286 lines
36 KiB
XML
286 lines
36 KiB
XML
<document id="5CAD8A6B3E1E3469DB9FEF4996D57487" ID-CLB-Dataset="45813" ID-DOI="10.11646/zootaxa.3795.2.7" ID-GBIF-Dataset="c58f6542-8063-48c0-a888-e584b7b13ec3" ID-ISSN="1175-5326" ID-Zenodo-Dep="228024" ID-ZooBank="564EFF9C-4734-4206-AE7E-E3C38F6E143D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460418351797" checkinUser="plazi" docAuthor="Dow, Rory A." docDate="2014" docId="03B18228921DAF69E3A91BCFFEA7D657" docLanguage="en" docName="zt03795p186.pdf" docOrigin="Zootaxa 3795 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onychogomphus marijanmatoki Dow, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="186" masterDocId="FF88FA50921DAF6CE33E1D5FFFAAD74A" masterDocTitle="Onychogomphus marijanmatoki, a new species from Sarawak, Borneo (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="186" masterPageNumber="181" pageNumber="181" updateTime="1698329334006" updateUser="ExternalLinkService">
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<mods:titleInfo id="71F7C550D2DF06BF599377C98B24529B">
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<mods:title id="B334FA19A8F9D541577AA42187004687">Onychogomphus marijanmatoki, a new species from Sarawak, Borneo (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:name id="EAC85E6DD222135365C0A0B35AEA4CF7" type="personal">
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<mods:roleTerm id="FD7423294FCA65CF1B4728A5537DAC49">Author</mods:roleTerm>
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<mods:namePart id="8F1E17CC17463EFF27055D524B14D488">Dow, Rory A.</mods:namePart>
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<mods:typeOfResource id="02996DDAF6C252E9EED81D0605E29B1A">text</mods:typeOfResource>
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<mods:title id="A9B5ACAC6F7B92770D867F7B53D934DC">Zootaxa</mods:title>
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<mods:part id="9685DDD14CE8D9771474B07164839AFB">
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<mods:date id="3C7B67637A29C7F85C75FF63F33690EE">2014</mods:date>
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<mods:number id="FE64661BE484559B6757738A3F40EF31">3795</mods:number>
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<treatment id="03B18228921DAF69E3A91BCFFEA7D657" ID-DOI="http://doi.org/10.5281/zenodo.6124462" ID-GBIF-Taxon="119426064" ID-Zenodo-Dep="6124462" LSID="urn:lsid:plazi:treatment:03B18228921DAF69E3A91BCFFEA7D657" httpUri="http://treatment.plazi.org/id/03B18228921DAF69E3A91BCFFEA7D657" lastPageId="5" lastPageNumber="186" pageId="0" pageNumber="181">
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<subSubSection id="C30260B5921DAF6CE3A91BCFFE8AD186" pageId="0" pageNumber="181" type="nomenclature">
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<paragraph id="8BA7333E921DAF6CE3A91BCFFD29D1E1" blockId="0.[151,643,1680,1741]" box="[151,643,1680,1707]" pageId="0" pageNumber="181">
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<heading id="D0EF8452921DAF6CE3A91BCFFD29D1E1" bold="true" box="[151,643,1680,1707]" fontSize="11" level="1" pageId="0" pageNumber="181" reason="1">
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<emphasis id="B96CEF2C921DAF6CE3A91BCFFD29D1E1" bold="true" box="[151,643,1680,1707]" pageId="0" pageNumber="181">
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<taxonomicName id="4C1848BD921DAF6CE3A91BCFFDB4D1E0" ID-CoL="9X3TX" box="[151,542,1680,1706]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="marijanmatoki" status="sp. nov.">
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<emphasis id="B96CEF2C921DAF6CE3A91BCFFDB4D1E0" bold="true" box="[151,542,1680,1706]" italics="true" pageId="0" pageNumber="181">Onychogomphus marijanmatoki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F5257921DAF6CE1181BCEFD29D1E1" box="[550,643,1681,1707]" pageId="0" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA7333E921DAF6CE3A91BEBFE8AD186" blockId="0.[151,643,1680,1741]" box="[151,288,1716,1741]" pageId="0" pageNumber="181">
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(
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||
<figureCitation id="13232FBB921DAF6CE3A11BEBFEBDD186" box="[159,279,1716,1741]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 5" captionStart-2="FIGURES 6 – 7" captionStart-3="FIGURES 8 – 9" captionStart-4="FIGURES 10 – 12" captionStartId-0="1.[151,250,1470,1492]" captionStartId-1="2.[151,264,1558,1581]" captionStartId-2="2.[151,264,1905,1928]" captionStartId-3="3.[151,264,1267,1290]" captionStartId-4="4.[151,264,1877,1900]" captionTargetBox-0="[151,1436,339,1445]" captionTargetBox-1="[151,1436,445,1536]" captionTargetBox-2="[160,1425,1671,1876]" captionTargetBox-3="[165,1410,176,1235]" captionTargetBox-4="[384,1197,208,1841]" captionTargetId-0="figure@1.[151,1436,339,1446]" captionTargetId-1="figure@2.[151,1436,445,1537]" captionTargetId-2="figure@2.[151,1436,1661,1884]" captionTargetId-3="figure@3.[151,1436,161,1246]" captionTargetId-4="figure@4.[365,1221,193,1856]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionTargetPageId-4="4" captionText-0="FIGURE 1. Onychogomphus marijanmatoki holotype habitus." captionText-1="FIGURES 2 – 5. Onychogomphus marijanmatoki holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view." captionText-2="FIGURES 6 – 7. Onychogomphus marijanmatoki holotype, wings: (6) left forewing; (7) left hindwing." captionText-3="FIGURES 8 – 9. Onychogomphus marijanmatoki holotype: (8) accessory genitalia, lateral view; (9) terminal segment of genital ligula, ventral view." captionText-4="FIGURES 10 – 12. Onychogomphus marijanmatoki holotype, anal appendages: (10) lateral view; (11) ventral view; (12) dorsal view." httpUri-0="https://zenodo.org/record/228025/files/figure.png" httpUri-1="https://zenodo.org/record/228026/files/figure.png" httpUri-2="https://zenodo.org/record/228027/files/figure.png" httpUri-3="https://zenodo.org/record/228028/files/figure.png" httpUri-4="https://zenodo.org/record/228029/files/figure.png" pageId="0" pageNumber="181">Figs. 1–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30260B5921DAF6CE3A91BA4FAD0D072" pageId="0" pageNumber="181" type="materials_examined">
