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<document id="1EEF9B141A0A2556B328BA5B32B71D0E" ID-DOI="10.11646/zootaxa.3272.1.1" ID-GBIF-Dataset="02e53dfe-221c-4c20-aa11-82fc7e6e5318" ID-ISSN="1175-5326" ID-Zenodo-Dep="215455" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1460472935642" checkinUser="plazi" docAuthor="Rooijen, Johan Van &amp; Vogel, Gernot" docDate="2012" docId="039579531E0C8F552E86FDFDDE60F8BB" docLanguage="en" docName="zt03272p025.pdf" docOrigin="Zootaxa 3272" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dendrelaphis terrificus Peters 1872" docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FFAC012B1E1E8F462E11FFD9DB07FFD1" masterDocTitle="A revision of the taxonomy of Dendrelaphis caudolineatus (Gray, 1834) (Serpentes: Colubridae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="19" updateTime="1698306253900" updateUser="plazi">
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<mods:title id="45604461AE7036B1053230B3C9FB2808">A revision of the taxonomy of Dendrelaphis caudolineatus (Gray, 1834) (Serpentes: Colubridae)</mods:title>
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<mods:namePart id="53D6D92E9FFA7D7FA6E232686DBC3016">Rooijen, Johan Van</mods:namePart>
<mods:affiliation id="4D84D92BC9E1CA757AA82E368EBECC10">Netherlands Centre for Biodiversity Naturalis, P. O. Box 9517, 2300 RA Leiden, The Netherlands &amp; Tulpentuin 313, 2272 EH, Voorburg, The Netherlands</mods:affiliation>
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<mods:namePart id="5AE0462DC9A541419DACD9E848D71248">Vogel, Gernot</mods:namePart>
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<treatment id="039579531E0C8F552E86FDFDDE60F8BB" ID-DOI="http://doi.org/10.5281/zenodo.6318696" ID-GBIF-Taxon="119515812" ID-Zenodo-Dep="6318696" LSID="urn:lsid:plazi:treatment:039579531E0C8F552E86FDFDDE60F8BB" httpUri="http://treatment.plazi.org/id/039579531E0C8F552E86FDFDDE60F8BB" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C3269BCE1E0C8F542E86FDFDD965FDEC" box="[151,610,547,574]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B83C8451E0C8F542E86FDFDD965FDEC" blockId="18.[151,610,547,606]" box="[151,610,547,574]" pageId="18" pageNumber="19">
<heading id="D0CB7F291E0C8F542E86FDFDD965FDEC" bold="true" box="[151,610,547,574]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C3CB3C61E0C8F542E86FDFDD965FDEC" authority="Peters, 1872" authorityName="Peters" authorityYear="1872" box="[151,610,547,574]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B94814571E0C8F542E86FDFDD965FDEC" bold="true" box="[151,610,547,574]" pageId="18" pageNumber="19">
<emphasis id="B94814571E0C8F542E86FDFDDAA8FDEF" bold="true" box="[151,431,548,574]" italics="true" pageId="18" pageNumber="19">Dendrelaphis terrificus</emphasis>
(
<bibRefCitation id="EFADB5B41E0C8F542FAEFDFAD95EFDEC" author="Peters" box="[447,601,547,573]" firstAuthor="Peters" pageId="18" pageNumber="19" pagination="581" refId="ref14582" refString="Peters, W. C. H. (1872) Mittheilung uber einige von Hrn. Dr. A. B. Meyer bei Gorontalo und auf den Togian-Inseln gesammelte Amphibien. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1872, 581 - 585." type="journal article" year="1872">Peters, 1872</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3269BCE1E0C8F542E86FD9FDE8DFCAF" pageId="18" pageNumber="19" type="description">
<paragraph id="8B83C8451E0C8F542E86FD9FDA25FD8F" blockId="18.[151,610,547,606]" box="[151,290,582,606]" pageId="18" pageNumber="19">
<figureCitation id="1307D4C01E0C8F542E86FD9FDBF4FD8F" box="[151,243,582,606]" captionStart="FIGURE 14" captionStartId="18.[151,249,1927,1950]" captionTargetBox="[151,1435,948,1905]" captionTargetId="figure@18.[151,1436,948,1907]" captionTargetPageId="18" captionText="FIGURE 14. Dendrelaphis terrificus (Peters, 1872) juvenile from Sulawesi, lectotype (ZMB 7416; TL 30.5 cm.)." httpUri="https://zenodo.org/record/215469/files/figure.png" pageId="18" pageNumber="19">Figs. 14</figureCitation>
,
<figureCitation id="1307D4C01E0C8F542EEEFD9FDA19FD8F" box="[255,286,582,606]" captionStart="FIGURE 15" captionStartId="19.[151,249,1195,1218]" captionTargetBox="[151,1435,185,1173]" captionTargetId="figure@19.[151,1436,185,1174]" captionTargetPageId="19" captionText="FIGURE 15. Dendrelaphis terrificus (Peters, 1872) adult from Sulawesi (RMNH 40179 (1); TL 114.5 cm.)." httpUri="https://zenodo.org/record/215470/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
.
