBF623A2572E43D7495766449D">1</mods:number>
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<mods:start id="85B958156E5E0961E11935C0EB84B5FC">67</mods:start>
<mods:end id="9027123F15C56728DE9AA81CDA43EA1E">74</mods:end>
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<mods:classification id="A9623FFB60493A679BE9AEA165A02032">journal article</mods:classification>
<mods:identifier id="B75DD96501ACA2350A31FE1E22F6F8FE" type="DOI">10.11646/zootaxa.4147.1.4</mods:identifier>
<mods:identifier id="C9F412861E02B284DB85E59E80F21F79" type="GBIF-Dataset">3166e193-d052-4efa-953b-4e5ac1b6f568</mods:identifier>
<mods:identifier id="0234C0E17D446E564704167A32D39CD3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DB471A260CBEA584865CBB9EF4BEA703" type="Zenodo-Dep">266251</mods:identifier>
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<treatment id="03E087A1E553FFCE88BE9A9464A87F22" ID-DOI="http://doi.org/10.5281/zenodo.6087203" ID-GBIF-Taxon="123513976" ID-Zenodo-Dep="6087203" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22" httpUri="http://treatment.plazi.org/id/03E087A1E553FFCE88BE9A9464A87F22" lastPageId="6" lastPageNumber="73" pageId="4" pageNumber="71">
<subSubSection id="C353653CE553FFCC88BE9A9466E67AE8" pageId="4" pageNumber="71" type="nomenclature">
<paragraph id="8BF636B7E553FFCC88BE9A9465337A89" blockId="4.[151,802,1695,1755]" box="[151,802,1695,1722]" pageId="4" pageNumber="71">
<heading id="D0BE81DBE553FFCC88BE9A9465337A89" bold="true" box="[151,802,1695,1722]" fontSize="11" level="1" pageId="4" pageNumber="71" reason="1">
<emphasis id="B93DEAA5E553FFCC88BE9A9465337A89" bold="true" box="[151,802,1695,1722]" pageId="4" pageNumber="71">
<taxonomicName id="4C494D34E553FFCC88BE9A9464AC7A89" ID-CoL="4XSB4" authority="Daniel &amp; Davis" authorityName="Daniel &amp; Davis" box="[151,701,1695,1722]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="neobornemisszanus" status="sp. nov.">
<emphasis id="B93DEAA5E553FFCC88BE9A9467E97A8A" bold="true" box="[151,504,1695,1721]" italics="true" pageId="4" pageNumber="71">Sisyphus neobornemisszanus</emphasis>
Daniel &amp; Davis
</taxonomicName>
<taxonomicNameLabel id="A20E57DEE553FFCC8AEC9AAB65337A89" box="[709,802,1696,1722]" pageId="4" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B7E553FFCC88BE9AC966E67AE8" blockId="4.[151,802,1695,1755]" box="[151,247,1730,1755]" pageId="4" pageNumber="71">
(
<figureCitation id="13722A32E553FFCC88B69AC966FE7AE8" box="[159,239,1730,1755]" captionStart="FIGURE 3. A. S" captionStartId="5.[151,250,906,928]" captionTargetBox="[364,1231,200,875]" captionTargetId="figure@5.[340,1247,193,883]" captionTargetPageId="5" captionText="FIGURE 3. A. S. neobornessmizanus sp. nov. habitus. Diagnostic features: B. Tufts of setae on elytra. C. Aedeagus." httpUri="https://zenodo.org/record/266254/files/figure.png" pageId="4" pageNumber="71">Fig. 3a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C353653CE553FFCC88BE9B0164637B13" box="[151,626,1802,1824]" pageId="4" pageNumber="71" type="description">
<paragraph id="8BF636B7E553FFCC88BE9B0164637B13" blockId="4.[151,626,1802,1824]" box="[151,626,1802,1824]" pageId="4" pageNumber="71">
<taxonomicName id="4C494D34E553FFCC88BE9B01643E7B13" authority="Endrodi, 1983" authorityName="Endrodi" authorityYear="1983" box="[151,559,1802,1824]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="bornemisszanus">
<emphasis id="B93DEAA5E553FFCC88BE9B0167897B13" box="[151,408,1802,1824]" italics="true" pageId="4" pageNumber="71">Sisyphus bornemisszanus</emphasis>
Endrödi, 1983
</taxonomicName>
(
<emphasis id="B93DEAA5E553FFCC8A149B00647B7B13" box="[573,618,1803,1824]" italics="true" pageId="4" pageNumber="71">pars</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="C353653CE553FFCC88BE9B46631D7BC6" pageId="4" pageNumber="71" type="diagnosis">
<paragraph id="8BF636B7E553FFCC88BE9B46631D7BC6" blockId="4.[151,1436,1869,2038]" pageId="4" pageNumber="71">
<emphasis id="B93DEAA5E553FFCC88BE9B4667007B55" bold="true" box="[151,273,1869,1894]" pageId="4" pageNumber="71">Diagnosis.</emphasis>
This species is close to
<taxonomicName id="4C494D34E553FFCC8A0E9B4665A07B55" authority="Olivier, 1789" authorityName="Olivier" authorityYear="1789" box="[551,945,1869,1894]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="muricatus">
<emphasis id="B93DEAA5E553FFCC8A0E9B4665107B56" box="[551,769,1869,1893]" italics="true" pageId="4" pageNumber="71">Sisyphus muricatus</emphasis>
(Olivier, 1789)
</taxonomicName>
and
<taxonomicName id="4C494D34E553FFCC8BC39B4663847B55" authority="Boheman, 1857" authorityName="Boheman" authorityYear="1857" box="[1002,1429,1869,1894]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="B93DEAA5E553FFCC8BC39B4662CA7B56" box="[1002,1243,1869,1893]" italics="true" pageId="4" pageNumber="71">Sisyphus fasciculatus</emphasis>
Boheman, 1857
</taxonomicName>
. Like
<taxonomicName id="4C494D34E553FFCC88FB9B7A67D87BBA" box="[210,457,1905,1929]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="neobornemisszanus">
<emphasis id="B93DEAA5E553FFCC88FB9B7A67D87BBA" box="[210,457,1905,1929]" italics="true" pageId="4" pageNumber="71">S. neobornemisszanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C494D34E553FFCC89F19B7A64747BBA" box="[472,613,1905,1929]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="muricatus">
<emphasis id="B93DEAA5E553FFCC89F19B7A64747BBA" box="[472,613,1905,1929]" italics="true" pageId="4" pageNumber="71">S. muricatus</emphasis>
</taxonomicName>
also has tufts of setae only on the elytra with uniformly distributed setae on the pronotum. However, the carinae on the head of
<taxonomicName id="4C494D34E553FFCC8B279B9D658D7B9E" box="[782,924,1942,1965]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="muricatus">
<emphasis id="B93DEAA5E553FFCC8B279B9D658D7B9E" box="[782,924,1942,1965]" italics="true" pageId="4" pageNumber="71">S. muricatus</emphasis>
</taxonomicName>
are mostly more developed with deeper and very strong lateral emargination on the pronotum.
