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<document id="5F96110717CC76D3E90C42ADC008A7C6" ID-DOI="10.1051/acarologia/20152151" ID-ISSN="2107-7207" ID-Zenodo-Dep="5401730" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630335126760" checkinUser="marcus" docAuthor="Kontschán, J. &amp; Seeman, O. D." docDate="2015" docId="03A98790FF85FFF82EF0FE4A0D03F823" docLanguage="en" docName="Acarologia.2015.55.1.19-31.pdf" docOrigin="Acarologia 55 (1)" docSource="http://dx.doi.org/10.1051/acarologia/20152151" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Myrmozercon brevipes Berlese 1902" docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FF90FFE8FF86FFF32D6FFFF70911FFBF" masterDocTitle="Rediscovery and redescription of the type species of Myrmozercon, Myrmozercon brevipes Berlese, 1902 (Acari: Mesostigmata: Laelapidae)" masterLastPageNumber="31" masterPageNumber="19" pageNumber="21" updateTime="1699232923288" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="6BE94BE66FD3ACE712FBAA15AF7B9132">Rediscovery and redescription of the type species of Myrmozercon, Myrmozercon brevipes Berlese, 1902 (Acari: Mesostigmata: Laelapidae)</mods:title>
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<mods:namePart id="C61773F0DC8AA5D8C267C6ADDF0AD483">Kontschán, J.</mods:namePart>
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<mods:title id="955108D4CF816C314BAAAFA80A2D93CD">Acarologia</mods:title>
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<treatment id="03A98790FF85FFF82EF0FE4A0D03F823" LSID="urn:lsid:plazi:treatment:03A98790FF85FFF82EF0FE4A0D03F823" httpUri="http://treatment.plazi.org/id/03A98790FF85FFF82EF0FE4A0D03F823" lastPageId="11" lastPageNumber="29" pageId="3" pageNumber="21">
<subSubSection id="C31A650DFF85FFF02EF0FE4A0C21FE6B" box="[927,1328,444,468]" pageId="3" pageNumber="21" type="nomenclature">
<paragraph id="8BBF3686FF85FFF02EF0FE4A0C21FE6B" blockId="3.[927,1328,444,468]" box="[927,1328,444,468]" pageId="3" pageNumber="21">
<heading id="D0F781EAFF85FFF02EF0FE4A0C21FE6B" box="[927,1328,444,468]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="21" reason="4">
<taxonomicName id="4C004D05FF85FFF02EF0FE4A0C21FE6B" authority="Berlese, 1902" authorityName="Berlese" authorityYear="1902" box="[927,1328,444,468]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF85FFF02EF0FE4A0D82FE6B" box="[927,1171,444,468]" italics="true" pageId="3" pageNumber="21">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF85FFF029F5FE4B0C21FE6B" author="Berlese A." box="[1178,1328,444,468]" pageId="3" pageNumber="21" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese, 1902</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="8BBF3686FF85FFF02E55FE040C30FDB5" blockId="3.[826,1429,498,1948]" box="[826,1313,498,522]" pageId="3" pageNumber="21">
<treatmentCitationGroup id="AB1011A8FF85FFF02E55FE040C30FDB5" box="[826,1313,498,522]" pageId="3" pageNumber="21">
<treatmentCitation id="0AA11097FF85FFF02E55FE040C0CFDB5" author="Berlese A." box="[826,1309,498,522]" page="700" pageId="3" pageNumber="21" year="1902">
<taxonomicName id="4C004D05FF85FFF02E55FE040C0CFDB5" authority="Berlese 1902: p. 700" authorityName="Berlese" authorityPageNumber="700" authorityYear="1902" box="[826,1309,498,522]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF85FFF02E55FE040D3FFDB5" box="[826,1070,498,522]" italics="true" pageId="3" pageNumber="21">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF85FFF0295AFE050DD6FDB5" author="Berlese A." box="[1077,1223,498,522]" pageId="3" pageNumber="21" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
: p. 700
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBF3686FF85FFF02E55FDE10D9DFDCC" blockId="3.[826,1429,498,1948]" pageId="3" pageNumber="21">
<treatmentCitationGroup id="AB1011A8FF85FFF02E55FDE10D9DFDCC" pageId="3" pageNumber="21">
<treatmentCitation id="0AA11097FF85FFF02E55FDE10C24FD92" author="Berlese A." box="[826,1333,533,557]" page="313" pageId="3" pageNumber="21" year="1904">
<taxonomicName id="4C004D05FF85FFF02E55FDE10C24FD92" authority=": Berlese 1904: p. 313" authorityName="Berlese" authorityPageNumber="313" authorityYear="1904" box="[826,1333,533,557]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF85FFF02E55FDE10D21FD92" box="[826,1072,533,557]" italics="true" pageId="3" pageNumber="21">Myrmozercon brevipes</emphasis>
:
<bibRefCitation id="EF914B77FF85FFF0292CFDE20DC9FD92" author="Berlese A." box="[1091,1240,533,557]" pageId="3" pageNumber="21" pagination="299 - 474" refId="ref4419" refString="Berlese A. 1904 - Illustrazione iconografica degli Acari mirmecofili - Redia, 1: 299 - 474." type="journal article" year="1904">Berlese 1904</bibRefCitation>
: p. 313
</taxonomicName>
</treatmentCitation>
(female chelicera), p. 314 (male chelicera), p. 445 (redescription). Table 19, Figs. 169-172.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="8BBF3686FF85FFF02E0FFD7D0DE6FCCB" blockId="3.[826,1429,498,1948]" pageId="3" pageNumber="21">
Material examined — Four females and
<specimenCount id="9D06FD0FFF85FFF02805FD7C0A62FD7A" pageId="3" pageNumber="21" type="male">one male</specimenCount>
(
<collectionCode id="ED11AE43FF85FFF02EEAFD5A0AF4FD7A" box="[901,997,685,709]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="21">HNHM</collectionCode>
).
<materialsCitation id="3B683CDBFF85FFF02E94FD5A0AD1FCCB" collectingDate="2011-05-06" collectionCode="T" country="Greece" county="Trifos" elevation="330" latitude="38.8066" location="Aetolia-Acarnania" longLatPrecision="1" longitude="21.094166" municipality="Akarnanika Mts" pageId="3" pageNumber="21" specimenCount="1">
<collectingCountry id="F3177616FF85FFF02E94FD5A0D5FFD7A" box="[1019,1102,685,709]" name="Greece" pageId="3" pageNumber="21">Greece</collectingCountry>
,
<location id="8EDF605DFF85FFF02933FD5A0C25FD7A" LSID="urn:lsid:plazi:treatment:03A98790FF85FFF82EF0FE4A0D03F823:8EDF605DFF85FFF02933FD5A0C25FD7A" box="[1116,1332,685,709]" country="Greece" county="Trifos" latitude="38.8066" longLatPrecision="1" longitude="21.094166" municipality="Akarnanika Mts" name="Aetolia-Acarnania" pageId="3" pageNumber="21">Aetolia-Acarnania</location>
peripheral unit,
<collectingMunicipality id="6BDBACFCFF85FFF02ED8FD270D6FFD57" box="[951,1150,720,744]" pageId="3" pageNumber="21">Akarnanika Mts</collectingMunicipality>
,
<collectingCounty id="62DE4E0AFF85FFF029FEFD270DC7FD57" box="[1169,1238,720,744]" pageId="3" pageNumber="21">Trifos</collectingCounty>
, small artificial pond and its shore vegetation S of the village,
<geoCoordinate id="EE345041FF85FFF02E55FCE10ADEFC91" box="[826,975,790,814]" degrees="38" direction="north" minutes="48.396" orientation="latitude" pageId="3" pageNumber="21" precision="1" value="38.8066">38°48.396 N</geoCoordinate>
,
<geoCoordinate id="EE345041FF85FFF02EB3FCE10D7DFC91" box="[988,1132,790,814]" degrees="21" direction="east" minutes="05.650" orientation="longitude" pageId="3" pageNumber="21" precision="1" value="21.094166">21°05.650 E</geoCoordinate>
,
<quantity id="4CF89B63FF85FFF02917FCE10DD0FC91" box="[1144,1217,790,814]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="21" unit="m" value="330.0">
<elevation id="002DD1B5FF85FFF02917FCE10DD0FC91" box="[1144,1217,790,814]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="21" unit="m" value="330.0">330 m</elevation>
</quantity>
,
<date id="FFBE1046FF85FFF029A1FCE10C48FC91" box="[1230,1369,790,815]" pageId="3" pageNumber="21" value="2011-05-06">
<collectingDate id="EFFAE9AEFF85FFF029A1FCE10C48FC91" box="[1230,1369,790,815]" pageId="3" pageNumber="21" value="2011-05-06">6 May 2011</collectingDate>
</date>
, nest of
<taxonomicName id="4C004D05FF85FFF02E38FCCD0D89FCEE" authority="Latreille" authorityName="Latreille" box="[855,1176,825,849]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="erraticum">
<emphasis id="B974EA94FF85FFF02E38FCCD0D3EFCEE" box="[855,1071,826,849]" italics="true" pageId="3" pageNumber="21">Tapinoma erraticum</emphasis>
Latreille
</taxonomicName>
, coll. J. KontschAEn, D. MurAEnyi,
<collectionCode id="ED11AE43FF85FFF02EC1FCAA0AD1FCCB" box="[942,960,861,884]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="21">T</collectionCode>
</materialsCitation>
. Szederjesi and Zs. UjvAEri.