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<paragraph id="8BA7333E921DAF6CE3A91BA4FAD0D072" blockId="0.[151,1437,1787,2031]" pageId="0" pageNumber="181">
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<emphasis id="B96CEF2C921DAF6CE3A91BA4FE2CD05E" bold="true" box="[151,390,1787,1812]" pageId="0" pageNumber="181">
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Material.
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<typeStatus id="54A38D9C921DAF6CE22E1BA3FE2BD05E" box="[272,385,1788,1812]" pageId="0" pageNumber="181" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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♂ (SAR09_10_GOM30),
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<collectingCountry id="F30F73AE921DAF6CE1871BA3FC88D05F" box="[697,802,1788,1813]" name="Malaysia" pageId="0" pageNumber="181">Malaysia</collectingCountry>
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,
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<collectingCountry id="F30F73AE921DAF6CE00E1BA3FC39D05F" box="[816,915,1788,1813]" name="Malaysia" pageId="0" pageNumber="181">Sarawak</collectingCountry>
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, Miri Division, Gunung Mulu National Park, Sungai Melinau Paku, flying over shallow riffle,
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<date id="FFA615FE921DAF6CE1821A7FFC8BD072" box="[700,801,1824,1848]" pageId="0" pageNumber="181" value="2010-05-07">7 v 2010</date>
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, leg. R. A. Dow. To be deposited in
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<collectionCode id="ED09ABFB921DAF6CE7891A7FFAA1D07D" LSID="urn:lsid:biocol.org:col:34992" box="[1207,1291,1824,1847]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="0" pageNumber="181">RMNH</collectionCode>
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(Leiden).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30260B5921DAF6CE3F91A18FE42D0ED" pageId="0" pageNumber="181" type="etymology">
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<paragraph id="8BA7333E921DAF6CE3F91A18FE42D0ED" blockId="0.[151,1437,1787,2031]" pageId="0" pageNumber="181">
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<emphasis id="B96CEF2C921DAF6CE3F91A18FEE4D015" bold="true" box="[199,334,1863,1887]" pageId="0" pageNumber="181">Etymology.</emphasis>
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<taxonomicName id="4C1848BD921DAF6CE26A1A18FE53D015" box="[340,505,1863,1887]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="marijanmatoki">
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<emphasis id="B96CEF2C921DAF6CE26A1A18FE53D015" box="[340,505,1863,1887]" italics="true" pageId="0" pageNumber="181">marijanmatoki</emphasis>
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</taxonomicName>
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, a noun in the genitive case, named after Marijan Matok (born
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<date id="FFA615FE921DAF6CE7831A18FADCD015" box="[1213,1398,1859,1887]" pageId="0" pageNumber="181" value="1972-03-28">28th March 1972</date>
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) of Ulm-Söflingen,
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<collectingCountry id="F30F73AE921DAF6CE26D1A34FE16D0C9" box="[339,444,1899,1923]" name="Germany" pageId="0" pageNumber="181">Germany</collectingCountry>
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, in appreciation of his support of odonatological research in
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<collectingCountry id="F30F73AE921DAF6CE79C1A35FAA2D0C9" box="[1186,1288,1898,1923]" name="Malaysia" pageId="0" pageNumber="181">Malaysia</collectingCountry>
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through the International Dragonfly Fund.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30260B5921DAF6FE3F91AEDFA9FD222" lastPageId="3" lastPageNumber="184" pageId="0" pageNumber="181" type="description">
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<paragraph id="8BA7333E921DAF6CE3F91AEDFA37D0A5" blockId="0.[151,1437,1787,2031]" pageId="0" pageNumber="181">
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<emphasis id="B96CEF2C921DAF6CE3F91AEDFD95D081" bold="true" box="[199,575,1970,1995]" pageId="0" pageNumber="181">
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Description of
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<typeStatus id="54A38D9C921DAF6CE2B61AECFE45D081" box="[392,495,1971,1995]" pageId="0" pageNumber="181" type="holotype">holotype</typeStatus>
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male.