<taxonomicName id="4C494D34E553FFCC8B169BB265E37BE2" box="[831,1010,1977,2001]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="B93DEAA5E553FFCC8B169BB265E37BE2" box="[831,1010,1977,2001]" italics="true" pageId="4" pageNumber="71">S. fasciculatus</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="4C494D34E553FFCC8DAE9BB267637BC6" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="neobornemisszanus" status="sp. nov.">
<emphasis id="B93DEAA5E553FFCC8DAE9BB267637BC6" italics="true" pageId="4" pageNumber="71">S. neobornemisszanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57DEE553FFCC89509BD667C27BC5" box="[377,467,2013,2038]" pageId="4" pageNumber="71" rank="species">
<emphasis id="B93DEAA5E553FFCC89509BD667C27BC5" bold="true" box="[377,467,2013,2038]" pageId="4" pageNumber="71">sp. nov.</emphasis>
</taxonomicNameLabel>
by bearing black tufts of setae on both the pronotum and elytra
<emphasis id="B93DEAA5E553FFCC8C899BD663147BC5" bold="true" box="[1184,1285,2013,2038]" pageId="4" pageNumber="71">
(
<figureCitation id="13722A32E553FFCC8C819BD662FD7BC5" box="[1192,1260,2013,2038]" captionStart="FIGURE 3. A. S" captionStartId="5.[151,250,906,928]" captionTargetBox="[364,1231,200,875]" captionTargetId="figure@5.[340,1247,193,883]" captionTargetPageId="5" captionText="FIGURE 3. A. S. neobornessmizanus sp. nov. habitus. Diagnostic features: B. Tufts of setae on elytra. C. Aedeagus." httpUri="https://zenodo.org/record/266254/files/figure.png" pageId="4" pageNumber="71">Fig. 3</figureCitation>
b)
</emphasis>
.
</paragraph>
</subSubSection>
<caption id="DF36663FE552FFCD88BE9F81632F7F93" box="[151,1342,906,928]" httpUri="https://zenodo.org/record/266254/files/figure.png" pageId="5" pageNumber="72" targetBox="[364,1231,200,875]" targetPageId="5">
<paragraph id="8BF636B7E552FFCD88BE9F81632F7F93" blockId="5.[151,1342,906,928]" box="[151,1342,906,928]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88BE9F81673A7F93" bold="true" box="[151,299,906,928]" pageId="5" pageNumber="72">FIGURE 3. A</emphasis>
.
<taxonomicName id="4C494D34E552FFCD89119F8064067F93" box="[312,535,906,928]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="neobornessmizanus" status="sp. nov.">
<emphasis id="B93DEAA5E552FFCD89119F8064067F93" box="[312,535,906,928]" italics="true" pageId="5" pageNumber="72">S. neobornessmizanus</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E552FFCD8A379F81647F7F93" bold="true" box="[542,622,906,928]" pageId="5" pageNumber="72">
<taxonomicNameLabel id="A20E57DEE552FFCD8A379F81647F7F93" box="[542,622,906,928]" pageId="5" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B93DEAA5E552FFCD8A5C9F81658A7F93" box="[629,923,906,928]" italics="true" pageId="5" pageNumber="72">habitus. Diagnostic features:</emphasis>
<emphasis id="B93DEAA5E552FFCD8B8B9F8165A27F93" bold="true" box="[930,947,906,928]" pageId="5" pageNumber="72">B</emphasis>
. Tufts of setae on elytra.
<emphasis id="B93DEAA5E552FFCD8C9D9F8162D77F93" bold="true" box="[1204,1222,906,928]" pageId="5" pageNumber="72">C</emphasis>
. Aedeagus.
</paragraph>
</caption>
<subSubSection id="C353653CE552FFCD88EE9FC367D87B72" pageId="5" pageNumber="72" type="description">
<paragraph id="8BF636B7E552FFCD88EE9FC367137837" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE9FC367CF7FD2" bold="true" box="[199,478,968,993]" pageId="5" pageNumber="72">Description. Size: Male</emphasis>
: Length:
<quantity id="4CB19B52E552FFCD8A679FC364C97FD3" box="[590,728,968,993]" metricMagnitude="-3" metricUnit="m" metricValue="8.649999999999999" metricValueMax="9.7" metricValueMin="7.6" pageId="5" pageNumber="72" unit="mm" value="8.649999999999999" valueMax="9.7" valueMin="7.6">7.6–9.7 mm</quantity>
, width:
<quantity id="4CB19B52E552FFCD8B1C9FC365D17FD3" box="[821,960,968,993]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.6" metricValueMin="4.6" pageId="5" pageNumber="72" unit="mm" value="5.1" valueMax="5.6" valueMin="4.6">4.6–5.6 mm</quantity>
;
<emphasis id="B93DEAA5E552FFCD8BE59FC362327FD2" bold="true" box="[972,1059,968,993]" pageId="5" pageNumber="72">Female</emphasis>
: Length: 8.0–
<quantity id="4CB19B52E552FFCD8CEF9FC263077FD3" box="[1222,1302,969,993]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="72" unit="mm" value="10.0">10 mm</quantity>
, width
<quantity id="4CB19B52E552FFCD8D429FC266EF7837" metricMagnitude="-3" metricUnit="m" metricValue="5.01" metricValueMax="5.62" metricValueMin="4.4" pageId="5" pageNumber="72" unit="mm" value="5.01" valueMax="5.62" valueMin="4.4">4.4– 5.62 mm</quantity>
.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE981B670E787E" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE981B6735781A" bold="true" box="[199,292,1040,1065]" pageId="5" pageNumber="72">Colour:</emphasis>
Black, opaque; except mouth parts, and tarsi which are reddish brown; Antennal club dark gray or dark brown.