</paragraph>
<paragraph id="8BBF3686FF85FFF02E0FFC7C0C08FB8F" blockId="3.[826,1429,498,1948]" pageId="3" pageNumber="21">
Five females,
<specimenCount id="9D06FD0FFF85FFF02967FC7B0D66FC1C" box="[1032,1143,907,931]" pageId="3" pageNumber="21" type="male">one male</specimenCount>
(
<collectionCode id="ED11AE43FF85FFF029E6FC7C0DF7FC1C" box="[1161,1254,907,931]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="21">HNHM</collectionCode>
) and
<specimenCount id="9D06FD0FFF85FFF0285FFC7C0A6FFC79" pageId="3" pageNumber="21" type="female">three females</specimenCount>
(
<collectionCode id="ED11AE43FF85FFF02EFCFC580AD4FC78" box="[915,965,943,967]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="3" pageNumber="21" type="Museum">QM</collectionCode>
).
<materialsCitation id="3B683CDBFF85FFF02EB1FC580AF3FB90" collectingDate="2011-05-06" collectionCode="T" country="Greece" elevation="50" latitude="38.722717" location="Lefkada" longLatPrecision="1" longitude="20.690067" municipality="Rahi" pageId="3" pageNumber="21" specimenCount="1">
<collectingCountry id="F3177616FF85FFF02EB1FC580D20FC78" box="[990,1073,943,967]" name="Greece" pageId="3" pageNumber="21">Greece</collectingCountry>
, Ionian Islands,
<location id="8EDF605DFF85FFF0286CFC590C73FC79" LSID="urn:lsid:plazi:treatment:03A98790FF85FFF82EF0FE4A0D03F823:8EDF605DFF85FFF0286CFC590C73FC79" box="[1283,1378,942,966]" country="Greece" latitude="38.722717" longLatPrecision="1" longitude="20.690067" municipality="Rahi" name="Lefkada" pageId="3" pageNumber="21">Lefkada</location>
peripheral unit,
<collectingMunicipality id="6BDBACFCFF85FFF02EB0FC260D09FC56" box="[991,1048,977,1001]" pageId="3" pageNumber="21">Rahi</collectingMunicipality>
,
<geoCoordinate id="EE345041FF85FFF0294BFC260DA3FC56" box="[1060,1202,977,1001]" degrees="38" direction="north" minutes="43.363" orientation="latitude" pageId="3" pageNumber="21" precision="1" value="38.722717">38°43.363 N</geoCoordinate>
,
<geoCoordinate id="EE345041FF85FFF029D0FC250C56FC56" box="[1215,1351,978,1002]" degrees="20" direction="east" minutes="41.404" orientation="longitude" pageId="3" pageNumber="21" precision="1" value="20.690067">20°41.404 E</geoCoordinate>
,
<quantity id="4CF89B63FF85FFF0283BFC260C81FC56" box="[1364,1424,977,1001]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="21" unit="m" value="50.0">
<elevation id="002DD1B5FF85FFF0283BFC260C81FC56" box="[1364,1424,977,1001]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="21" unit="m" value="50.0">50 m</elevation>
</quantity>
,
<date id="FFBE1046FF85FFF02E55FC030ADEFBB2" box="[826,975,1012,1037]" pageId="3" pageNumber="21" value="2011-05-06">
<collectingDate id="EFFAE9AEFF85FFF02E55FC030ADEFBB2" box="[826,975,1012,1037]" pageId="3" pageNumber="21" value="2011-05-06">06 May 2011</collectingDate>
</date>
, nest of
<taxonomicName id="4C004D05FF85FFF02940FC020DA5FBB3" box="[1071,1204,1013,1036]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="erraticum">
<emphasis id="B974EA94FF85FFF02940FC020DA5FBB3" box="[1071,1204,1013,1036]" italics="true" pageId="3" pageNumber="21">T. erraticum</emphasis>
</taxonomicName>
, coll. J. KontschAEn, D. MurAEnyi,
<collectionCode id="ED11AE43FF85FFF02EBFFBEF0AF3FB90" box="[976,994,1048,1071]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="21">T</collectionCode>
</materialsCitation>
. Szederjesi and Zs. UjvAEri.
</paragraph>
<paragraph id="8BBF3686FF85FFF02E0FFBB10C3DF930" blockId="3.[826,1429,498,1948]" pageId="3" pageNumber="21">
Diagnosis (based on female) — Dorsal shield densely hypertrichous, setal length 20 40; posterior margin of shield rounded, not truncated; soft cuticle posterior to shield hypertrichous. Sternal shield horseshoe-shaped (inverted U-shape); setae
<emphasis id="B974EA94FF85FFF02E0BFB010A94FAB1" box="[868,901,1270,1294]" italics="true" pageId="3" pageNumber="21">st4</emphasis>
present; epigynal shield sculpturing lineatereticulate, shield bearing setae
<emphasis id="B974EA94FF85FFF029C7FAEE0DD8FA8F" box="[1192,1225,1305,1328]" italics="true" pageId="3" pageNumber="21">st5</emphasis>
only; one pair of metapodal shields present; anal shield peltate. Palp setal count (trochanter to tibia) 1-5-5-11; subcapitular groove with 8 rows of denticles, with one large denticle per row; palp coxal seta absent. Chelicerae highly modified, terminating in small membranous flange; fixed digit absent. All legs short, leg I distinctly shorter (280 295) than all other legs (leg II 370 410; leg III 385 425; leg IV 410 445); setal counts (coxa to tibia) for legs I-IV: 2-4-8-8-8, 2-4-8-8- 7, 2-5-6-10-9, 2-5-8-8-7; all leg setae simple.