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</emphasis>
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Head (
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<figureCitation id="13232FBB921DAF6CE19C1AEDFCB2D081" box="[674,792,1970,1995]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1558,1581]" captionTargetBox="[151,1436,445,1536]" captionTargetId="figure@2.[151,1436,445,1537]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Onychogomphus marijanmatoki holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view." httpUri="https://zenodo.org/record/228026/files/figure.png" pageId="0" pageNumber="181">
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Figs.
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<docIssue id="CF7B4CD1921DAF6CE1D41AECFD50D081" box="[746,762,1971,1995]" pageId="0" pageNumber="181">2</docIssue>
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–3
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</figureCitation>
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): Labium mostly pale. Labrum mostly greyish-green becoming pale brownish cream laterally, narrow dark brown bands along free margin and adjacent to anteclypeus.
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</paragraph>
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<paragraph id="8BA7333E921CAF6DE3A91DC8FA3CD657" blockId="1.[151,1436,151,285]" pageId="1" pageNumber="182">
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Mandible bases and genae pale. Anteclypeus with lower margin slightly expanded so overlapping labrum (
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<figureCitation id="13232FBB921CAF6DE6771DC8FA24D7FA" box="[1353,1422,151,176]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1558,1581]" captionTargetBox="[151,1436,445,1536]" captionTargetId="figure@2.[151,1436,445,1537]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Onychogomphus marijanmatoki holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view." httpUri="https://zenodo.org/record/228026/files/figure.png" pageId="1" pageNumber="182">Fig. 2</figureCitation>
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), yellowish-brown coloured. Postclypeus dark centrally with yellowish-brown marks in lower lateral quarters. Ante- and postfrons not very sharply divided, dark brown with a yellow marking occupying most of postfrons (
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<figureCitation id="13232FBB921CAF6DE6761D80FA27D7B2" box="[1352,1421,223,248]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1558,1581]" captionTargetBox="[151,1436,445,1536]" captionTargetId="figure@2.[151,1436,445,1537]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Onychogomphus marijanmatoki holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view." httpUri="https://zenodo.org/record/228026/files/figure.png" pageId="1" pageNumber="182">Fig. 3</figureCitation>
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), narrowly divided centrally. Vertex and occiput black and brown. Tubercules behind lateral ocelli, ocelli yellowish.
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</paragraph>
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<caption id="DF6763B6921CAF6DE3A918E1FCBED29E" box="[151,788,1470,1492]" httpUri="https://zenodo.org/record/228025/files/figure.png" pageId="1" pageNumber="182" targetBox="[151,1436,339,1445]" targetPageId="1">
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<paragraph id="8BA7333E921CAF6DE3A918E1FCBED29E" blockId="1.[151,788,1470,1492]" box="[151,788,1470,1492]" pageId="1" pageNumber="182">
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<emphasis id="B96CEF2C921CAF6DE3A918E1FEB9D299" bold="true" box="[151,275,1470,1492]" pageId="1" pageNumber="182">FIGURE 1.</emphasis>
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<taxonomicName id="4C1848BD921CAF6DE22718E1FDF4D29E" box="[281,606,1470,1492]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="marijanmatoki">
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<emphasis id="B96CEF2C921CAF6DE22718E1FDF4D29E" box="[281,606,1470,1492]" italics="true" pageId="1" pageNumber="182">Onychogomphus marijanmatoki</emphasis>
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</taxonomicName>
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holotype habitus.