</paragraph>
<paragraph id="8BF636B7E552FFCD88F99853638C78A7" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88F99853671E7842" bold="true" box="[208,271,1112,1137]" pageId="5" pageNumber="72">Head</emphasis>
: Apex of clypeus broadly and deeply emarginate, bordered by two acute medial teeth and two more obtuse lateral teeth, genae angular anteriorly and curved laterally, surface of clypeus pubescent and finely punctate.
</paragraph>
<paragraph id="8BF636B7E552FFCD88FA98AB636C78EE" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88FA98AB6744788A" bold="true" box="[211,341,1184,1209]" pageId="5" pageNumber="72">Pronotum:</emphasis>
Convex with dense cover of setae, much broader than long, lateral edges curved anteriorly, narrowed posteriorly, complete lateral prothoracic ridge separating prothoracic disc and prothoracic episternum.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE98E363817916" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE98E367027932" bold="true" box="[199,275,1256,1281]" pageId="5" pageNumber="72">Elytra</emphasis>
: narrowed posteriorly, interstriae 1, 3, 5, and 7 with rows of black setae arranged in tufts, interstriae 2, 4, and 6 with sparsely arranged single setae, a rudimentary row of setae along outer margin of abdominal pleurae.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE993B624D797A" blockId="5.[151,1437,968,2004]" box="[199,1116,1328,1353]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE993B672B797A" bold="true" box="[199,314,1328,1353]" pageId="5" pageNumber="72">Pygidium</emphasis>
: V-shaped, with indistinct rows of brown setae and ocellate punctation.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE995F624B79A2" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE995F6728795E" bold="true" box="[199,313,1364,1389]" pageId="5" pageNumber="72">Sternites:</emphasis>
Six visible abdominal sternites with setae and ocellate punctures; meso- and metasternum black, shiny and pubescent with ocellate punctation; meso-metasternal suture clearly visible.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE999762247A76" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE999766EE7986" bold="true" box="[199,255,1436,1461]" pageId="5" pageNumber="72">Legs</emphasis>
: Profemur punctate ventrally and setose laterally; meso- and metafemur pubescent with ocellate punctation; mesotrochanter short and contiguous with femur; metatrochanter finely punctate, slightly angular and projected away from femur; Protibia with three teeth and a single terminal spurs; meso- and metatibia serrated, setose with two terminal spurs; pro- meso- and metatarsus five segmented with two claws; the first tarsal segment of the meso- and meta-thoracic legs with a row of strong setae on the ventral edge.
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE9A5B66D37A82" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE9A5B67557A5A" bold="true" box="[199,324,1616,1641]" pageId="5" pageNumber="72">Aedeagus:</emphasis>
Phallobase slightly curved dorsally as is common in the genus
<taxonomicName id="4C494D34E552FFCD8C359A5B626F7A5B" box="[1052,1150,1616,1640]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEAA5E552FFCD8C359A5B626F7A5B" box="[1052,1150,1616,1640]" italics="true" pageId="5" pageNumber="72">Sisyphus</emphasis>
</taxonomicName>
; parameres symmetrical, short, triangular, separated by membranous area dorsally, sides almost straight and slightly impressed above (
<emphasis id="B93DEAA5E552FFCD8D479A7F66A37A82" bold="true" pageId="5" pageNumber="72">
<figureCitation id="13722A32E552FFCD8D479A7F66B47A82" captionStart="FIGURE 3. A. S" captionStartId="5.[151,250,906,928]" captionTargetBox="[364,1231,200,875]" captionTargetId="figure@5.[340,1247,193,883]" captionTargetPageId="5" captionText="FIGURE 3. A. S. neobornessmizanus sp. nov. habitus. Diagnostic features: B. Tufts of setae on elytra. C. Aedeagus." httpUri="https://zenodo.org/record/266254/files/figure.png" pageId="5" pageNumber="72">Fig. 3</figureCitation>
c
</emphasis>
).
</paragraph>
<paragraph id="8BF636B7E552FFCD88EE9AB767D87B72" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE9AB764067AE6" bold="true" box="[199,535,1724,1749]" pageId="5" pageNumber="72">Sexual dimorphism. Female</emphasis>
: Lateral carina of pronotum somewhat stronger and more curved than in male; protibial teeth longer; meta- and mesotibia almost straight; last visible abdominal sternite not constricted medially.
<emphasis id="B93DEAA5E552FFCD88BE9B0F66C57B2E" bold="true" box="[151,212,1796,1821]" pageId="5" pageNumber="72">Male</emphasis>
: Protibial teeth strong and short with rudimentary lateral setae; meta- and mesotibia curved; last abdominal sternite narrowed medially.