</paragraph>
<paragraph id="8BBF3686FF85FFF02E0FF9510C23F901" blockId="3.[826,1429,498,1948]" box="[864,1330,1702,1726]" pageId="3" pageNumber="21">
Description —
<emphasis id="B974EA94FF85FFF0297FF9510D4AF901" box="[1040,1115,1702,1726]" italics="true" pageId="3" pageNumber="21">Female</emphasis>
(
<figureCitation id="133B2A03FF85FFF02904F9510DC1F901" box="[1131,1232,1702,1726]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStart-4="FIGURE" captionStart-5="FIGURE" captionStartId-0="2.[171,183,1751,1770]" captionStartId-1="4.[159,171,1841,1860]" captionStartId-2="5.[159,171,1922,1941]" captionStartId-3="6.[159,171,1543,1562]" captionStartId-4="7.[159,171,1823,1842]" captionStartId-5="8.[159,171,1855,1874]" captionTargetBox-0="[314,1302,491,1688]" captionTargetBox-1="[171,1432,393,1797]" captionTargetBox-2="[294,1239,307,1890]" captionTargetBox-3="[240,1377,639,1443]" captionTargetBox-4="[164,1587,346,1783]" captionTargetBox-5="[317,1279,326,1817]" captionTargetId-0="figure-5@2.[265,1323,472,1719]" captionTargetId-1="figure-5@4.[159,1444,382,1809]" captionTargetId-2="figure-4@5.[279,1307,302,1890]" captionTargetId-3="figure-5@6.[196,1391,624,1511]" captionTargetId-4="figure-4@7.[159,1662,344,1791]" captionTargetId-5="figure-5@8.[288,1300,312,1824]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1: Myrmozercon brevipes Berlese 1902, female: Dorsal shield. Lyrifissures, pores and sigillae are shown on the right-hand side." captionText-1="FIGURE 2: Scanning electron micrographs of Myrmozercon brevipes Berlese 1902, female: A Dorsum; B Venter; C Ventral gnathosoma." captionText-2="FIGURE 3: Myrmozercon brevipes Berlese 1902, female: A Venter; extensive setation of opisthogaster partially illustrated (mp = metapo-" captionText-3="FIGURE 4: Myrmozercon brevipes Berlese 1902, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen)." captionText-4="FIGURE 5: Myrmozercon brevipes Berlese 1902, female, chelicerae from three different specimens, with two interpretations of cheliceral morphology labelled (see Remarks). cs = vestigial cheliceral seta; dps = distal pseudosegment of second cheliceral digit; ly = lyrifissure; md = movable digit; pps = proximal pseudosegment of second cheliceral digit;? = novel structure." captionText-5="FIGURE 6: Myrmozercon brevipes Berlese 1902, female: A Tarsus I (circle = blunt-tipped, ribbed seta; diamond = minute seta; x = abruptly tapering seta); B Leg I, coxa to tibia; C Leg II, coxa to tarsus; D Leg III, trochanter to tarsus; E Leg IV, trochanter to tarsus;" figureDoi-0="http://doi.org/10.5281/zenodo.5401732" figureDoi-1="http://doi.org/10.5281/zenodo.5401734" figureDoi-2="http://doi.org/10.5281/zenodo.5401738" figureDoi-3="http://doi.org/10.5281/zenodo.5401740" figureDoi-4="http://doi.org/10.5281/zenodo.5401742" figureDoi-5="http://doi.org/10.5281/zenodo.5401744" httpUri-0="https://zenodo.org/record/5401732/files/figure.png" httpUri-1="https://zenodo.org/record/5401734/files/figure.png" httpUri-2="https://zenodo.org/record/5401738/files/figure.png" httpUri-3="https://zenodo.org/record/5401740/files/figure.png" httpUri-4="https://zenodo.org/record/5401742/files/figure.png" httpUri-5="https://zenodo.org/record/5401744/files/figure.png" pageId="3" pageNumber="21">Figs. 1-6</figureCitation>
; n = 12).
</paragraph>
<paragraph id="8BBF3686FF85FFFB2E0FF9220890F835" blockId="3.[826,1429,498,1948]" lastBlockId="8.[159,1429,1855,1930]" lastPageId="8" lastPageNumber="26" pageId="3" pageNumber="21">
Dorsal idiosoma (
<figureCitation id="133B2A03FF85FFF02959F9220D9DF952" box="[1078,1164,1749,1773]" captionStart="FIGURE" captionStartId="2.[171,183,1751,1770]" captionTargetBox="[314,1302,491,1688]" captionTargetId="figure-5@2.[265,1323,472,1719]" captionTargetPageId="2" captionText="FIGURE 1: Myrmozercon brevipes Berlese 1902, female: Dorsal shield. Lyrifissures, pores and sigillae are shown on the right-hand side." figureDoi="http://doi.org/10.5281/zenodo.5401732" httpUri="https://zenodo.org/record/5401732/files/figure.png" pageId="3" pageNumber="21">Figs. 1</figureCitation>
,
<figureCitation id="133B2A03FF85FFF029F2F9220DD1F952" box="[1181,1216,1749,1773]" captionStart="FIGURE" captionStartId="4.[159,171,1841,1860]" captionTargetBox="[171,1432,393,1797]" captionTargetId="figure-5@4.[159,1444,382,1809]" captionTargetPageId="4" captionText="FIGURE 2: Scanning electron micrographs of Myrmozercon brevipes Berlese 1902, female: A Dorsum; B Venter; C Ventral gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.5401734" httpUri="https://zenodo.org/record/5401734/files/figure.png" pageId="3" pageNumber="21">2A</figureCitation>
) — Dorsal shield length 640 760, width 550 640, subcircular, cuticle smooth, with several hundred fine setae, length 20 40. Most marginal setae short; dorsal marginal setae become longer posteriorly (longest 50 75) on shield and soft cuticle. Dorsal shield with 16 pairs of dal platelet); B Tritosternum. F Ambulacrum III.
</paragraph>
<caption id="DF7F660EFF82FFF72DF0F8C60C87F8FC" ID-DOI="http://doi.org/10.5281/zenodo.5401734" ID-Zenodo-Dep="5401734" box="[159,1430,1840,1860]" httpUri="https://zenodo.org/record/5401734/files/figure.png" pageId="4" pageNumber="22" startId="4.[159,171,1841,1860]" targetBox="[171,1432,393,1797]" targetPageId="4">
<paragraph id="8BBF3686FF82FFF72DF0F8C60C87F8FC" blockId="4.[159,1430,1840,1860]" box="[159,1430,1840,1860]" pageId="4" pageNumber="22">
FIGURE 2: Scanning electron micrographs of
<taxonomicName id="4C004D05FF82FFF72F2FF8C60A6DF8FC" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[576,892,1840,1859]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF82FFF72F2FF8C60A12F8FC" box="[576,771,1840,1859]" italics="true" pageId="4" pageNumber="22">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF82FFF72E68F8C70A6DF8FC" author="Berlese A." box="[775,892,1840,1859]" pageId="4" pageNumber="22" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, female: A Dorsum; B Venter; C Ventral gnathosoma.
</paragraph>
</caption>
<caption id="DF7F660EFF83FFF62DF0F8750C84F82A" ID-DOI="http://doi.org/10.5281/zenodo.5401738" ID-Zenodo-Dep="5401738" box="[159,1429,1921,1941]" httpUri="https://zenodo.org/record/5401738/files/figure.png" pageId="5" pageNumber="23" startId="5.[159,171,1922,1941]" targetBox="[294,1239,307,1890]" targetPageId="5">
<paragraph id="8BBF3686FF83FFF62DF0F8750C84F82A" blockId="5.[159,1429,1921,1965]" box="[159,1429,1921,1941]" pageId="5" pageNumber="23">
FIGURE 3:
<taxonomicName id="4C004D05FF83FFF62C6AF8750B52F82B" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[261,579,1921,1940]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF83FFF62C6AF87508D8F82B" box="[261,457,1921,1940]" italics="true" pageId="5" pageNumber="23">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF83FFF62CA1F8760B52F82B" author="Berlese A." box="[462,579,1921,1940]" pageId="5" pageNumber="23" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, female: A Venter; extensive setation of opisthogaster partially illustrated (mp = metapo-
</paragraph>
</caption>
<caption id="DF7F660EFF80FFF52DF0F9F00895F9EE" ID-DOI="http://doi.org/10.5281/zenodo.5401740" ID-Zenodo-Dep="5401740" httpUri="https://zenodo.org/record/5401740/files/figure.png" pageId="6" pageNumber="24" startId="6.[159,171,1543,1562]" targetBox="[240,1377,639,1443]" targetPageId="6">
<paragraph id="8BBF3686FF80FFF52DF0F9F00895F9EE" blockId="6.[159,1429,1543,1617]" pageId="6" pageNumber="24">
FIGURE 4:
<taxonomicName id="4C004D05FF80FFF52C6AF9F00B52F9A5" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[261,579,1543,1562]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="24" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF80FFF52C6AF9F008D9F9A5" box="[261,456,1543,1562]" italics="true" pageId="6" pageNumber="24">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF80FFF52CA2F9F00B52F9A5" author="Berlese A." box="[461,579,1543,1562]" pageId="6" pageNumber="24" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen).