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</paragraph>
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</caption>
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<paragraph id="8BA7333E921CAF6DE3F918A3FEF8D037" blockId="1.[151,1437,1532,2024]" pageId="1" pageNumber="182">
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Thorax: Prothorax almost entirely dark brown and black, pale along carina of anterior pronotal lobe. Synthorax dark brown with pale yellow markings as follows (
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<figureCitation id="13232FBB921CAF6DE1E21B40FCE1D172" box="[732,843,1567,1592]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1558,1581]" captionTargetBox="[151,1436,445,1536]" captionTargetId="figure@2.[151,1436,445,1537]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Onychogomphus marijanmatoki holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view." httpUri="https://zenodo.org/record/228026/files/figure.png" pageId="1" pageNumber="182">Figs. 4–5</figureCitation>
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): short mesothoracic collar narrowly separated from narrow antehumeral stripes that reach to approximately the apex of the antealar triangle. A broad stripe occupying most of mesepimeron, narrowing somewhat toward wing bases. A broad stripe occupying all of metepimeron except narrowly below metepleural suture. Venter obscurely dark and pale. Legs (
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<figureCitation id="13232FBB921CAF6DE7551BD3FB1BD1EE" box="[1131,1201,1676,1701]" captionStart="FIGURE 1" captionStartId="1.[151,250,1470,1492]" captionTargetBox="[151,1436,339,1445]" captionTargetId="figure@1.[151,1436,339,1446]" captionTargetPageId="1" captionText="FIGURE 1. Onychogomphus marijanmatoki holotype habitus." httpUri="https://zenodo.org/record/228025/files/figure.png" pageId="1" pageNumber="182">Fig. 1</figureCitation>
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) with coxae mixed yellowish, grey and brown, trochanters yellow and brown, rest mostly black, with brownish-yellow marks on flexor surfaces of femora, these most extensive on posterior legs where extending onto upper part of extensor surface. Wings (
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<figureCitation id="13232FBB921CAF6DE26E1BA8FE6AD05A" box="[336,448,1783,1808]" captionStart="FIGURES 6 – 7" captionStartId="2.[151,264,1905,1928]" captionTargetBox="[160,1425,1671,1876]" captionTargetId="figure@2.[151,1436,1661,1884]" captionTargetPageId="2" captionText="FIGURES 6 – 7. Onychogomphus marijanmatoki holotype, wings: (6) left forewing; (7) left hindwing." httpUri="https://zenodo.org/record/228027/files/figure.png" pageId="1" pageNumber="182">Figs. 6–7</figureCitation>
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): Sectors of arculus well separated at origin with 2 (left) or 3 (right) crossveins before first bifurcation of superior sector in Fw and
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||
<quantity id="4CE09EDB921CAF6DE1B11A43FD10D07F" box="[655,698,1820,1845]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="182" unit="in" value="1.0">1 in</quantity>
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Hw. Discoidal field with two rows of cells from origin to ca level of nodus in both wings. 13 Ax in Fw,
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<quantity id="4CE09EDB921CAF6DE1721A1FFDD3D012" box="[588,633,1855,1880]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="1" pageNumber="182" unit="in" value="9.0">9 in</quantity>
|
||
Hw, 11 (left) or 10 (right) Px in Fw,
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||
<quantity id="4CE09EDB921CAF6DE7181A1FFBF8D012" box="[1062,1106,1855,1880]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="1" pageNumber="182" unit="in" value="9.0">9 in</quantity>
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||
Hw. Pt brown, covering 4–5 underlying cells.