</paragraph>
<paragraph id="8B83C8451E0C8F542E86FD57DE8DFCAF" blockId="18.[151,1436,654,894]" pageId="18" pageNumber="19">
<emphasis id="B94814571E0C8F542E86FD57DA2EFD76" bold="true" box="[151,297,654,679]" pageId="18" pageNumber="19">Description.</emphasis>
A species of the
<taxonomicName id="4C3CB3C61E0C8F542FF8FD57D823FD77" box="[489,804,654,678]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="caudolineatus">
<emphasis id="B94814571E0C8F542FF8FD57D823FD77" box="[489,804,654,678]" italics="true" pageId="18" pageNumber="19">Dendrelaphis caudolineatus</emphasis>
</taxonomicName>
group, characterized by the combination of 1) a reduced pattern of black longitudinal stripes, stripes along the entire length of the body, 6 stripes at midbody; 2) a pale ventrolateral stripe along the entire length of the body; 3) a black postocular stripe which extends onto the neck where it becomes one the upper longitudinal stripe that borders the pale ventrolateral line; 4) 170175 ventrals (males), 174181 ventrals (females); 5) 94102 subcaudals; 6) relative tail-length 0.2450.260 (males), 0.2550.265 (females); 7) 67 temporal scales on each side; 8) 911 infralabials on each side; 9) 45 scales touching the posterior borders of the parietal scales; 10) maximum total length males
<quantity id="4CC465A01E0C8F542A44FCBFDFB5FCAF" box="[1109,1202,870,894]" metricMagnitude="-1" metricUnit="m" metricValue="9.75" pageId="18" pageNumber="19" unit="cm" value="97.5">97.5 cm</quantity>
, females 116.0 cm.
</paragraph>
</subSubSection>
<caption id="DF4398CD1E0C8F542E86F85EDE1FF84F" box="[151,1304,1927,1950]" httpUri="https://zenodo.org/record/215469/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1435,948,1905]" targetPageId="18">
<paragraph id="8B83C8451E0C8F542E86F85EDE1FF84F" blockId="18.[151,1304,1927,1950]" box="[151,1304,1927,1950]" pageId="18" pageNumber="19">
<emphasis id="B94814571E0C8F542E86F85EDA18F84C" bold="true" box="[151,287,1927,1950]" pageId="18" pageNumber="19">FIGURE 14.</emphasis>
<taxonomicName id="4C3CB3C61E0C8F542F34F851D9A2F84F" authority="(Peters, 1872)" authorityName="Peters" authorityYear="1872" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[293,677,1928,1950]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B94814571E0C8F542F34F851D916F84F" box="[293,529,1928,1950]" italics="true" pageId="18" pageNumber="19">Dendrelaphis terrificus</emphasis>
(
<bibRefCitation id="EFADB5B41E0C8F542C0FF851D999F84F" author="Peters" box="[542,670,1928,1950]" firstAuthor="Peters" pageId="18" pageNumber="19" pagination="581" refId="ref14582" refString="Peters, W. C. H. (1872) Mittheilung uber einige von Hrn. Dr. A. B. Meyer bei Gorontalo und auf den Togian-Inseln gesammelte Amphibien. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1872, 581 - 585." type="journal article" year="1872">Peters, 1872</bibRefCitation>
)
</taxonomicName>
juvenile from Sulawesi, lectotype (ZMB 7416; TL 30.5 cm.).