</paragraph>
</caption>
<caption id="DF7F660EFF81FFF42DF0F8E80D80F8D6" ID-DOI="http://doi.org/10.5281/zenodo.5401742" ID-Zenodo-Dep="5401742" httpUri="https://zenodo.org/record/5401742/files/figure.png" pageId="7" pageNumber="25" startId="7.[159,171,1823,1842]" targetBox="[164,1587,346,1783]" targetPageId="7">
<paragraph id="8BBF3686FF81FFF42DF0F8E80D80F8D6" blockId="7.[159,1429,1822,1897]" pageId="7" pageNumber="25">
FIGURE 5:
<taxonomicName id="4C004D05FF81FFF42C66F8E80B5AF88D" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[265,587,1822,1842]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF81FFF42C66F8E808DFF88E" box="[265,462,1822,1841]" italics="true" pageId="7" pageNumber="25">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF81FFF42CBBF8E90B5AF88D" author="Berlese A." box="[468,587,1822,1842]" pageId="7" pageNumber="25" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, female, chelicerae from three different specimens, with two interpretations of cheliceral morphology labelled (see Remarks). cs = vestigial cheliceral seta; dps = distal pseudosegment of second cheliceral digit; ly = lyrifissure; md = movable digit; pps = proximal pseudosegment of second cheliceral digit;? = novel structure.
</paragraph>
</caption>
<caption id="DF7F660EFF8EFFFB2DF0F8C80C7BF8D1" ID-DOI="http://doi.org/10.5281/zenodo.5401744" ID-Zenodo-Dep="5401744" httpUri="https://zenodo.org/record/5401744/files/figure.png" pageId="8" pageNumber="26" startId="8.[159,171,1855,1874]" targetBox="[317,1279,326,1817]" targetPageId="8">
<paragraph id="8BBF3686FF8EFFFB2DF0F8C80C7BF8D1" blockId="8.[159,1429,1855,1930]" pageId="8" pageNumber="26">
FIGURE 6:
<taxonomicName id="4C004D05FF8EFFFB2C6CF8B70B2EF8ED" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[259,575,1855,1874]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="26" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8EFFFB2C6CF8B708D7F8ED" box="[259,454,1855,1874]" italics="true" pageId="8" pageNumber="26">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF8EFFFB2CA5F8C80B2EF8ED" author="Berlese A." box="[458,575,1855,1874]" pageId="8" pageNumber="26" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, female: A Tarsus I (circle = blunt-tipped, ribbed seta; diamond = minute seta; x = abruptly tapering seta); B Leg I, coxa to tibia; C Leg II, coxa to tarsus; D Leg III, trochanter to tarsus; E Leg IV, trochanter to tarsus;
</paragraph>
</caption>
<paragraph id="8BBF3686FF8FFFFA2DF0FEC00BE7FE07" blockId="9.[159,761,311,440]" pageId="9" pageNumber="27">visible pore-like structures (ten slit-like lyrifissures; all others rounded, surrounded by lacunae). Sigillae small arranged in medial groups on podontal region; groups on opisthonotal region more lateral.</paragraph>
<paragraph id="8BBF3686FF8FFFFA2DAAFE220BAEF993" blockId="9.[159,762,469,1580]" pageId="9" pageNumber="27">
Ventral idiosoma (
<figureCitation id="133B2A03FF8FFFFA2CF2FE220B11FE52" box="[413,512,469,493]" captionStart="FIGURE" captionStartId="4.[159,171,1841,1860]" captionTargetBox="[171,1432,393,1797]" captionTargetId="figure-5@4.[159,1444,382,1809]" captionTargetPageId="4" captionText="FIGURE 2: Scanning electron micrographs of Myrmozercon brevipes Berlese 1902, female: A Dorsum; B Venter; C Ventral gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.5401734" httpUri="https://zenodo.org/record/5401734/files/figure.png" pageId="9" pageNumber="27">Figs. 2B</figureCitation>
,
<figureCitation id="133B2A03FF8FFFFA2F63FE220B3FFE52" box="[524,558,469,493]" captionStart="FIGURE" captionStartId="5.[159,171,1922,1941]" captionTargetBox="[294,1239,307,1890]" captionTargetId="figure-4@5.[279,1307,302,1890]" captionTargetPageId="5" captionText="FIGURE 3: Myrmozercon brevipes Berlese 1902, female: A Venter; extensive setation of opisthogaster partially illustrated (mp = metapo-" figureDoi="http://doi.org/10.5281/zenodo.5401738" httpUri="https://zenodo.org/record/5401738/files/figure.png" pageId="9" pageNumber="27">3A</figureCitation>
) — Tritosternum (
<figureCitation id="133B2A03FF8FFFFA2DC8FE0F0816FDAF" box="[167,263,504,528]" captionStart="FIGURE" captionStartId="5.[159,171,1922,1941]" captionTargetBox="[294,1239,307,1890]" captionTargetId="figure-4@5.[279,1307,302,1890]" captionTargetPageId="5" captionText="FIGURE 3: Myrmozercon brevipes Berlese 1902, female: A Venter; extensive setation of opisthogaster partially illustrated (mp = metapo-" figureDoi="http://doi.org/10.5281/zenodo.5401738" httpUri="https://zenodo.org/record/5401738/files/figure.png" pageId="9" pageNumber="27">Fig. 3B</figureCitation>
) with short laciniae, length 40 45, unfused, laciniae each with a pair of tiny spines 14 16 from base, base hyaline, evanescent. Presternal area membranous, presternal shields absent. Sternal shield narrow, highly concave, horse-shoe shaped, medial length 30 34, maximum length 240 260, length to seta
<emphasis id="B974EA94FF8FFFFA2CFDFD3D08A2FD5D" box="[402,435,714,738]" italics="true" pageId="9" pageNumber="27">st4</emphasis>
160 185, width at level of setae
<emphasis id="B974EA94FF8FFFFA2D8DFD190812FCB9" box="[226,259,750,774]" italics="true" pageId="9" pageNumber="27">st2</emphasis>
125 140, width at level of
<emphasis id="B974EA94FF8FFFFA2F35FD190B6AFCB9" box="[602,635,750,774]" italics="true" pageId="9" pageNumber="27">st4</emphasis>
250 270, ratio medial length / length to st4 0.2; sternal shield smooth, bearing smooth slender setae
<emphasis id="B974EA94FF8FFFFA2F31FCC30B84FCF3" box="[606,661,820,844]" italics="true" pageId="9" pageNumber="27">st1-4</emphasis>
(39, 35 37, 38 39, 43 45, respectively) and lyrifissures
<emphasis id="B974EA94FF8FFFFA2FA0FCA009BDFC2D" italics="true" pageId="9" pageNumber="27">iv1- 3</emphasis>
. Endopodal shields absent, or reduced and fused to exopodal shield around posterior margin of coxa IV. Epigynal shield (full length 410 420; 180 195 from
<emphasis id="B974EA94FF8FFFFA2DB0FC1409EEFC45" box="[223,255,995,1018]" italics="true" pageId="9" pageNumber="27">st5</emphasis>
to posterior tip) with well-defined broadly rounded anterior margin, with longitudinal reticulation, pointed posteriorly, maximum width 265 270; setae
<emphasis id="B974EA94FF8FFFFA2C4FFBBA0850FBDB" box="[288,321,1101,1124]" italics="true" pageId="9" pageNumber="27">st5</emphasis>
30 35; lyrifissures
<emphasis id="B974EA94FF8FFFFA2F2BFBBA0B77FBDB" box="[580,614,1101,1124]" italics="true" pageId="9" pageNumber="27">iv5</emphasis>
not present on shield (nor detected in soft cuticle). Metapodal platelets elongate, weakly formed (55 63 × 12 14). Peritremal shield small, smooth, free posteriorly and surrounding only posterior half of peritreme, not fused to dorsal shield, bearing one gland pore; peritreme extending to level of anterior coxa III. Anal shield peltate (length 150 170 × width 105 110), shield with few longitudinal reticulations, with post-anal seta (36 37) shorter than para-anal setae (100 105); cribrum a dense field of spicules extending from just behind post-anal seta to posterior margin; sigillae absent. Soft cuticle hypertrichous, with 350 400 setae, setal length 30 45.