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</paragraph>
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<paragraph id="8BA7333E921CAF6EE3F91AD8FC01D6CD" blockId="1.[151,1437,1532,2024]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182">
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Abdomen: In dorsal view S1–2 moderately broad, contracting in apical half of S3, slender to S7 where it expands in apical half, maximum width at S8–9, then slightly more narrow on S10. Black with pale markings as follows (
|
||
<figureCitation id="13232FBB921CAF6DE3C31A90FEE8D0A2" box="[253,322,1999,2024]" captionStart="FIGURE 1" captionStartId="1.[151,250,1470,1492]" captionTargetBox="[151,1436,339,1445]" captionTargetId="figure@1.[151,1436,339,1446]" captionTargetPageId="1" captionText="FIGURE 1. Onychogomphus marijanmatoki holotype habitus." httpUri="https://zenodo.org/record/228025/files/figure.png" pageId="1" pageNumber="182">Fig. 1</figureCitation>
|
||
): S1–2 pale yellow laterally, including auricles, this colour deepening in distal half of S2, and a narrow yellow mid dorsal stripe, tapering to rear of S2. S3–6 with deep yellow basal markings, S7 with a more extensive deep yellow marking, occupying basal ca two–thirds of tergite. On S3 this marking partly divided dorsally, on S4–S7 narrowly but completely divided dorsally. S8–9 with lower part of sides orangey brown. S10 dark basally, largely yellow brown apically. Cerci yellow-brown in basal ca three-quarters, turning to brown apically, shaped as shown in
|
||
<figureCitation id="13232FBB921FAF6EE2E31C78FDCED60A" box="[477,612,295,320]" captionStart="FIGURES 10 – 12" captionStartId="4.[151,264,1877,1900]" captionTargetBox="[384,1197,208,1841]" captionTargetId="figure@4.[365,1221,193,1856]" captionTargetPageId="4" captionText="FIGURES 10 – 12. Onychogomphus marijanmatoki holotype, anal appendages: (10) lateral view; (11) ventral view; (12) dorsal view." httpUri="https://zenodo.org/record/228029/files/figure.png" pageId="2" pageNumber="183">Figs. 10, 11</figureCitation>
|
||
. Epiproct (
|
||
<figureCitation id="13232FBB921FAF6EE1DE1C78FCCDD60A" box="[736,871,295,320]" captionStart="FIGURES 10 – 12" captionStartId="4.[151,264,1877,1900]" captionTargetBox="[384,1197,208,1841]" captionTargetId="figure@4.[365,1221,193,1856]" captionTargetPageId="4" captionText="FIGURES 10 – 12. Onychogomphus marijanmatoki holotype, anal appendages: (10) lateral view; (11) ventral view; (12) dorsal view." httpUri="https://zenodo.org/record/228029/files/figure.png" pageId="2" pageNumber="183">Figs. 10, 12</figureCitation>
|
||
) dark brown becoming black apically, longer than cerci, deeply divided, the branches slowly dividing until apex, where abruptly out-turned. Accessory genitalia as shown in
|
||
<figureCitation id="13232FBB921FAF6EE23A1C30FEC4D6C2" box="[260,366,367,392]" captionStart="FIGURES 8 – 9" captionStartId="3.[151,264,1267,1290]" captionTargetBox="[165,1410,176,1235]" captionTargetId="figure@3.[151,1436,161,1246]" captionTargetPageId="3" captionText="FIGURES 8 – 9. Onychogomphus marijanmatoki holotype: (8) accessory genitalia, lateral view; (9) terminal segment of genital ligula, ventral view." httpUri="https://zenodo.org/record/228028/files/figure.png" pageId="2" pageNumber="183">Figs. 8–9</figureCitation>
|
||
, terminal segment of genital ligula without cornua.
|
||
</paragraph>
|
||
<caption id="DF6763B6921FAF6EE3A91B49FDF3D101" httpUri="https://zenodo.org/record/228026/files/figure.png" pageId="2" pageNumber="183" targetBox="[151,1436,445,1536]" targetPageId="2">
|
||
<paragraph id="8BA7333E921FAF6EE3A91B49FDF3D101" blockId="2.[151,1436,1558,1611]" pageId="2" pageNumber="183">
|
||
<emphasis id="B96CEF2C921FAF6EE3A91B49FE94D166" bold="true" box="[151,318,1558,1581]" pageId="2" pageNumber="183">FIGURES 2–5.</emphasis>
|
||
<taxonomicName id="4C1848BD921FAF6EE2771B48FD38D167" box="[329,658,1559,1581]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921FAF6EE2771B48FD38D167" box="[329,658,1559,1581]" italics="true" pageId="2" pageNumber="183">Onychogomphus marijanmatoki</emphasis>
|
||
</taxonomicName>
|
||
holotype: (2) head, anterior view; (3) head, dorsal view; (4) pterothorax lateral view; (5) mesepisternum dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6763B6921FAF6EE3A91A2EFB09D0C2" box="[151,1187,1905,1928]" httpUri="https://zenodo.org/record/228027/files/figure.png" pageId="2" pageNumber="183" targetBox="[160,1425,1671,1876]" targetPageId="2">