</paragraph>
</subSubSection>
<subSubSection id="C353653CE552FFCD88EE9B4767C37BBB" pageId="5" pageNumber="72" type="etymology">
<paragraph id="8BF636B7E552FFCD88EE9B4767C37BBB" blockId="5.[151,1437,968,2004]" pageId="5" pageNumber="72">
<emphasis id="B93DEAA5E552FFCD88EE9B47675F7B56" bold="true" box="[199,334,1868,1893]" pageId="5" pageNumber="72">Etymology.</emphasis>
Species named as
<taxonomicName id="4C494D34E552FFCD8A0C9B46650C7B56" box="[549,797,1869,1893]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="neobornemisszanus">
<emphasis id="B93DEAA5E552FFCD8A0C9B46650C7B56" box="[549,797,1869,1893]" italics="true" pageId="5" pageNumber="72">S. neobornemisszanus</emphasis>
</taxonomicName>
, because the designated
<typeStatus id="54F28815E552FFCD8C119B47628B7B56" box="[1080,1178,1868,1893]" pageId="5" pageNumber="72" type="holotype">holotype</typeStatus>
was in the mixed
<typeStatus id="54F28815E552FFCD8D459B46638D7B56" box="[1388,1436,1869,1893]" pageId="5" pageNumber="72">type</typeStatus>
series of
<taxonomicName id="4C494D34E552FFCD88D59B7A67DA7BBB" box="[252,459,1904,1928]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="bornemisszanus">
<emphasis id="B93DEAA5E552FFCD88D59B7A67DA7BBB" box="[252,459,1904,1928]" italics="true" pageId="5" pageNumber="72">S. bornemisszanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C353653CE552FFCE88EE9B9C64A87F22" lastPageId="6" lastPageNumber="73" pageId="5" pageNumber="72" type="materials_examined">
<paragraph id="8BF636B7E552FFCE88EE9B9C639C7EB1" blockId="5.[151,1437,968,2004]" lastBlockId="6.[151,1437,151,786]" lastPageId="6" lastPageNumber="73" pageId="5" pageNumber="72">
<materialsCitation id="3B213CEAE552FFCD88EE9B9C621F7B83" ID-GBIF-Occurrence="1290984908" box="[199,1038,1943,1968]" collectionCode="TMSA" collectorName="Examined" country="South Africa" location="Zululand" pageId="5" pageNumber="72" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E552FFCD88EE9B9C67D47B83" bold="true" box="[199,453,1943,1968]" pageId="5" pageNumber="72">
<collectorName id="26BC5361E552FFCD88EE9B9C67517B83" box="[199,320,1943,1968]" pageId="5" pageNumber="72">Examined</collectorName>
material. (
</emphasis>
<specimenCount id="9D4FFD3EE552FFCD89EC9B9367CB7B9C" box="[453,474,1944,1967]" pageId="5" pageNumber="72" type="male">♂</specimenCount>
<emphasis id="B93DEAA5E552FFCD89C89B9C65897B83" bold="true" box="[481,920,1943,1968]" pageId="5" pageNumber="72">
<collectionCode id="ED58AE72E552FFCD89C89B9C64257B83" box="[481,564,1943,1968]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="72">TMSA</collectionCode>
)
<typeStatus id="54F28815E552FFCD8A6A9B9C64A57B83" box="[579,692,1943,1968]" pageId="5" pageNumber="72" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F35E7627E552FFCD8AE89B9C65897B83" box="[705,920,1943,1968]" name="South Africa" pageId="5" pageNumber="72">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<location id="8E96606CE552FFCD8B8F9B9C621F7B83" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE552FFCD8B8F9B9C621F7B83" box="[934,1038,1943,1968]" country="South Africa" name="Zululand" pageId="5" pageNumber="72">Zululand</location>
</materialsCitation>
,
<materialsCitation id="3B213CEAE552FFCD8C339B9364A67BE7" ID-GBIF-Occurrence="1290984914" collectingDate="1975-12-07" collectorName="Endrody-Younga" country="Saint Lucia" location="St. Lucia" pageId="5" pageNumber="72" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F35E7627E552FFCD8C339B93626E7B83" box="[1050,1151,1943,1968]" name="Saint Lucia" pageId="5" pageNumber="72">St. Lucia</collectingCountry>
28[◦] 13[’] 12[’’] S; 32[◦] 15[’] 00[’’] E,
<date id="FFF71077E552FFCD89699BB067A57BE7" box="[320,436,1979,2004]" pageId="5" pageNumber="72" value="1975-12-07">
<collectingDate id="EFB3E99FE552FFCD89699BB067A57BE7" box="[320,436,1979,2004]" pageId="5" pageNumber="72" value="1975-12-07">7.xii.1975</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E552FFCD89DD9BB064A67BE7" box="[500,695,1979,2004]" pageId="5" pageNumber="72">Endrody-Younga</collectorName>
</materialsCitation>
.
<materialsCitation id="3B213CEAE552FFCE8AED9BB064857C81" ID-GBIF-Occurrence="1290984902" collectionCode="TMSA" country="South Africa" lastPageId="6" lastPageNumber="73" location="Mission Rock" municipality="Zululand" pageId="5" pageNumber="72" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B93DEAA5E552FFCD8AED9BB065547BE7" bold="true" box="[708,837,1979,2004]" pageId="5" pageNumber="72">
<typeStatus id="54F28815E552FFCD8AED9BB065507BE7" box="[708,833,1979,2004]" pageId="5" pageNumber="72" type="paratype">Paratypes</typeStatus>
:
</emphasis>
(
<specimenCount id="9D4FFD3EE552FFCD8B709BB7656B7BE7" box="[857,890,1980,2004]" pageId="5" pageNumber="72" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E552FFCD8BAB9BB065C27BE7" bold="true" box="[898,979,1979,2004]" pageId="5" pageNumber="72">
<collectionCode id="ED58AE72E552FFCD8BAB9BB065C27BE7" box="[898,979,1979,2004]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="72">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E552FFCD8BCC9BB062AC7BE7" bold="true" box="[997,1213,1979,2004]" pageId="5" pageNumber="72">
<collectingCountry id="F35E7627E552FFCD8BCC9BB062AC7BE7" box="[997,1213,1979,2004]" name="South Africa" pageId="5" pageNumber="72">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<collectingMunicipality id="6B92ACCDE552FFCD8CE49BB063247BE7" box="[1229,1333,1979,2004]" pageId="5" pageNumber="72">Zululand</collectingMunicipality>
,
<location id="8E96606CE552FFCE8D6B9BB066C47C80" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE552FFCE8D6B9BB066C47C80" country="South Africa" lastPageId="6" lastPageNumber="73" municipality="Zululand" name="Mission Rock" pageId="5" pageNumber="72">Mission Rock</location>
, 28[◦] 15[’] 36[’’] S; 32[◦] 17[’] 24[’’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE8A899C9166EE7CE9" ID-GBIF-Occurrence="1290984913" collectingDate="1975-12-07" collectionCode="TMSA" collectorName="Endrody-Younga." country="South Africa" location="Zululand" pageId="6" pageNumber="73" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="FFF71077E551FFCE8A899C9165057C80" box="[672,788,154,179]" pageId="6" pageNumber="73" value="1975-12-07">
<collectingDate id="EFB3E99FE551FFCE8A899C9165057C80" box="[672,788,154,179]" pageId="6" pageNumber="73" value="1975-12-07">7.xii.1975</collectingDate>
</date>
, legs:
<collectorName id="26BC5361E551FFCE8B759C9162327C80" box="[860,1059,154,179]" pageId="6" pageNumber="73">Endrody-Younga.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8C1B9C9062477C80" box="[1074,1110,155,179]" pageId="6" pageNumber="73" type="male">2♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE8C749C9162BF7C80" bold="true" box="[1117,1198,154,179]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8C749C9162BF7C80" box="[1117,1198,154,179]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="73">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE8C969C9163847C80" bold="true" box="[1215,1429,154,179]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8C969C9163847C80" box="[1215,1429,154,179]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<location id="8E96606CE551FFCE88BE9CCA66EE7CE9" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE88BE9CCA66EE7CE9" box="[151,255,193,218]" country="South Africa" name="Zululand" pageId="6" pageNumber="73">Zululand</location>
</materialsCitation>
, St. Lucia 28[◦] 13[’] 12[’’] S; 32[◦] 15[’] 00[’’] E,
<materialsCitation id="3B213CEAE551FFCE8B4D9CCA67F27CCD" ID-GBIF-Occurrence="1290984911" collectingDate="1971-08-28" collectionCode="TMSA" collectorName="Bornemissza." country="South Africa" location="Zululand" pageId="6" pageNumber="73" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="FFF71077E551FFCE8B4D9CCA65FE7CE9" box="[868,1007,193,218]" pageId="6" pageNumber="73" value="1971-08-28">
<collectingDate id="EFB3E99FE551FFCE8B4D9CCA65FE7CE9" box="[868,1007,193,218]" pageId="6" pageNumber="73" value="1971-08-28">28.viii.1971</collectingDate>
</date>
. leg. GF.