</paragraph>
<paragraph id="8BBF3686FF8FFFFA2DAAF9BE0A6FFC1C" blockId="9.[159,762,1609,1948]" lastBlockId="9.[826,1429,311,931]" pageId="9" pageNumber="27">
Gnathosoma (
<figureCitation id="133B2A03FF8FFFFA2C05F9BE08DDF9DE" box="[362,460,1609,1633]" captionStart="FIGURE" captionStartId="4.[159,171,1841,1860]" captionTargetBox="[171,1432,393,1797]" captionTargetId="figure-5@4.[159,1444,382,1809]" captionTargetPageId="4" captionText="FIGURE 2: Scanning electron micrographs of Myrmozercon brevipes Berlese 1902, female: A Dorsum; B Venter; C Ventral gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.5401734" httpUri="https://zenodo.org/record/5401734/files/figure.png" pageId="9" pageNumber="27">Figs. 2C</figureCitation>
,
<figureCitation id="133B2A03FF8FFFFA2CB8F9BE08F4F9DE" box="[471,485,1609,1633]" captionStart="FIGURE" captionStartId="6.[159,171,1543,1562]" captionTargetBox="[240,1377,639,1443]" captionTargetId="figure-5@6.[196,1391,624,1511]" captionTargetPageId="6" captionText="FIGURE 4: Myrmozercon brevipes Berlese 1902, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen)." figureDoi="http://doi.org/10.5281/zenodo.5401740" httpUri="https://zenodo.org/record/5401740/files/figure.png" pageId="9" pageNumber="27">4</figureCitation>
) — Gnathotectum (
<figureCitation id="133B2A03FF8FFFFA2FA2F9BE09ACF93B" captionStart="FIGURE" captionStartId="6.[159,171,1543,1562]" captionTargetBox="[240,1377,639,1443]" captionTargetId="figure-5@6.[196,1391,624,1511]" captionTargetPageId="6" captionText="FIGURE 4: Myrmozercon brevipes Berlese 1902, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen)." figureDoi="http://doi.org/10.5281/zenodo.5401740" httpUri="https://zenodo.org/record/5401740/files/figure.png" pageId="9" pageNumber="27">Fig. 4B</figureCitation>
) with anterior margin smooth, rounded. Subcapitulum (
<figureCitation id="133B2A03FF8FFFFA2C46F978089AF918" box="[297,395,1679,1703]" captionStart="FIGURE" captionStartId="6.[159,171,1543,1562]" captionTargetBox="[240,1377,639,1443]" captionTargetId="figure-5@6.[196,1391,624,1511]" captionTargetPageId="6" captionText="FIGURE 4: Myrmozercon brevipes Berlese 1902, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen)." figureDoi="http://doi.org/10.5281/zenodo.5401740" httpUri="https://zenodo.org/record/5401740/files/figure.png" pageId="9" pageNumber="27">Fig. 4A</figureCitation>
) with eight deutosternal rows each comprising a single denticle; corniculi (21 24) hyaline, adpressed to anterior margin of subcapitulum, tips acuminate directed medially; subcapitulum not extending medially beyond corniculi; internal malae not detectable or perhaps absent; labrum (16 30), reduced, extending to level of anterior margin of palp trochanter to mid-femur; hy- postomal setae fine,
<emphasis id="B974EA94FF8FFFFA2947FEC00D52FEF0" box="[1064,1091,311,335]" italics="true" pageId="9" pageNumber="27">h1</emphasis>
longer (27) than outer posterior setae
<emphasis id="B974EA94FF8FFFFA2EC2FEAD0AD9FECD" box="[941,968,346,370]" italics="true" pageId="9" pageNumber="27">h2</emphasis>
(15), inner posterior setae
<emphasis id="B974EA94FF8FFFFA2990FEAD0C08FECD" box="[1279,1305,346,370]" italics="true" pageId="9" pageNumber="27">h3</emphasis>
(16); illustrated specimen with duplication of seta
<emphasis id="B974EA94FF8FFFFA287FFE8A0C3AFE2A" box="[1296,1323,381,405]" italics="true" pageId="9" pageNumber="27">h1</emphasis>
on righthand side, scanning electron micrographed specimen lacking seta
<emphasis id="B974EA94FF8FFFFA2960FE340D3BFE64" box="[1039,1066,451,475]" italics="true" pageId="9" pageNumber="27">h2</emphasis>
on right-hand side (
<figureCitation id="133B2A03FF8FFFFA2849FE340C99FE64" box="[1318,1416,451,475]" captionStart="FIGURE" captionStartId="4.[159,171,1841,1860]" captionTargetBox="[171,1432,393,1797]" captionTargetId="figure-5@4.[159,1444,382,1809]" captionTargetPageId="4" captionText="FIGURE 2: Scanning electron micrographs of Myrmozercon brevipes Berlese 1902, female: A Dorsum; B Venter; C Ventral gnathosoma." figureDoi="http://doi.org/10.5281/zenodo.5401734" httpUri="https://zenodo.org/record/5401734/files/figure.png" pageId="9" pageNumber="27">Fig. 2C</figureCitation>
); palpcoxal setae absent; setae
<emphasis id="B974EA94FF8FFFFA29FEFE110DBDFE41" box="[1169,1196,486,510]" italics="true" pageId="9" pageNumber="27">h2</emphasis>
40 42 apart.
<emphasis id="B974EA94FF8FFFFA2835FE110C84FE41" box="[1370,1429,486,510]" italics="true" pageId="9" pageNumber="27">Palps</emphasis>
(
<figureCitation id="133B2A03FF8FFFFA2E2DFDFE0A88FD9E" box="[834,921,521,545]" captionStart="FIGURE" captionStartId="6.[159,171,1543,1562]" captionTargetBox="[240,1377,639,1443]" captionTargetId="figure-5@6.[196,1391,624,1511]" captionTargetPageId="6" captionText="FIGURE 4: Myrmozercon brevipes Berlese 1902, female: A Subcapitulum (c = corniculus; lb = labrum); B Gnathotectum (ch = chelicera; gt = gnathotectum; p = palp); C Palp (x = tarsal seta; blunt-tipped sensory setae are coloured black); D Detail of palp apotele (ap) (different specimen)." figureDoi="http://doi.org/10.5281/zenodo.5401740" httpUri="https://zenodo.org/record/5401740/files/figure.png" pageId="9" pageNumber="27">Figs. 4</figureCitation>
C-D): length 92 94 with simple setae except lateral setae on palp femur and genu thickened, dorsal setae on palp femur, genu and tibia thickened; femur without dorsal anterolateral spine; tarsus placed entirely ventrally; setation of palp segments from trochanter to tarsus: 1-5-5-11-10, tibia with two apical round-tipped hollow tibial setae on small tubercles, three tarsal setae of similar form; palp tarsal claw two-tined, with tines tapered, claw flanked by small cusp, ventral tine shorter (2 3 from fork) than dorsal tine (6 from fork, 10 11 from base).