|
||
<paragraph id="8BA7333E921FAF6EE3A91A2EFB09D0C2" blockId="2.[151,1187,1905,1928]" box="[151,1187,1905,1928]" pageId="2" pageNumber="183">
|
||
<emphasis id="B96CEF2C921FAF6EE3A91A2EFE90D0CD" bold="true" box="[151,314,1905,1928]" pageId="2" pageNumber="183">FIGURES 6–7.</emphasis>
|
||
<taxonomicName id="4C1848BD921FAF6EE27E1A2DFD2CD0C2" box="[320,646,1906,1928]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921FAF6EE27E1A2DFD2CD0C2" box="[320,646,1906,1928]" italics="true" pageId="2" pageNumber="183">Onychogomphus marijanmatoki</emphasis>
|
||
</taxonomicName>
|
||
holotype, wings: (6) left forewing; (7) left hindwing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6763B6921EAF6FE3A919ACFECBD262" httpUri="https://zenodo.org/record/228028/files/figure.png" pageId="3" pageNumber="184" targetBox="[165,1410,176,1235]" targetPageId="3">
|
||
<paragraph id="8BA7333E921EAF6FE3A919ACFECBD262" blockId="3.[151,1436,1267,1320]" pageId="3" pageNumber="184">
|
||
<emphasis id="B96CEF2C921EAF6FE3A919ACFE90D242" bold="true" box="[151,314,1267,1290]" pageId="3" pageNumber="184">FIGURES 8–9.</emphasis>
|
||
<taxonomicName id="4C1848BD921EAF6FE27E19ACFD2FD243" box="[320,645,1267,1289]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921EAF6FE27E19ACFD2FD243" box="[320,645,1267,1289]" italics="true" pageId="3" pageNumber="184">Onychogomphus marijanmatoki</emphasis>
|
||
</taxonomicName>
|
||
holotype: (8) accessory genitalia, lateral view; (9) terminal segment of genital ligula, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7333E921EAF6FE3F9180FFA9FD222" blockId="3.[151,1437,1359,2032]" box="[199,1333,1359,1384]" pageId="3" pageNumber="184">
|
||
<emphasis id="B96CEF2C921EAF6FE3F9180FFEDCD222" bold="true" box="[199,374,1360,1384]" pageId="3" pageNumber="184">Measurements</emphasis>
|
||
[mm]: Hw 25.5, abdomen without anal appendages 28.5, cerci ca 2.75, epiproct ca 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30260B5921EAF6FE3F9182DFCA5D10A" pageId="3" pageNumber="184" type="diagnosis">
|
||
<paragraph id="8BA7333E921EAF6FE3F9182DFCA5D10A" blockId="3.[151,1437,1359,2032]" pageId="3" pageNumber="184">
|
||
<emphasis id="B96CEF2C921EAF6FE3F9182DFEEBD2C1" bold="true" box="[199,321,1394,1419]" pageId="3" pageNumber="184">Diagnosis.</emphasis>
|
||
A small member of the Onychogomphinae, with terminal segment of genital ligula not bearing cornua and epiproct just longer than cerci. Differs from all other named members of the Onychogomphinae for which the male is known and with epiproct not markedly shorter than cerci by the following combination of characters: terminal segment of genital ligula without cornua, seminal vesicle large but not very large and not rounded in profile, epiproct without basal third strongly downturned and without any dorsal spine or tooth visible in lateral view, merely a rounded hump in the basal half.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30260B5921EAF69E3F91B14FEA7D657" lastPageId="5" lastPageNumber="186" pageId="3" pageNumber="184" type="discussion">
|
||
<paragraph id="8BA7333E921EAF6FE3F91B14FDAAD0BA" blockId="3.[151,1437,1359,2032]" pageId="3" pageNumber="184">
|
||
<emphasis id="B96CEF2C921EAF6FE3F91B14FE91D129" bold="true" box="[199,315,1611,1635]" pageId="3" pageNumber="184">Remarks.</emphasis>
|
||
<taxonomicName id="4C1848BD921EAF6FE27C1B13FDA3D129" box="[322,521,1611,1635]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921EAF6FE27C1B13FDA3D129" box="[322,521,1611,1635]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE27C1B13FEF3D129" box="[322,345,1612,1635]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. marijanmatoki
|
||
</emphasis>
|
||
</taxonomicName>
|
||
differs from the species described by Karube & Sasamoto (in press) in markings, details of accessory genitalia and penis structure and of the anal appendages. The male of
|
||
<taxonomicName id="4C1848BD921EAF6FE7DE1B30FE10D1E6" authority="Laidlaw, 1902" authorityName="Laidlaw" authorityYear="1902" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="nigrescens">
|
||
<emphasis id="B96CEF2C921EAF6FE7DE1B30FEA4D1E1" italics="true" pageId="3" pageNumber="184">Onychogomphus nigrescens</emphasis>
|
||