<collectorName id="26BC5361E551FFCE8C4E9CCA62EE7CE9" box="[1127,1279,193,218]" pageId="6" pageNumber="73">Bornemissza.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8D3A9CC963277CE9" box="[1299,1334,194,218]" pageId="6" pageNumber="73" type="male">2♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE8D6B9CCA63827CE9" bold="true" box="[1346,1427,193,218]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8D6B9CCA63827CE9" box="[1346,1427,193,218]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="73">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE88BE9CEE677C7CCD" bold="true" box="[151,365,229,254]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE88BE9CEE677C7CCD" box="[151,365,229,254]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<location id="8E96606CE551FFCE89529CEE67F27CCD" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE89529CEE67F27CCD" box="[379,483,229,254]" country="South Africa" name="Zululand" pageId="6" pageNumber="73">Zululand</location>
</materialsCitation>
, St. Lucia, Estuary.
<materialsCitation id="3B213CEAE551FFCE8AEA9CEE620F7D17" ID-GBIF-Occurrence="1290984907" collectingDate="1971-08-28" collectionCode="TMSA" collectorName="Bornemissza." country="South Africa" location="Sodwana Bay" municipality="Zululand" pageId="6" pageNumber="73" specimenCount="28" specimenCount-female="13" specimenCount-male="15" typeStatus="paratype">
<date id="FFF71077E551FFCE8AEA9CEE655C7CCD" box="[707,845,229,254]" pageId="6" pageNumber="73" value="1971-08-28">
<collectingDate id="EFB3E99FE551FFCE8AEA9CEE655C7CCD" box="[707,845,229,254]" pageId="6" pageNumber="73" value="1971-08-28">28.viii.1971</collectingDate>
</date>
, leg: GF.
<collectorName id="26BC5361E551FFCE8B909CEE62437CCD" box="[953,1106,229,254]" pageId="6" pageNumber="73">Bornemissza.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8C489CEE62847CCD" box="[1121,1173,229,254]" pageId="6" pageNumber="73" type="male">15♂</specimenCount>
,
<specimenCount id="9D4FFD3EE551FFCE8C889CED62C37CCD" box="[1185,1234,230,254]" pageId="6" pageNumber="73" type="female">13♀</specimenCount>
<emphasis id="B93DEAA5E551FFCE8CF09CEE633B7CCD" bold="true" box="[1241,1322,229,254]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8CF09CEE633B7CCD" box="[1241,1322,229,254]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="73">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE8D139CEE67157D16" bold="true" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8D139CEE67157D16" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<collectingMunicipality id="6B92ACCDE551FFCE893A9D07676A7D16" box="[275,379,268,293]" pageId="6" pageNumber="73">Zululand</collectingMunicipality>
,
<location id="8E96606CE551FFCE89AE9D0764357D16" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE89AE9D0764357D16" box="[391,548,268,293]" country="South Africa" municipality="Zululand" name="Sodwana Bay" pageId="6" pageNumber="73">Sodwana Bay</location>
,
<quantity id="4CB19B52E551FFCE8A199D0764747D16" box="[560,613,268,293]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="73" unit="km" value="5.0">5km</quantity>
, 27[◦] 21[’] 0[’’] S; 32[◦] 23[’] 24[’’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE8C029D0762B67D78" ID-GBIF-Occurrence="1290984910" collectingDate="1992-11-02" collectionCode="UPSA" collectorName="Endrody-Younga." country="South Africa" location="Richard's Bay" pageId="6" pageNumber="73" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingDate id="EFB3E99FE551FFCE8C029D0762867D16" box="[1067,1175,268,293]" pageId="6" pageNumber="73" value="1992-11-02">
<date id="FFF71077E551FFCE8C029D0762867D16" box="[1067,1175,268,293]" pageId="6" pageNumber="73" value="1992-11-02">2.xi.1992</date>
</collectingDate>
, leg.
<collectorName id="26BC5361E551FFCE8CFC9D07638C7D16" box="[1237,1437,268,293]" pageId="6" pageNumber="73">Endrody-Younga.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE888B9D3866D27D78" box="[162,195,307,331]" pageId="6" pageNumber="73" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E551FFCE88E39D3867027D7F" bold="true" box="[202,275,307,332]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE88E39D3867027D7F" box="[202,275,307,332]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">UPSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE890A9D3867E87D7F" bold="true" box="[291,505,307,332]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE890A9D3867E87D7F" box="[291,505,307,332]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN.