</paragraph>
</caption>
<caption id="DF4398CD1E0D8F552E86FB72DFE6FB10" box="[151,1249,1195,1218]" httpUri="https://zenodo.org/record/215470/files/figure.png" pageId="19" pageNumber="20" targetBox="[151,1435,185,1173]" targetPageId="19">
<paragraph id="8B83C8451E0D8F552E86FB72DFE6FB10" blockId="19.[151,1249,1195,1218]" box="[151,1249,1195,1218]" pageId="19" pageNumber="20">
<emphasis id="B94814571E0D8F552E86FB72DA18FB11" bold="true" box="[151,287,1195,1218]" pageId="19" pageNumber="20">FIGURE 15.</emphasis>
<taxonomicName id="4C3CB3C61E0D8F552F34FB72D9A2FB13" authority="(Peters, 1872)" authorityName="Peters" authorityYear="1872" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[293,677,1195,1218]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B94814571E0D8F552F34FB72D916FB10" box="[293,529,1195,1217]" italics="true" pageId="19" pageNumber="20">Dendrelaphis terrificus</emphasis>
(
<bibRefCitation id="EFADB5B41E0D8F552C0FFB75D999FB13" author="Peters" box="[542,670,1196,1218]" firstAuthor="Peters" pageId="19" pageNumber="20" pagination="581" refId="ref14582" refString="Peters, W. C. H. (1872) Mittheilung uber einige von Hrn. Dr. A. B. Meyer bei Gorontalo und auf den Togian-Inseln gesammelte Amphibien. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1872, 581 - 585." type="journal article" year="1872">Peters, 1872</bibRefCitation>
)
</taxonomicName>
adult from Sulawesi (RMNH 40179 (1); TL 114.5 cm.).
</paragraph>
</caption>
<subSubSection id="C3269BCE1E0D8F552ED6FAC8DE6BF9BF" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B83C8451E0D8F552ED6FAC8DE6BF9BF" blockId="19.[151,1436,1297,1898]" pageId="19" pageNumber="20">
<emphasis id="B94814571E0D8F552ED6FAC8D908FAFB" bold="true" box="[199,527,1297,1322]" pageId="19" pageNumber="20">
Description of the
<typeStatus id="548776E71E0D8F552FB5FACBD908FAFB" box="[420,527,1298,1322]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
</emphasis>
.
<materialsCitation id="3B54C2181E0D8F552C0FFACADE5BFAFA" ID-GBIF-Occurrence="3753898304" box="[542,1372,1298,1323]" pageId="19" pageNumber="20">
<collectionCode id="ED2D50801E0D8F552C0FFACAD95CFAFB" box="[542,603,1299,1322]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="19" pageNumber="20">ZMB</collectionCode>
7416, juvenile from Gorontalo, Sulawesi, collected by A. B. Meyer
</materialsCitation>
; SVL 23.0 cm; TAIL
<quantity id="4CC465A01E0D8F552F55FAEEDA94FA9F" box="[324,403,1335,1358]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="19" pageNumber="20" unit="cm" value="7.7">7.7 cm</quantity>
; 172 ventral scales; 94 subcaudal scales; anal shield divided; dorsal scales in
<date id="FF82EE851E0D8F552AE0FAEFDE50FA9F" box="[1265,1367,1334,1358]" pageId="19" pageNumber="20" value="1913-11-13">13-13-11</date>
rows; 9 supralabials, supralabials 5,6 touch eye; 9 infralalabials, infralabials 6 and 7 touch first sublabial; infralabials 15 touch first chinshield, infralabials 5,6 touch second chinshield; 2 postoculars; temporal formula 2:2:2/2:2:2; vertebral scales not enlarged; eye-diameter
<quantity id="4CC465A01E0D8F552C94FA7BD9DBFA6B" box="[645,732,1442,1466]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="19" pageNumber="20" unit="mm" value="3.1">3.1 mm</quantity>
/ 3.0 mm; eye-nostril distance
<quantity id="4CC465A01E0D8F552A51FA7BDF90FA6B" box="[1088,1175,1442,1466]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
/
<quantity id="4CC465A01E0D8F552ABEFA7BDE0EFA6B" box="[1199,1289,1442,1466]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="20" unit="mm" value="2.6">2.6 mm</quantity>
; snout-width
<quantity id="4CC465A01E0D8F552E86FA1FDBF5FA0F" box="[151,242,1478,1502]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="20" unit="mm" value="2.8">2.8 mm</quantity>
; posterior edge of parietal scales touched by 4 scales; head-length
<quantity id="4CC465A01E0D8F552DE8FA1FDF65FA0F" box="[1017,1122,1478,1502]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="mm" value="13.5">13.5 mm</quantity>
; ground-color olive-brown; supralabials, chin and throat yellow; venter yellowish; pronounced black temporal stripe covers nearly all of temporal region, extends onto neck, runs along whole body as black stripe; 4 black stripes anteriorly (including extensions of temporal stripes), 6 black stripes posteriorly; lower two black stripes enclose pale ventrolateral line that covers upper half of first dorsal row and lower half of second dorsal row; black stripe on underside of tail.