<bibRefCitation id="EF894ECF921EAF6FE2291BCCFE10D1E6" author="Laidlaw" box="[279,442,1683,1708]" pageId="3" pageNumber="184" refString="Laidlaw, F. F. (1902) On a collection of dragonflies made by members of the Skeat expedition in the Malay Peninsula in 1899 - 1900. Proceedings of the Zoological Society of London, 72 (1), 63 - 101, pls. v - vi, excl. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1902. tb 08207. x" type="journal article" year="1902">Laidlaw, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
is not known, but the female, which is known only from peninsular
|
||
<collectingCountry id="F30F73AE921EAF6FE7E21BCCFAECD1E6" box="[1244,1350,1683,1708]" name="Malaysia" pageId="3" pageNumber="184">Malaysia</collectingCountry>
|
||
, differs from
|
||
<taxonomicName id="4C1848BD921EAF6FE3E41BE7FE0CD185" box="[218,422,1719,1743]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921EAF6FE3E41BE7FE0CD185" box="[218,422,1719,1743]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE3E41BE7FF5BD185" box="[218,241,1720,1743]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. marijanmatoki
|
||
</emphasis>
|
||
</taxonomicName>
|
||
in the markings of the thorax, which are stated by
|
||
<bibRefCitation id="EF894ECF921EAF6FE7231BE8FAAAD19A" author="Laidlaw" box="[1053,1280,1719,1744]" pageId="3" pageNumber="184" refString="Laidlaw, F. F. (1902) On a collection of dragonflies made by members of the Skeat expedition in the Malay Peninsula in 1899 - 1900. Proceedings of the Zoological Society of London, 72 (1), 63 - 101, pls. v - vi, excl. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1902. tb 08207. x" type="journal article" year="1902">Laidlaw (1902: 80)</bibRefCitation>
|
||
to be “as in
|
||
<taxonomicName id="4C1848BD921EAF6FE3A91B83FE8AD1B9" box="[151,288,1756,1779]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="geometricus">
|
||
<emphasis id="B96CEF2C921EAF6FE3A91B83FE8AD1B9" box="[151,288,1756,1779]" italics="true" pageId="3" pageNumber="184">geometricus</emphasis>
|
||
</taxonomicName>
|
||
;” the markings of the pterothorax of
|
||
<taxonomicName id="4C1848BD921EAF6FE1EC1B83FBBBD1BE" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[722,1041,1755,1780]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="geometricus">
|
||
<emphasis id="B96CEF2C921EAF6FE1EC1B83FCD5D1B9" box="[722,895,1756,1779]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE1EC1B83FD43D1B9" box="[722,745,1756,1779]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. geometricus
|
||
</emphasis>
|
||
Selys, 1854
|
||
</taxonomicName>
|
||
, are illustrated in
|
||
<bibRefCitation id="EF894ECF921EAF6FE7DD1B84FE8ED052" author="Lieftinck" pageId="3" pageNumber="184" refString="Lieftinck, M. A (1929) Contributions to the dragonfly fauna of the Sondaic area. Tijdschrift voor Entomologie, 72, 109 - 147." type="journal article" year="1929">Lieftinck (1929: 132, fig. 24)</bibRefCitation>
|
||
. Although
|
||
<taxonomicName id="4C1848BD921EAF6FE2A31A5FFD15D052" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[413,703,1791,1816]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saundersii">
|
||
<emphasis id="B96CEF2C921EAF6FE2A31A5FFD99D05D" box="[413,563,1791,1815]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE2A31A5FFE1ED05D" box="[413,436,1792,1815]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. saundersii
|
||
</emphasis>
|
||
Selys, 1854
|
||
</taxonomicName>
|
||
, has been recorded from Sundaland,
|
||
<bibRefCitation id="EF894ECF921EAF6FE7671BA0FF04D076" author="Lieftinck" pageId="3" pageNumber="184" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202." type="journal article" year="1954">Lieftinck (1954: 93, footnote 1)</bibRefCitation>
|
||
noted that there is “a strong element of doubt about the occurrence of the genuine
|
||
<taxonomicName id="4C1848BD921EAF6FE7781A7CFB11D071" box="[1094,1211,1827,1851]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saundersii">
|
||
<emphasis id="B96CEF2C921EAF6FE7781A7CFB11D071" box="[1094,1211,1827,1851]" italics="true" pageId="3" pageNumber="184">saundersii</emphasis>
|
||
</taxonomicName>
|
||
SELYS-HAGEN in Sumatra, and even within the boundaries of the Malaysian subregion.” In any case, the anal appendages and accessory genitalia of
|
||
<taxonomicName id="4C1848BD921EAF6FE2A21A33FD9DD0C9" box="[412,567,1899,1923]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saundersii">