</paragraph>
<paragraph id="8BBF3686FF8FFFFA2E0FFC260DDBFB60" blockId="9.[826,1429,977,1247]" pageId="9" pageNumber="27">
Chelicerae (
<figureCitation id="133B2A03FF8FFFFA2E86FC260D24FC56" box="[1001,1077,977,1001]" captionStart="FIGURE" captionStartId="7.[159,171,1823,1842]" captionTargetBox="[164,1587,346,1783]" captionTargetId="figure-4@7.[159,1662,344,1791]" captionTargetPageId="7" captionText="FIGURE 5: Myrmozercon brevipes Berlese 1902, female, chelicerae from three different specimens, with two interpretations of cheliceral morphology labelled (see Remarks). cs = vestigial cheliceral seta; dps = distal pseudosegment of second cheliceral digit; ly = lyrifissure; md = movable digit; pps = proximal pseudosegment of second cheliceral digit;? = novel structure." figureDoi="http://doi.org/10.5281/zenodo.5401742" httpUri="https://zenodo.org/record/5401742/files/figure.png" pageId="9" pageNumber="27">Figs. 5</figureCitation>
A-C) — Second cheliceral segment seemingly split into two segments (see Remarks); proximal pseudosegment segment 52 55, bearing vestigial cheliceral seta and lyrifissure, and distal membranous process; distal pseudosegment 17 19; movable digit (12) a hyaline lobe bearing some narrow, straight sclerotised elements; arthrodial corona and fixed digit absent.
</paragraph>
<paragraph id="8BBF3686FF8FFFFA2E0FFAF90C52F823" blockId="9.[826,1429,1293,1949]" pageId="9" pageNumber="27">
Legs (
<figureCitation id="133B2A03FF8FFFFA2EC0FAFA0D08FA9A" box="[943,1049,1293,1317]" captionStart="FIGURE" captionStartId="8.[159,171,1855,1874]" captionTargetBox="[317,1279,326,1817]" captionTargetId="figure-5@8.[288,1300,312,1824]" captionTargetPageId="8" captionText="FIGURE 6: Myrmozercon brevipes Berlese 1902, female: A Tarsus I (circle = blunt-tipped, ribbed seta; diamond = minute seta; x = abruptly tapering seta); B Leg I, coxa to tibia; C Leg II, coxa to tarsus; D Leg III, trochanter to tarsus; E Leg IV, trochanter to tarsus;" figureDoi="http://doi.org/10.5281/zenodo.5401744" httpUri="https://zenodo.org/record/5401744/files/figure.png" pageId="9" pageNumber="27">Figs. 6</figureCitation>
A-F) — Excluding ambulacra, lengths of leg I 280 295, leg II 370 410, leg III 385 425, leg IV 410 445. Tarsus I (72 77) with elongate ambulacrum (20 24), without claws; ambulacra II- IV (48 55), each with a dorsal ribbed leaf-like pretarsus (
<figureCitation id="133B2A03FF8FFFFA2EFEFA4A0AF9FA6A" box="[913,1000,1469,1493]" captionStart="FIGURE" captionStartId="8.[159,171,1855,1874]" captionTargetBox="[317,1279,326,1817]" captionTargetId="figure-5@8.[288,1300,312,1824]" captionTargetPageId="8" captionText="FIGURE 6: Myrmozercon brevipes Berlese 1902, female: A Tarsus I (circle = blunt-tipped, ribbed seta; diamond = minute seta; x = abruptly tapering seta); B Leg I, coxa to tibia; C Leg II, coxa to tarsus; D Leg III, trochanter to tarsus; E Leg IV, trochanter to tarsus;" figureDoi="http://doi.org/10.5281/zenodo.5401744" httpUri="https://zenodo.org/record/5401744/files/figure.png" pageId="9" pageNumber="27">Fig. 6F</figureCitation>
); claws I absent, II-IV hyaline. Setation of leg segments I-IV: coxae 2-2-2-2; trochanters 4-4-4-5 (TrI-III with larval complement for
<taxonomicName id="4C004D05FF8FFFFA285AF9F40A76F981" authorityName="A.Berlese" authorityYear="1892" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="27" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
); femora 8-8-8-8 or, as (
<emphasis id="B974EA94FF8FFFFA2914F9D10DA1F981" box="[1147,1200,1574,1598]" italics="true" pageId="9" pageNumber="27">al -ad</emphasis>
/
<emphasis id="B974EA94FF8FFFFA29AFF9D00DC8F981" box="[1216,1241,1575,1598]" italics="true" pageId="9" pageNumber="27">av</emphasis>
,
<emphasis id="B974EA94FF8FFFFA2987F9D10C12F981" box="[1256,1283,1574,1598]" italics="true" pageId="9" pageNumber="27">pd</emphasis>
/
<emphasis id="B974EA94FF8FFFFA287DF9D00C3CF981" box="[1298,1325,1575,1598]" italics="true" pageId="9" pageNumber="27">pv</emphasis>
,
<emphasis id="B974EA94FF8FFFFA2854F9D10C41F981" box="[1339,1360,1574,1598]" italics="true" pageId="9" pageNumber="27">pl</emphasis>
), I-IV (1-2/1, 2/1-1); genua 8-8-8-8, or I-IV (1-2/1, 2/1-1); tibiae 8-7-7-7, or I (1-2/1, 2/1-1), II-IV (1-1/1, 2/1- 1); tarsi II-IV 17-17-17, with
<emphasis id="B974EA94FF8FFFFA29EDF9780DC8F918" box="[1154,1241,1679,1703]" italics="true" pageId="9" pageNumber="27">ad1-pd1</emphasis>
seta-like, apical processes 0.4-0.5 × length of pretarsi,
<emphasis id="B974EA94FF8FFFFA2993F9450C33F975" box="[1276,1314,1714,1738]" italics="true" pageId="9" pageNumber="27">ad1</emphasis>
subequal to
<emphasis id="B974EA94FF8FFFFA2E34F9220A92F952" box="[859,899,1749,1773]" italics="true" pageId="9" pageNumber="27">pd1</emphasis>
. Tarsus I with 30 setae: four ribbed blunttipped setae (length 16 26) dorsodistally among dorsodistal cluster of nine abruptly tapering setae plus one tiny seta (3), and 16 proximal and ventral strongly tapered setae. All setae on other leg segments unspecialised, smooth, length 13 35.
</paragraph>
<caption id="DF7F660EFF8CFFF92DF0F8BA0A68F8C3" ID-DOI="http://doi.org/10.5281/zenodo.5401746" ID-Zenodo-Dep="5401746" httpUri="https://zenodo.org/record/5401746/files/figure.png" pageId="10" pageNumber="28" startId="10.[159,171,1869,1888]" targetBox="[326,1237,346,1789]" targetPageId="10">
<paragraph id="8BBF3686FF8CFFF92DF0F8BA0A68F8C3" blockId="10.[159,1414,1869,1916]" pageId="10" pageNumber="28">
FIGURE 7:
<taxonomicName id="4C004D05FF8CFFF92C6AF8BA0B52F8DF" authority="Berlese 1902" authorityName="Berlese" authorityYear="1902" box="[261,579,1869,1888]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="28" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8CFFF92C6AF8BA08D9F8DF" box="[261,456,1869,1888]" italics="true" pageId="10" pageNumber="28">Myrmozercon brevipes</emphasis>
<bibRefCitation id="EF914B77FF8CFFF92CA2F8BA0B52F8DF" author="Berlese A." box="[461,579,1869,1888]" pageId="10" pageNumber="28" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese 1902</bibRefCitation>
</taxonomicName>
, male: A Venter; extensive setation of opisthogaster partially illustrated; B Chelicerae; C Ventral gnathosoma (c = corniculus; im = internal malae; lb = labrum).
</paragraph>
</caption>
<paragraph id="8BBF3686FF8DFFF82DAAFEC00876FEF0" blockId="11.[159,762,311,1949]" box="[197,359,311,335]" pageId="11" pageNumber="29">
<emphasis id="B974EA94FF8DFFF82DAAFEC009EDFEF0" box="[197,252,311,335]" italics="true" pageId="11" pageNumber="29">Male</emphasis>
(
<figureCitation id="133B2A03FF8DFFF82C65FEC00848FEF0" box="[266,345,311,335]" captionStart="FIGURE" captionStartId="10.[159,171,1869,1888]" captionTargetBox="[326,1237,346,1789]" captionTargetId="figure-5@10.[282,1306,325,1837]" captionTargetPageId="10" captionText="FIGURE 7: Myrmozercon brevipes Berlese 1902, male: A Venter; extensive setation of opisthogaster partially illustrated; B Chelicerae; C Ventral gnathosoma (c = corniculus; im = internal malae; lb = labrum)." figureDoi="http://doi.org/10.5281/zenodo.5401746" httpUri="https://zenodo.org/record/5401746/files/figure.png" pageId="11" pageNumber="29">Figs. 7</figureCitation>
).
</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAFE9108B8FE7B" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="29">Dorsal idiosoma — Dorsal shield (length 650 660 × width 550 560), similar in ornamentation and setation to female.</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAFE2C0881FC28" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="29">
Ventral idiosoma (
<figureCitation id="133B2A03FF8DFFF82CC4FE2C0B0BFE4C" box="[427,538,475,499]" captionStart="FIGURE" captionStartId="10.[159,171,1869,1888]" captionTargetBox="[326,1237,346,1789]" captionTargetId="figure-5@10.[282,1306,325,1837]" captionTargetPageId="10" captionText="FIGURE 7: Myrmozercon brevipes Berlese 1902, male: A Venter; extensive setation of opisthogaster partially illustrated; B Chelicerae; C Ventral gnathosoma (c = corniculus; im = internal malae; lb = labrum)." figureDoi="http://doi.org/10.5281/zenodo.5401746" httpUri="https://zenodo.org/record/5401746/files/figure.png" pageId="11" pageNumber="29">Fig. 7A</figureCitation>
) — Sternoventral shield (420 long × 200 wide at level of
<emphasis id="B974EA94FF8DFFF82F0BFE090B95FDA9" box="[612,644,510,534]" italics="true" pageId="11" pageNumber="29">st4</emphasis>
), fused to exopodal shield, bearing slender sternal setae and 21 opisthogastric setae at posterior margin; of setae
<emphasis id="B974EA94FF8DFFF82DA1FD900815FDC0" box="[206,260,615,639]" italics="true" pageId="11" pageNumber="29">st1-5</emphasis>
,
<emphasis id="B974EA94FF8DFFF82C76FD90082BFDC1" box="[281,314,615,638]" italics="true" pageId="11" pageNumber="29">st5</emphasis>
longest (36), other sternal setae (30 32); longitudinal reticulation throughout except near anterolateral margin. Genital operculum in anterior concavity of sternal shield. Anal shield similar to female (length 135 × width 100). Soft cuticle hypertrichous, setal length 30 80, longer posteriorly. Poroids on soft cuticle and anal shield absent; lyrifissure
<emphasis id="B974EA94FF8DFFF82C09FCAA0899FCCB" box="[358,392,861,884]" italics="true" pageId="11" pageNumber="29">iv5</emphasis>
near posterolateral margin of sternoventral shield.
</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAFC590BC0FB60" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="29">
Gnathosoma (
<figureCitation id="133B2A03FF8DFFF82C04FC5908DFFC79" box="[363,462,942,966]" captionStart="FIGURE" captionStartId="10.[159,171,1869,1888]" captionTargetBox="[326,1237,346,1789]" captionTargetId="figure-5@10.[282,1306,325,1837]" captionTargetPageId="10" captionText="FIGURE 7: Myrmozercon brevipes Berlese 1902, male: A Venter; extensive setation of opisthogaster partially illustrated; B Chelicerae; C Ventral gnathosoma (c = corniculus; im = internal malae; lb = labrum)." figureDoi="http://doi.org/10.5281/zenodo.5401746" httpUri="https://zenodo.org/record/5401746/files/figure.png" pageId="11" pageNumber="29">Figs. 7C</figureCitation>
) — Similar to female, setae
<emphasis id="B974EA94FF8DFFF82DA5FC2609F4FC56" box="[202,229,977,1001]" italics="true" pageId="11" pageNumber="29">h1</emphasis>
28,
<emphasis id="B974EA94FF8DFFF82C76FC260825FC56" box="[281,308,977,1001]" italics="true" pageId="11" pageNumber="29">h2</emphasis>
14,
<emphasis id="B974EA94FF8DFFF82C07FC260893FC56" box="[360,386,977,1001]" italics="true" pageId="11" pageNumber="29">h3</emphasis>
11, palpcoxal seta absent, palp length 95, corniculi similar to female. Subcapitulum extends medially as two finely spiculate processes reaching level of mid-palp femur; labrum finely spiculate, larger, thicker than in female, reaching level of mid-palp femur.
<emphasis id="B974EA94FF8DFFF82CBFFB760B2FFB26" box="[464,574,1153,1177]" italics="true" pageId="11" pageNumber="29">Chelicerae</emphasis>
(
<figureCitation id="133B2A03FF8DFFF82F21FB760BBBFB26" box="[590,682,1153,1177]" captionStart="FIGURE" captionStartId="10.[159,171,1869,1888]" captionTargetBox="[326,1237,346,1789]" captionTargetId="figure-5@10.[282,1306,325,1837]" captionTargetPageId="10" captionText="FIGURE 7: Myrmozercon brevipes Berlese 1902, male: A Venter; extensive setation of opisthogaster partially illustrated; B Chelicerae; C Ventral gnathosoma (c = corniculus; im = internal malae; lb = labrum)." figureDoi="http://doi.org/10.5281/zenodo.5401746" httpUri="https://zenodo.org/record/5401746/files/figure.png" pageId="11" pageNumber="29">Fig. 7B</figureCitation>
): fixed digit absent; movable digit represented by hyaline sheath that encloses spermatodactyl (length 73).
</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAFB010BE8FAEB" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="29">Legs — Chaetotaxy as in female, legs subequal in length compared with female. Tarsus I as in female; all other leg setae unspecialised, as in female.</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAFA9C0B41F9EA" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="29">
Remarks — The only previous record for
<taxonomicName id="4C004D05FF8DFFF82FC5FA9C09C7FA1A" authorityName="Berlese" authorityYear="1902" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8DFFF82FC5FA9C09C7FA1A" italics="true" pageId="11" pageNumber="29">M. brevipes</emphasis>
</taxonomicName>
is from
<collectingCountry id="F3177616FF8DFFF82C56FA79087FFA19" box="[313,366,1422,1446]" name="Italy" pageId="11" pageNumber="29">Italy</collectingCountry>
with the ant
<taxonomicName id="4C004D05FF8DFFF82F60FA7809F1FA76" authority="(Berlese 1904)" baseAuthorityName="Berlese" baseAuthorityYear="1904" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="erraticum">
<emphasis id="B974EA94FF8DFFF82F60FA780B84FA1A" box="[527,661,1422,1446]" italics="true" pageId="11" pageNumber="29">T. erraticum</emphasis>
(
<bibRefCitation id="EF914B77FF8DFFF82FCBFA7909C9FA76" author="Berlese A." pageId="11" pageNumber="29" pagination="299 - 474" refId="ref4419" refString="Berlese A. 1904 - Illustrazione iconografica degli Acari mirmecofili - Redia, 1: 299 - 474." type="journal article" year="1904">Berlese 1904</bibRefCitation>
)
</taxonomicName>
. Our new specimens are from
<collectingCountry id="F3177616FF8DFFF82F01FA460BD0FA76" box="[622,705,1457,1481]" name="Greece" pageId="11" pageNumber="29">Greece</collectingCountry>
, but from the same host species, and we expect
<taxonomicName id="4C004D05FF8DFFF82FC7FA2209C7F9B0" authorityName="Berlese" authorityYear="1902" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8DFFF82FC7FA2209C7F9B0" italics="true" pageId="11" pageNumber="29">M. brevipes</emphasis>
</taxonomicName>
to be present throughout the intervening terrain in the Balkan Peninsula and central Europe where
<taxonomicName id="4C004D05FF8DFFF82D81F9C90862F9EA" baseAuthorityName="Berlese" baseAuthorityYear="1904" box="[238,371,1598,1621]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="erraticum">
<emphasis id="B974EA94FF8DFFF82D81F9C90862F9EA" box="[238,371,1598,1621]" italics="true" pageId="11" pageNumber="29">T. erraticum</emphasis>
</taxonomicName>
is known to occur.
</paragraph>
<paragraph id="8BBF3686FF8DFFF82DAAF99B0C3EFC1C" blockId="11.[159,762,311,1949]" lastBlockId="11.[826,1429,311,931]" pageId="11" pageNumber="29">
Species of
<taxonomicName id="4C004D05FF8DFFF82C2DF99A08C8F93B" authorityName="Berlese" authorityYear="1902" box="[322,473,1645,1668]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA94FF8DFFF82C2DF99A08C8F93B" box="[322,473,1645,1668]" italics="true" pageId="11" pageNumber="29">Myrmozercon</emphasis>
</taxonomicName>
are highly host specific, with no species known from more than one species of ant, although in three known cases two species of
<taxonomicName id="4C004D05FF8DFFF82DF0F9210827F952" authorityName="Berlese" authorityYear="1902" box="[159,310,1750,1773]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA94FF8DFFF82DF0F9210827F952" box="[159,310,1750,1773]" italics="true" pageId="11" pageNumber="29">Myrmozercon</emphasis>
</taxonomicName>
are known to utilize the same species of ant (Shawand Seeman 2009). In these cases, the species pairs conform to a Hutchinsonian size ratio (Shawand Seeman 2009). Therefore, the host species and size of the mite is informative. Our specimens match
<bibRefCitation id="EF914B77FF8DFFF82C1FF8730B35F822" author="Berlese A." box="[368,548,1924,1949]" pageId="11" pageNumber="29" pagination="299 - 474" refId="ref4419" refString="Berlese A. 1904 - Illustrazione iconografica degli Acari mirmecofili - Redia, 1: 299 - 474." type="journal article" year="1904">Berleses (1904)</bibRefCitation>
illustrations in all regards, including the bizarre chelicerae of the female and male. Their size is also similar.
<bibRefCitation id="EF914B77FF8DFFF8282EFEAD0A90FE2A" author="Berlese A." pageId="11" pageNumber="29" pagination="697 - 700" refId="ref4369" refString="Berlese A. 1902 - Specie di Acari nuovi - Zool. Anz., 25: 697 - 700." type="journal article" year="1902">Berlese (1902)</bibRefCitation>
measured the female as 800 µm × 609 µm and the male at 700 µm × 560 µm. Dr Roberto Nannelli (Berlese Collection, Florence) measured all available intact females, including the
<typeStatus id="54BB8824FF8DFFF829D2FE110C34FE41" box="[1213,1317,486,510]" pageId="11" pageNumber="29" type="holotype">holotype</typeStatus>
(n = 4, on slides 5/39 and 5/40,
<typeStatus id="54BB8824FF8DFFF82925FDFE0DA3FD9E" box="[1098,1202,521,545]" pageId="11" pageNumber="29" type="holotype">holotype</typeStatus>
measurement first followed by range of values based on three other females at the Berlese Collection), and recorded the length as 792 µm (720-768) and width as 600 µm (552 576). Berleses male was remeasured at 720 µm × 552 µm. Our females (640 760 µm × 550 640 µm) and males (650 660 µm × 550 560 µm) tended to be slightly shorter, and
<bibRefCitation id="EF914B77FF8DFFF82919FD080C0BFCA8" author="Berlese A." box="[1142,1306,767,791]" pageId="11" pageNumber="29" pagination="299 - 474" refId="ref4419" refString="Berlese A. 1904 - Illustrazione iconografica degli Acari mirmecofili - Redia, 1: 299 - 474." type="journal article" year="1904">Berlese (1904)</bibRefCitation>
also illustrated the species with a slightly tapering idiosoma, while ours are ovate. We regard our specimens as the same species, attributing the small differences in size and shape to intraspecific variation.
</paragraph>
<paragraph id="8BBF3686FF8DFFF82E0FFC260C16FA6B" blockId="11.[826,1429,977,1492]" pageId="11" pageNumber="29">
The chelicerae of
<taxonomicName id="4C004D05FF8DFFF82946FC250DB7FC56" authorityName="Berlese" authorityYear="1902" box="[1065,1190,977,1001]" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8DFFF82946FC250DB7FC56" box="[1065,1190,977,1001]" italics="true" pageId="11" pageNumber="29">M. brevipes</emphasis>
</taxonomicName>
are unlike any other member of the genus (or perhaps the Acari) and are difficult to interpret. Two hypotheses are given. In the first, the lateral cheliceral lyrifissure has enlarged to such a size that it separates the second cheliceral segment into two pseudosegments (
<figureCitation id="133B2A03FF8DFFF82807FB760A4BFB03" captionStart="FIGURE" captionStartId="7.[159,171,1823,1842]" captionTargetBox="[164,1587,346,1783]" captionTargetId="figure-4@7.[159,1662,344,1791]" captionTargetPageId="7" captionText="FIGURE 5: Myrmozercon brevipes Berlese 1902, female, chelicerae from three different specimens, with two interpretations of cheliceral morphology labelled (see Remarks). cs = vestigial cheliceral seta; dps = distal pseudosegment of second cheliceral digit; ly = lyrifissure; md = movable digit; pps = proximal pseudosegment of second cheliceral digit;? = novel structure." figureDoi="http://doi.org/10.5281/zenodo.5401742" httpUri="https://zenodo.org/record/5401742/files/figure.png" pageId="11" pageNumber="29">Fig. 5B</figureCitation>
). The dorsal lyrifissure and vestigial cheliceral seta remain on the proximal pseudosegment, while the distal pseudosegment terminates in a membranous movable digit that includes small rod-like sclerotised elements. The dorsal, membranous flange at the margin of the pseudosegmental division is a novel structure. The lyrifissures and setae are absent or not visible in males. The terminology in the description is based on this hypothesis.
</paragraph>
<paragraph id="8BBF3686FF8DFFF82E0FF9F40D03F823" blockId="11.[826,1429,1539,1948]" pageId="11" pageNumber="29">
An alternative interpretation is that the movable digit is equivalent to the distal pseudosegment of the first hypothesis, and this bears a novel structure distally (
<figureCitation id="133B2A03FF8DFFF82EB9F99B0D3FF93B" box="[982,1070,1644,1668]" captionStart="FIGURE" captionStartId="7.[159,171,1823,1842]" captionTargetBox="[164,1587,346,1783]" captionTargetId="figure-4@7.[159,1662,344,1791]" captionTargetPageId="7" captionText="FIGURE 5: Myrmozercon brevipes Berlese 1902, female, chelicerae from three different specimens, with two interpretations of cheliceral morphology labelled (see Remarks). cs = vestigial cheliceral seta; dps = distal pseudosegment of second cheliceral digit; ly = lyrifissure; md = movable digit; pps = proximal pseudosegment of second cheliceral digit;? = novel structure." figureDoi="http://doi.org/10.5281/zenodo.5401742" httpUri="https://zenodo.org/record/5401742/files/figure.png" pageId="11" pageNumber="29">Fig. 5C</figureCitation>
). In support of this is
<bibRefCitation id="EF914B77FF8DFFF82841F99B0A90F918" author="Berlese A." pageId="11" pageNumber="29" pagination="299 - 474" refId="ref4419" refString="Berlese A. 1904 - Illustrazione iconografica degli Acari mirmecofili - Redia, 1: 299 - 474." type="journal article" year="1904">Berleses (1904)</bibRefCitation>
drawing of cheliceral musculature for
<taxonomicName id="4C004D05FF8DFFF8282AF9670A60F975" authorityName="Berlese" authorityYear="1902" class="Arachnida" family="Laelapidae" genus="Myrmozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B974EA94FF8DFFF8282AF9670A60F975" italics="true" pageId="11" pageNumber="29">M. brevipes</emphasis>
</taxonomicName>
, showing the adductor muscle of the movable digit attaching to the first separation in the second cheliceral digit. In this interpretation, the small membranous flange is a much reduced fixed digit. Cheliceral musculature was poorly visible in our specimens, but what was visible supported Berleses drawing.
</paragraph>
</subSubSection>
</treatment>
</document>