<location id="8E96606CE551FFCE8A649D3864FC7D7F" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE8A649D3864FC7D7F" box="[589,749,307,332]" country="South Africa" name="Richard's Bay" pageId="6" pageNumber="73">Richard’s Bay</location>
, 28[◦] 37[’] 39[’’] S, 32[◦] 17[’] 24[’’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE8C9B9D3863867D41" ID-GBIF-Occurrence="1290984903" collectingDate="2000-01-27" collectionCode="UPSA" collectorName="Davis &amp; Delport." country="South Africa" location="Richard's Bay" pageId="6" pageNumber="73" specimenCount="8" specimenCount-female="2" specimenCount-male="6" typeStatus="paratype">
<date id="FFF71077E551FFCE8C9B9D38630F7D7F" box="[1202,1310,307,332]" pageId="6" pageNumber="73" value="2000-01-27">
<collectingDate id="EFB3E99FE551FFCE8C9B9D38630F7D7F" box="[1202,1310,307,332]" pageId="6" pageNumber="73" value="2000-01-27">27.i.2000</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E551FFCE8D729D38638D7D7F" box="[1371,1436,307,332]" pageId="6" pageNumber="73">Davis</collectorName>
&amp;
<collectorName id="26BC5361E551FFCE889F9D5167027D40" box="[182,275,346,371]" pageId="6" pageNumber="73">Delport.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE890C9D5167597D41" box="[293,328,346,370]" pageId="6" pageNumber="73" type="female">2♀</specimenCount>
,
<specimenCount id="9D4FFD3EE551FFCE89719D51676A7D40" box="[344,379,346,371]" pageId="6" pageNumber="73" type="male">6♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE89AD9D5167DC7D40" bold="true" box="[388,461,346,371]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE89AD9D5167DC7D40" box="[388,461,346,371]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">UPSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE89F69D5164A97D40" bold="true" box="[479,696,346,371]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE89F69D5164A97D40" box="[479,696,346,371]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN.
<location id="8E96606CE551FFCE8B389D5165A57D40" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE8B389D5165A57D40" box="[785,948,346,371]" country="South Africa" name="Richard's Bay" pageId="6" pageNumber="73">Richard’s Bay</location>
, 28[◦] 43[’] 28.5[’’] S; 32[◦] 10[’] 52[’’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE88BE9D8B67377D8C" ID-GBIF-Occurrence="1290984905" collectingDate="2000-01-27" collectionCode="UPSA" collectorName="Davis &amp; Delport." country="South Africa" location="Sileza Nature Reserve" pageId="6" pageNumber="73" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
<date id="FFF71077E551FFCE88BE9D8B67127DAA" box="[151,259,384,409]" pageId="6" pageNumber="73" value="2000-01-27">
<collectingDate id="EFB3E99FE551FFCE88BE9D8B67127DAA" box="[151,259,384,409]" pageId="6" pageNumber="73" value="2000-01-27">27.i.2000</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E551FFCE896A9D8B67957DAA" box="[323,388,384,409]" pageId="6" pageNumber="73">Davis</collectorName>
&amp;
<collectorName id="26BC5361E551FFCE89829D8B64197DAA" box="[427,520,384,409]" pageId="6" pageNumber="73">Delport.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8A339D8B642D7DAA" box="[538,572,384,409]" pageId="6" pageNumber="73" type="female">5♀</specimenCount>
,
<specimenCount id="9D4FFD3EE551FFCE8A659D8A647F7DAA" box="[588,622,385,409]" pageId="6" pageNumber="73" type="male">2♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE8A519D8B64D07DAA" bold="true" box="[632,705,384,409]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8A519D8B64D07DAA" box="[632,705,384,409]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">UPSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE8AFA9D8B65BB7DAA" bold="true" box="[723,938,384,409]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8AFA9D8B65BB7DAA" box="[723,938,384,409]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN,
<location id="8E96606CE551FFCE8C2A9D8B62EE7DAB" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE8C2A9D8B62EE7DAB" box="[1027,1279,384,409]" country="South Africa" name="Sileza Nature Reserve" pageId="6" pageNumber="73">Sileza Nature Reserve</location>
27[◦] 06[’] S; 32[◦] 36[’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE891A9DAC66FB7DD5" ID-GBIF-Occurrence="1290984904" collectingDate="1996-04-03" collectionCode="UPSA" collectorName="Van Resnburg." country="South Africa" location="Sodwana" pageId="6" pageNumber="73" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="FFF71077E551FFCE891A9DAC67BD7DF3" box="[307,428,423,448]" pageId="6" pageNumber="73" value="1996-04-03">
<collectingDate id="EFB3E99FE551FFCE891A9DAC67BD7DF3" box="[307,428,423,448]" pageId="6" pageNumber="73" value="1996-04-03">03.iv.1996</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E551FFCE89C29DA364807DF3" box="[491,657,423,448]" pageId="6" pageNumber="73">Van Resnburg.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8A8D9DA364D47DF3" box="[676,709,424,448]" pageId="6" pageNumber="73" type="female">1♀</specimenCount>
<emphasis id="B93DEAA5E551FFCE8AE49DAC65077DF3" bold="true" box="[717,790,423,448]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8AE49DAC65077DF3" box="[717,790,423,448]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">UPSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE8B0E9DAC65EE7DF3" bold="true" box="[807,1023,423,448]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8B0E9DAC65EE7DF3" box="[807,1023,423,448]" name="South Africa" pageId="6" pageNumber="73">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN,