|
||
<emphasis id="B96CEF2C921EAF6FE2A21A33FD9DD0C9" box="[412,567,1899,1923]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE2A21A33FE19D0C9" box="[412,435,1900,1923]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. saundersii
|
||
</emphasis>
|
||
</taxonomicName>
|
||
are illustrated in
|
||
<bibRefCitation id="EF894ECF921EAF6FE0351A34FB03D0CE" author="Selys" box="[779,1193,1899,1924]" pageId="3" pageNumber="184" refString="Selys Longchamps, E. de & Hagen, H. A. (1858) Monographie des Gomphines. Muquardt, Bruxelles - Leipzig & Roret, Paris, 460 pp., pls. 23, excl. http: // dx. doi. org / 10.5962 / bhl. title. 60465" type="book" year="1858">
|
||
Selys
|
||
<bibRefCitation id="EF894ECF921EAF6FE06D1A34FB03D0CE" author="Longchamps" box="[851,1193,1899,1924]" pageId="3" pageNumber="184" refString="Selys Longchamps, E. de & Hagen, H. A. (1858) Monographie des Gomphines. Muquardt, Bruxelles - Leipzig & Roret, Paris, 460 pp., pls. 23, excl. http: // dx. doi. org / 10.5962 / bhl. title. 60465" type="book" year="1858">Longchamps & Hagen (1858</bibRefCitation>
|
||
</bibRefCitation>
|
||
: plate 1, fig. 2), and although the anal appendages are somewhat similar to those of
|
||
<taxonomicName id="4C1848BD921EAF6FE05D1ACFFB81D0ED" box="[867,1067,1935,1959]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C921EAF6FE05D1ACFFB81D0ED" box="[867,1067,1935,1959]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE05D1ACFFCD0D0ED" box="[867,890,1936,1959]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. marijanmatoki
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, they differ in at least the length of the epiproct relative to the cerci and in the shape of the cercus; more importantly the penis of
|
||
<taxonomicName id="4C1848BD921EAF6FE7FA1AEBFAF1D081" box="[1220,1371,1971,1995]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saundersii">
|
||
<emphasis id="B96CEF2C921EAF6FE7FA1AEBFAF1D081" box="[1220,1371,1971,1995]" italics="true" pageId="3" pageNumber="184">
|
||
<collectingCountry id="F30F73AE921EAF6FE7FA1AEBFB71D081" box="[1220,1243,1972,1995]" name="Iceland" pageId="3" pageNumber="184">O</collectingCountry>
|
||
. saundersii
|
||
</emphasis>
|
||
</taxonomicName>
|
||
bears cornua on the terminal segment.
|
||
</paragraph>
|
||
<caption id="DF6763B69219AF68E3A91A0AFF64D0C0" httpUri="https://zenodo.org/record/228029/files/figure.png" pageId="4" pageNumber="185" targetBox="[384,1197,208,1841]" targetPageId="4">
|
||
<paragraph id="8BA7333E9219AF68E3A91A0AFF64D0C0" blockId="4.[151,1436,1877,1930]" pageId="4" pageNumber="185">
|
||
<emphasis id="B96CEF2C9219AF68E3A91A0AFEF9D021" bold="true" box="[151,339,1877,1900]" pageId="4" pageNumber="185">FIGURES 10–12.</emphasis>
|
||
<taxonomicName id="4C1848BD9219AF68E2641A0AFD35D021" box="[346,671,1877,1899]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="marijanmatoki">
|
||
<emphasis id="B96CEF2C9219AF68E2641A0AFD35D021" box="[346,671,1877,1899]" italics="true" pageId="4" pageNumber="185">Onychogomphus marijanmatoki</emphasis>
|
||
</taxonomicName>
|
||
holotype, anal appendages: (10) lateral view; (11) ventral view; (12) dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7333E9218AF69E3F91DC7FEA7D657" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="186">
|
||
A number of genera have been erected within the Asian Onychogomphinae over the decades, but many of these have not been accepted by all authors. I prefer to follow the same conservative course as
|
||
<bibRefCitation id="EF894ECF9218AF69E6131DE3FEDFD7B2" author="Muller" pageId="5" pageNumber="186" refString="Muller, R. A. & Hamalainen, M. (1993) Onychogomphus treadawayi n. sp., eine neue Libellenart von der Insel Busuanga, Philippinen (Odonata: Gomphidae). Entomologische Zeitschrift, Essen, 103 (4), 41 - 45." type="journal article" year="1993">Müller & Hämäläinen (1993)</bibRefCitation>
|
||
and place the new species in
|
||
<taxonomicName id="4C1848BD9218AF69E1D61DBFFC0ED7B2" box="[744,932,224,248]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="186" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96CEF2C9218AF69E1D61DBFFC0ED7B2" box="[744,932,224,248]" italics="true" pageId="5" pageNumber="186">Onychogomphus</emphasis>
|
||
</taxonomicName>
|
||
pending a thorough revision of the entire subfamily.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |