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<document ID-DOI="10.1206/0003-0082(2004)433&lt;0001:FSDICA&gt;2.0.CO;2" ID-GBIF-Dataset="51eea930-5f0d-43f6-8e03-3093edf36cd7" ID-ISSN="0003-0082" ID-Zenodo-Dep="4712338" checkinTime="1619135562207" checkinUser="felipe" docAuthor="BLAGODEROV, VLADIMIR &amp; GRIMALDI, DAVID" docDate="2004" docId="039A87C9FF90FF8FFF5BFBA2FC1FFD0F" docLanguage="en" docName="AmMusNovit.2004.3433.1-76.pdf" docOrigin="American Museum Novitates 3433 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&amp;doi=10.1206%2F0003-0082(2004)433%3C0001%3AFSDICA%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.1:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="1" docTitle="Protognoriste BLAGODEROV &amp; GRIMALDI 2004, new genus" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFA3FFB1FF80FF9FFFB0FFB9FFA1FFC4" masterDocTitle="Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae" masterLastPageNumber="76" masterPageNumber="1" pageNumber="17" updateTime="1661833164902" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>BLAGODEROV, VLADIMIR</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>GRIMALDI, DAVID</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>American Museum Novitates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2004</mods:date>
<mods:detail type="pubDate">
<mods:number>2004-02-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3433</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.bioone.org/perlserv/?request=get-abstract&amp;doi=10.1206%2F0003-0082(2004)433%3C0001%3AFSDICA%3E2.0.CO%3B2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0082(2004)433&lt;0001:FSDICA&gt;2.0.CO;2</mods:identifier>
<mods:identifier type="GBIF-Dataset">51eea930-5f0d-43f6-8e03-3093edf36cd7</mods:identifier>
<mods:identifier type="ISSN">0003-0082</mods:identifier>
<mods:identifier type="Zenodo-Dep">4712338</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5060480" ID-GBIF-Taxon="183235918" ID-Zenodo-Dep="5060480" LSID="urn:lsid:plazi:treatment:039A87C9FF90FF8FFF5BFBA2FC1FFD0F" httpUri="http://treatment.plazi.org/id/039A87C9FF90FF8FFF5BFBA2FC1FFD0F" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection box="[235,541,1051,1075]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph blockId="16.[235,541,1051,1075]" box="[235,541,1051,1075]" pageId="16" pageNumber="17">
<heading box="[235,541,1051,1075]" centered="true" fontSize="36" level="2" pageId="16" pageNumber="17" reason="8">
<emphasis bold="true" box="[235,405,1051,1075]" pageId="16" pageNumber="17">
<taxonomicName authority="BLAGODEROV &amp; GRIMALDI, 2004" authorityName="BLAGODEROV &amp; GRIMALDI" authorityYear="2004" box="[235,398,1051,1075]" class="Insecta" family="Diadocidiidae" genus="Protognoriste" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="new genus">
<emphasis bold="true" box="[235,398,1051,1075]" italics="true" pageId="16" pageNumber="17">Protognoriste</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[415,541,1051,1074]" pageId="16" pageNumber="17" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="diagnosis">
<paragraph blockId="16.[124,654,1102,1741]" pageId="16" pageNumber="17">
DIAGNOSIS: Wing membrane without macrotrichia, occiput without row of strong setae. Palpi short. Eyes bare. Face wide. Stem of M and M
<subScript attach="left" box="[267,276,1203,1217]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
absent. RS base distinct. Crossvein
<emphasis box="[188,230,1220,1243]" italics="true" pageId="16" pageNumber="17">r­m</emphasis>
meets M
<subScript attach="left" box="[352,383,1232,1246]" fontSize="6" pageId="16" pageNumber="17">3+4</subScript>
and CuA stem. CuA gently curved. Abdomen insertion broad. Fore tibiae shorter than femora.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="materials_examined">
<paragraph blockId="16.[124,654,1102,1741]" pageId="16" pageNumber="17">
<typeStatus box="[153,211,1308,1331]" pageId="16" pageNumber="17">TYPE</typeStatus>
SPECIES:
<taxonomicName authority="BLAGODEROV &amp; GRIMALDI, 2004" authorityName="BLAGODEROV &amp; GRIMALDI" authorityYear="2004" box="[334,645,1308,1331]" class="Insecta" family="Keroplatidae" genus="Protognoriste" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="Amplicauda" status="n.sp.">
<emphasis box="[334,645,1308,1331]" italics="true" pageId="16" pageNumber="17">Protognoriste amplicauda</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[124,179,1337,1360]" pageId="16" pageNumber="17" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="etymology">
<paragraph blockId="16.[124,654,1102,1741]" pageId="16" pageNumber="17">
ETYMOLOGY: The name is a combination of
<emphasis box="[163,237,1396,1419]" italics="true" pageId="16" pageNumber="17">protos</emphasis>
(Greek πρω´ τos, or first) and the genus name
<taxonomicName authorityName="Meigen" authorityYear="1818" box="[272,374,1425,1448]" class="Insecta" family="Mycetophilidae" genus="Gnoriste" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis box="[272,374,1425,1448]" italics="true" pageId="16" pageNumber="17">Gnoriste</emphasis>
</taxonomicName>
. The name is feminine.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="discussion">
<paragraph blockId="16.[124,654,1102,1741]" lastBlockId="16.[700,1229,193,715]" pageId="16" pageNumber="17">
COMMENTS: The following apomorphic characters refer the genera
<taxonomicName authorityName="BLAGODEROV &amp; GRIMALDI" authorityYear="2004" box="[485,653,1484,1507]" class="Insecta" family="Keroplatidae" genus="Plesiognoriste" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis box="[485,653,1484,1507]" italics="true" pageId="16" pageNumber="17">Plesiognoriste</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="BLAGODEROV &amp; GRIMALDI" authorityYear="2004" box="[183,344,1513,1536]" class="Insecta" family="Keroplatidae" genus="Protognoriste" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis box="[183,344,1513,1536]" italics="true" pageId="16" pageNumber="17">Protognoriste</emphasis>
</taxonomicName>
to the
<taxonomicName authorityName="Edwards" authorityYear="1925" box="[452,650,1513,1536]" class="Insecta" family="Lygistorrhinidae" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Lygistorrhinidae</taxonomicName>
: short palpi; small dorsal cleft of anepisternum; anepisternum and metepimeron not divided by laterotergite (possibly plesiomorphic); short, incomplete Sc; short R
<subScript attach="right" box="[527,536,1643,1657]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
; RS base and M stem reduced;
<emphasis box="[416,458,1660,1683]" italics="true" pageId="16" pageNumber="17">r­m</emphasis>
horizontal and fused with R
<subScript attach="left" box="[274,283,1701,1715]" fontSize="6" pageId="16" pageNumber="17">5</subScript>
; hind coxa smaller and broader than mid coxa; abdomen insertion narrow; cerci simple. Face wide, with three ocelli, situated almost in straight line, suggestive of
<taxonomicName authorityName="Edwards" authorityYear="1925" box="[700,827,252,275]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subFamily" subFamily="Manotinae">Manotinae</taxonomicName>
, but structures of palpi, katepisternum and metepimeron, and absence of strong setae, show no apomorphies with respect to that subfamily. Eye size, face width, vein vestiture, tibial spurs of equal size, and complicated shape of the gonostyli are more primitive conditions than occur in Recent lygistorrhinids, although some features of venation and gonostyli are apomorphic. The short fork of M
<subScript attach="left" box="[889,920,528,542]" fontSize="6" pageId="16" pageNumber="17">3+4</subScript>
and CuA and absence of a long proboscis are most similar to the genus
<taxonomicName authority="Matile. It" authorityName="Matile. It" box="[750,976,574,598]" class="Insecta" family="Lygistorrhinidae" genus="Seguyola" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis box="[750,857,575,598]" italics="true" pageId="16" pageNumber="17">Seguyola</emphasis>
Matile. It
</taxonomicName>
is possible that these two genera should be treated as a separate subfamily of
<taxonomicName authorityName="Edwards" authorityYear="1925" box="[864,1061,633,656]" class="Insecta" family="Lygistorrhinidae" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Lygistorrhinidae</taxonomicName>
. We prefer to expand the definition of the family to include newly described taxa.
</paragraph>
</subSubSection>
</treatment>
</document>