</paragraph>
</subSubSection>
<subSubSection id="C3269BCE1E0D8F552ED6F9A0DF39F967" pageId="19" pageNumber="20" type="description">
<paragraph id="8B83C8451E0D8F552ED6F9A0DF39F967" blockId="19.[151,1436,1297,1898]" pageId="19" pageNumber="20">
<emphasis id="B94814571E0D8F552ED6F9A0DAA8F943" bold="true" box="[199,431,1657,1682]" pageId="19" pageNumber="20">Sexual dimorphism</emphasis>
. On average, males have 172 ventral scales whereas females have 178 ventral scales (F = 9.6; df = 1; p = 0.03). In addition, females grow larger than males (see description).
</paragraph>
</subSubSection>
<subSubSection id="C3269BCE1E0D8F552ED6F918DE60F8BB" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8B83C8451E0D8F552ED6F918DE60F8BB" blockId="19.[151,1436,1297,1898]" pageId="19" pageNumber="20">
<emphasis id="B94814571E0D8F552ED6F918DA5EF90B" bold="true" box="[199,345,1729,1754]" pageId="19" pageNumber="20">Distribution</emphasis>
.
<taxonomicName id="4C3CB3C61E0D8F552F7AF91BD973F90B" box="[363,628,1730,1754]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B94814571E0D8F552F7AF91BD973F90B" box="[363,628,1730,1754]" italics="true" pageId="19" pageNumber="20">Dendrelaphis terrificus</emphasis>
</taxonomicName>
is endemic to Sulawesi. Specimens included in this study originated from Manado and Gorontalo, both locations situated on the northern arm of Sulawesi, and “North Sulawesi”. These localities are in agreement with the distribution given by
<bibRefCitation id="EFADB5B41E0D8F552D0BF8D2DF31F8F3" author="De Lang &amp; Vogel" box="[794,1078,1802,1827]" firstAuthor="De Lang" pageId="19" pageNumber="20" refId="ref13063" refString="De Lang, R. &amp; Vogel, G. (2005) The snakes of Sulawesi A field guide to the land snakes of Sulawesi with identification keys, Frankfurt am Main: Edition Chimaira." type="book" year="2005">
<collectingCountry id="F32B88D51E0D8F552D0BF8D2D83CF8F3" box="[794,827,1803,1826]" name="South Georgia and the South Sandwich Islands" pageId="19" pageNumber="20">De</collectingCountry>
Lang &amp; Vogel (2005)
</bibRefCitation>
, who provided six localities, all situated on the northern arm.
<bibRefCitation id="EFADB5B41E0D8F552FCFF8F6D9C4F897" author="Wanger" box="[478,707,1838,1863]" etAl="et al." firstAuthor="Wanger" pageId="19" pageNumber="20" pagination="17 - 29" refId="ref15686" refString="Wanger, T. C., Motzke, I., Saleh, S. &amp; Iskandar, D. T. (2011) The amphibians and reptiles of the Lore Lindu National Park area, Central Sulawesi, Indonesia. Salamandra, 47, 17 - 29." type="journal article" year="2011">
Wanger
<emphasis id="B94814571E0D8F552C2AF8F6D974F897" box="[571,627,1838,1862]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2011)
</bibRefCitation>
carried out a survey in Central Sulawesi but did not document
<taxonomicName id="4C3CB3C61E0D8F552B90F8F6DBF9F8BB" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B94814571E0D8F552B90F8F6DBF9F8BB" italics="true" pageId="19" pageNumber="20">D. terrificus</emphasis>
</taxonomicName>
. However, the species has meanwhile been recorded in other parts of Sulawesi (Koch, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>