<location id="8E96606CE551FFCE8C7F9DAC62D17DF3" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE8C7F9DAC62D17DF3" box="[1110,1216,423,448]" country="South Africa" name="Sodwana" pageId="6" pageNumber="73">Sodwana</location>
, 27[◦] 32[’] S; 32[◦] 41[’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE88DE9DC5640D7E3E" ID-GBIF-Occurrence="1290984909" collectingDate="1989-04-07" collectionCode="UPSA" collectorName="Mansfield." country="Mozambique" location="Maputo" pageId="6" pageNumber="73" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Maputo" typeStatus="paratype">
<date id="FFF71077E551FFCE88DE9DC567617DD4" box="[247,368,462,487]" pageId="6" pageNumber="73" value="1989-04-07">
<collectingDate id="EFB3E99FE551FFCE88DE9DC567617DD4" box="[247,368,462,487]" pageId="6" pageNumber="73" value="1989-04-07">07.iv.1989</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E551FFCE89869DC564397DD4" box="[431,552,462,487]" pageId="6" pageNumber="73">Mansfield.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8A139DC4644D7DD4" box="[570,604,463,487]" pageId="6" pageNumber="73" type="female">2♀</specimenCount>
,
<specimenCount id="9D4FFD3EE551FFCE8A459DC5649F7DD4" box="[620,654,462,487]" pageId="6" pageNumber="73" type="male">5♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE8ABE9DC564F17DD4" bold="true" box="[663,736,462,487]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE8ABE9DC564F17DD4" box="[663,736,462,487]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">UPSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE8ADB9DC565AF7DD4" bold="true" box="[754,958,462,487]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8ADB9DC565AF7DD4" box="[754,958,462,487]" name="Mozambique" pageId="6" pageNumber="73">MOZAMBIQUE</collectingCountry>
</emphasis>
,
<collectingRegion id="498DF855E551FFCE8BE59DC462377DD5" box="[972,1062,463,486]" country="Mozambique" name="Maputo" pageId="6" pageNumber="73">Maputo</collectingRegion>
, Elephant
<collectingRegion id="498DF855E551FFCE8CB69DC462E77DD5" box="[1183,1270,463,486]" country="Mozambique" name="Maputo" pageId="6" pageNumber="73">Maputo</collectingRegion>
Reserve, 26[◦] 22[’] 30[’’] S; 33[◦] 47[’] 50[’’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE8A039DFE64E07E07" ID-GBIF-Occurrence="1290984912" collectingDate="2003-01-14" collectingDateMax="2003-01-16" collectingDateMin="2003-01-14" collectionCode="UPSA" collectorName="Scholtz &amp; Holter." country="Mozambique" location="Maputo" pageId="6" pageNumber="73" specimenCount="2" specimenCount-female="2" stateProvince="Maputo" typeStatus="paratype">
<date id="FFF71077E551FFCE8A039DFE64AD7E3D" box="[554,700,501,526]" pageId="6" pageNumber="73" value="2003-01-14" valueMax="2003-01-16" valueMin="2003-01-14">
<collectingDate id="EFB3E99FE551FFCE8A039DFE64AD7E3D" box="[554,700,501,526]" pageId="6" pageNumber="73" value="2003-01-14" valueMax="2003-01-16" valueMin="2003-01-14">14-16.i.2003</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E551FFCE8AD49DFE65407E3D" box="[765,849,501,526]" pageId="6" pageNumber="73">Scholtz</collectorName>
&amp;
<collectorName id="26BC5361E551FFCE8B539DFE65D67E3D" box="[890,967,501,526]" pageId="6" pageNumber="73">Holter.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8BF39DFD65ED7E3D" box="[986,1020,502,526]" pageId="6" pageNumber="73" type="female">2♀</specimenCount>
<collectionCode id="ED58AE72E551FFCE8C2F9DFE625E7E3D" box="[1030,1103,501,526]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="6" pageNumber="73">
<emphasis id="B93DEAA5E551FFCE8C2F9DFE625E7E3D" bold="true" box="[1030,1103,501,526]" pageId="6" pageNumber="73">UPSA</emphasis>
</collectionCode>
)
<emphasis id="B93DEAA5E551FFCE8C4B9DFE633F7E3D" bold="true" box="[1122,1326,501,526]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE8C4B9DFE633F7E3D" box="[1122,1326,501,526]" name="Mozambique" pageId="6" pageNumber="73">MOZAMBIQUE</collectingCountry>
</emphasis>
,
<collectingRegion id="498DF855E551FFCE8D179DFD63897E3E" box="[1342,1432,502,525]" country="Mozambique" name="Maputo" pageId="6" pageNumber="73">Maputo</collectingRegion>
, Elephant
<collectingRegion id="498DF855E551FFCE892B9E1667487E07" box="[258,345,541,564]" country="Mozambique" name="Maputo" pageId="6" pageNumber="73">Maputo</collectingRegion>
Reserve, 26[◦] 39[’] S; 32[◦] 43[’] E
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE8AD59E1767D97E68" ID-GBIF-Occurrence="1290984906" collectingDate="2007-11-10" collectingDateMax="2007-11-16" collectingDateMin="2007-11-10" collectionCode="TMSA" collectorName="Strumpher &amp; Deschodt." country="Mozambique" location="MOCAMBIQUE" pageId="6" pageNumber="73" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="FFF71077E551FFCE8AD59E17658A7E06" box="[764,923,540,565]" pageId="6" pageNumber="73" value="2007-11-10" valueMax="2007-11-16" valueMin="2007-11-10">
<collectingDate id="EFB3E99FE551FFCE8AD59E17658A7E06" box="[764,923,540,565]" pageId="6" pageNumber="73" value="2007-11-10" valueMax="2007-11-16" valueMin="2007-11-10">10-16.xi.2007</collectingDate>
</date>
,
<collectingMethod id="52084EA0E551FFCE8B819E1762317E07" box="[936,1056,540,565]" pageId="6" pageNumber="73">pitfall trap</collectingMethod>
, leg:
<collectorName id="26BC5361E551FFCE8C749E1762C27E06" box="[1117,1235,540,565]" pageId="6" pageNumber="73">Strumpher</collectorName>
&amp;
<collectorName id="26BC5361E551FFCE8CDE9E1763787E06" box="[1271,1385,540,565]" pageId="6" pageNumber="73">Deschodt.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8D509E17638D7E07" box="[1401,1436,540,564]" pageId="6" pageNumber="73" type="male">1♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE88BE9E4966F97E68" bold="true" box="[151,232,578,603]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE88BE9E4966F97E68" box="[151,232,578,603]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="73">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE88D29E4967D97E68" bold="true" box="[251,456,578,603]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE88D29E4967D97E68" box="[251,456,578,603]" name="Mozambique" pageId="6" pageNumber="73">MOÇAMBIQUE</collectingCountry>
</emphasis>
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE89F39E4967D47EB1" ID-GBIF-Occurrence="1290984937" collectionCode="TMSA" collectorName="Inharrime &amp; Moor." country="Mozambique" location="Inhambane" pageId="6" pageNumber="73" specimenCount="1" specimenCount-male="1" stateProvince="Inhambane" typeStatus="paratype">
<collectingRegion id="498DF855E551FFCE89F39E4964487E68" box="[474,601,578,603]" country="Mozambique" name="Inhambane" pageId="6" pageNumber="73">Inhambane</collectingRegion>
,
<collectorName id="26BC5361E551FFCE8A419E4964C87E68" box="[616,729,578,603]" pageId="6" pageNumber="73">Inharrime</collectorName>
, 24[◦] 28[’] 37[’’] S; 35[◦] 01[’] 49[’’], 1974, leg:
<collectorName id="26BC5361E551FFCE8D0B9E4863777E69" box="[1314,1382,579,602]" pageId="6" pageNumber="73">Moor.</collectorName>
(
<specimenCount id="9D4FFD3EE551FFCE8D509E48638D7E68" box="[1401,1436,579,603]" pageId="6" pageNumber="73" type="male">1♂</specimenCount>
<emphasis id="B93DEAA5E551FFCE88BE9E6266F97EB1" bold="true" box="[151,232,617,642]" pageId="6" pageNumber="73">
<collectionCode id="ED58AE72E551FFCE88BE9E6266F97EB1" box="[151,232,617,642]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="73">TMSA</collectionCode>
</emphasis>
)
<emphasis id="B93DEAA5E551FFCE88D19E6267D47EB1" bold="true" box="[248,453,617,642]" pageId="6" pageNumber="73">
<collectingCountry id="F35E7627E551FFCE88D19E6267D47EB1" box="[248,453,617,642]" name="Mozambique" pageId="6" pageNumber="73">MOÇAMBIQUE</collectingCountry>
</emphasis>
</materialsCitation>
,
<materialsCitation id="3B213CEAE551FFCE89FA9E6263997EB1" ID-GBIF-Occurrence="1290984938" box="[467,1416,614,642]" collectingDate="1974-05-02" collectorName="A. Strydom" country="Mozambique" location="Inhambane" pageId="6" pageNumber="73" specimenCount="1" stateProvince="Inhambane" typeStatus="paratype">
<collectingRegion id="498DF855E551FFCE89FA9E6264427EB1" box="[467,595,617,642]" country="Mozambique" name="Inhambane" pageId="6" pageNumber="73">Inhambane</collectingRegion>
, Pomene 22[◦] 35[’] 24[’’] S; 35[◦] 21[’] 00[’’],
<date id="FFF71077E551FFCE8C4C9E6162D67EB1" box="[1125,1223,618,642]" pageId="6" pageNumber="73" value="1974-05-02">
<collectingDate id="EFB3E99FE551FFCE8C4C9E6162D67EB1" box="[1125,1223,618,642]" pageId="6" pageNumber="73" value="1974-05-02">2.v.1974</collectingDate>
</date>
, leg.
<collectorName id="26BC5361E551FFCE8D2A9E6163997EB1" box="[1283,1416,617,642]" pageId="6" pageNumber="73">A. Strydom</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636B7E551FFCE88EE9E8664A87F22" blockId="6.[151,1437,151,786]" pageId="6" pageNumber="73">
<materialsCitation id="3B213CEAE551FFCE88EE9E8664A87F22" ID-GBIF-Occurrence="1290984943" country="Mozambique" county="Pomene" latitude="-25.11" location="KwaZulu-Natal Province" longLatPrecision="749" longitude="33.73" municipality="South Africa" pageId="6" pageNumber="73" specimenCount="1" stateProvince="Maputo">
<emphasis id="B93DEAA5E551FFCE88EE9E86673B7E95" bold="true" box="[199,298,653,678]" pageId="6" pageNumber="73">Habitat.</emphasis>
This species has been recorded in
<location id="8E96606CE551FFCE8AEB9E8665CE7E95" LSID="urn:lsid:plazi:treatment:03E087A1E553FFCE88BE9A9464A87F22:8E96606CE551FFCE8AEB9E8665CE7E95" box="[706,991,653,678]" country="Mozambique" county="Pomene" latitude="-25.11" longLatPrecision="749" longitude="33.73" municipality="South Africa" name="KwaZulu-Natal Province" pageId="6" pageNumber="73" stateProvince="Maputo">KwaZulu-Natal Province</location>
(
<collectingMunicipality id="6B92ACCDE551FFCE8BD99E8662977E95" box="[1008,1158,653,678]" pageId="6" pageNumber="73">South Africa</collectingMunicipality>
) as well as
<collectingRegion id="498DF855E551FFCE8D3A9E85637B7E96" box="[1299,1386,654,677]" country="Mozambique" name="Maputo" pageId="6" pageNumber="73">Maputo</collectingRegion>
and
<collectingRegion id="498DF855E551FFCE88BE9EBA67057EF9" box="[151,276,689,714]" country="Mozambique" name="Inhambane" pageId="6" pageNumber="73">Inhambane</collectingRegion>
Provinces (
<collectingCountry id="F35E7627E551FFCE898B9EBA642D7EF9" box="[418,572,689,714]" name="Mozambique" pageId="6" pageNumber="73">Mozambique</collectingCountry>
), although records from
<collectingCounty id="62974E3BE551FFCE8B4D9EB965AF7EFA" box="[868,958,690,713]" pageId="6" pageNumber="73">Pomene</collectingCounty>
suggest a wider distribution northwards along the
<collectingCountry id="F35E7627E551FFCE892D9EDE67897EDD" box="[260,408,725,750]" name="Mozambique" pageId="6" pageNumber="73">Mozambique</collectingCountry>
coastline (
<geoCoordinate id="EE7D5070E551FFCE8A399EDE64797EDD" box="[528,616,725,750]" direction="south" orientation="latitude" pageId="6" pageNumber="73" precision="555" value="-25.11">25.11 S</geoCoordinate>
;
<geoCoordinate id="EE7D5070E551FFCE8A5D9EDD64DF7EDD" box="[628,718,726,750]" direction="east" orientation="longitude" pageId="6" pageNumber="73" precision="555" value="33.73">33.73 E</geoCoordinate>
). In Maputaland, it is found within coastal sand and dune forests where the canopy is closed and the floor is open.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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