treatments-xml/data/54/42/4E/54424E035AC3DAF2C040C4D72C1C1DF4.xml

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<mods:title id="BEADFAE275B8B8F937910D68D4CE057B">Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)</mods:title>
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<mods:namePart id="DC4E82D406EBE6EC12D06148E117FEA2">Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation id="43A06CE0E37EE23CEAA4C8B4CD35BF0B">Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:namePart id="4A636F52712DE885F0DD86DE3D6FA6F9">Deli, Tamas</mods:namePart>
<mods:affiliation id="BFECB7D46DCEB8F905CAA0AEBC51D8F9">Moricz Zsigmond u. 2, Gyomaendrod, H- 5500, Hungary</mods:affiliation>
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<mods:namePart id="C37CCEB9F6C6ADA71FABFD2C0ADD15C1">Eross, Zoltan Peter</mods:namePart>
<mods:affiliation id="891582306799CC61F904E82CBCF44023">Bem u. 36., Budapest, H- 1151, Hungary</mods:affiliation>
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<mods:namePart id="5EA77309D13EBDA705BBA709249F746E">Reischuetz, Peter L.</mods:namePart>
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<mods:namePart id="779210EE62674C30D05A92CA89D5ADDD">Reischuetz, Alexander</mods:namePart>
<mods:affiliation id="123C2CF61C834CEE2FBFF0AA6C69DB17">Puechhaimg. 52, A- 3580, Horn, Austria</mods:affiliation>
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<mods:namePart id="F8F94FC0B2F8DDA050E174700DE430D9">Feher, Zoltan</mods:namePart>
<mods:affiliation id="044032D15A9263CC31C70566233ED34D">Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H- 1088, Hungary</mods:affiliation>
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<treatment id="54424E035AC3DAF2C040C4D72C1C1DF4" ID-GBIF-Taxon="182229754" LSID="urn:lsid:plazi:treatment:54424E035AC3DAF2C040C4D72C1C1DF4" httpUri="http://treatment.plazi.org/id/54424E035AC3DAF2C040C4D72C1C1DF4" lastPageNumber="13" pageId="0" pageNumber="13" scope_class="Gastropoda" scope_family="Spelaeodiscidae">
<subSubSection id="8D917D0952E080A2547490D746EB7BCE" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="AC5423C03D6DA1E61A14ECD9DDE7E546" pageId="0" pageNumber="13">
Genus 
<taxonomicName id="E408A19306147E2E4D64D983689A1FD8" ID-CoL="7PH5P" authority="Brusina, 1886" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">Spelaeodiscus Brusina, 1886</taxonomicName>
</paragraph>
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<paragraph id="AA6BDFF57F4ADC4F512E508530ED56C2" pageId="0" pageNumber="13">
<taxonomicName id="F18C16ECB64FBA1E100AF2E8D062F5BE" authority="" class="Gastropoda" family="Spelaeodiscidae" genus="Patula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patula (Spelaeodiscus)" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subGenus" subGenus="Spelaeodiscus">Patula (Spelaeodiscus)</taxonomicName>
Brusina, 1886: 37.
</paragraph>
</subSubSection>
<subSubSection id="17E888633238C5F35FCA01AB98C6EA91" pageId="0" pageNumber="13" type="type species">
<paragraph id="EAAEC4D3E8A6D7FD62F5E982EAD7D8AE" pageId="0" pageNumber="13">Type species.</paragraph>
<paragraph id="A276F5DEE8F609E713FBFA1C26C3EEF3" pageId="0" pageNumber="13">
<emphasis id="9D1ED96FBAF9EEC2EBFCF5D1ADB52F86" italics="true" pageId="0" pageNumber="13">
<taxonomicName id="2C284F77D7094FEC626921CAEA2BC068" class="Gastropoda" family="Spelaeodiscidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">Helix</taxonomicName>
Hauffeni
</emphasis>
Schmidt, 1855.
</paragraph>
</subSubSection>
<subSubSection id="37D227D70EBB08242272F24D66808D3F" pageId="0" pageNumber="13" type="distribution">
<paragraph id="34C8954E0B3890E2A5238CC766CAC05D" pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph id="0CADF7F4D978E422ADE9A57846B9E121" pageId="0" pageNumber="13">
The genus 
<taxonomicName id="2D7A5931A5C7C7A8C45952ED41FD72CD" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="711B24B243B9F91F7E5548E7201A8F6A" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
has a disjunct distribution. One species (
<taxonomicName id="4BF89A8533CFD1CCAE1281C210202DDE" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis id="0D993F2A8CD5A2370890D49DEE4813E5" italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
) is only known from Slovenia, whereas the rest of the genus is distributed in the vicinity of the Skadar Lake Basin (also known as 
<normalizedToken id="61F0BE577E3A25F54CD0A21837173351" originalValue="Shkodër">Shkoder</normalizedToken>
Lake or Skutari Lake) in Montenegro and northern Albania (Figure 
<figureCitation id="05626BBD740AD46246E1BBD6C50D86A8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the genus Spelaeodiscus Brusina, 1886." figureDoi="10.3897/zookeys.769.25258.figure2" httpUri="https://binary.pensoft.net/fig/211836" pageId="0" pageNumber="13">2</figureCitation>
).
</paragraph>
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<paragraph id="13066A9548AD09353D257A548DF83F79" pageId="0" pageNumber="13">
<emphasis id="8A1507658DD05D8F8124985ED66692AB" bold="true" pageId="0" pageNumber="13">Figure 2.</emphasis>
Distribution of the genus 
<taxonomicName id="4F0C0E7B25A6C0CD4D772CFC23F06A26" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="61800972B29FD949B9699AE492BA3BA2" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
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Brusina, 1886.
</paragraph>
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</subSubSection>
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<paragraph id="7C2551F726BA02D9EB96D0177AA3BBC7" pageId="0" pageNumber="13">Included taxa.</paragraph>
<paragraph id="E4172DF65097EE4AAF2BEC0AA5E73CF1" pageId="0" pageNumber="13">
<taxonomicName id="658F520D3FB3D06776C5C1930798DC76" authority="albanicus" authorityName="albanicus (A. J. Wagner" authorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus subsp. albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="albanicus" subSpecies="albanicus">
<emphasis id="26E3C97DA0A4FEEE6D02C6423A422BBB" italics="true" pageId="0" pageNumber="13">Spelaeodiscus albanicus albanicus</emphasis>
</taxonomicName>
(A. J. Wagner, 1914), 
<taxonomicName id="BE825F74083FB83692DF3E5A5065F4FB" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. albanicus subsp. edentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="albanicus" subSpecies="edentatus">
<emphasis id="B645EE13324BE30B5B9E7E23ADF8C5F5" italics="true" pageId="0" pageNumber="13">S. albanicus edentatus</emphasis>
</taxonomicName>
<normalizedToken id="FA15D5FC6CCCD44DD66930233B70E810" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; P. L. 
<normalizedToken id="BEE27786CD246FE06FC11FBCE782A9BC" originalValue="Reischütz">Reischuetz</normalizedToken>
, ssp. n., 
<taxonomicName id="384278389F2331A303F7D1B0B00A53F8" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. dejongi" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="dejongi">
<emphasis id="CB6E2271FF1EE3F46892E981A63C6DF2" italics="true" pageId="0" pageNumber="13">S. dejongi</emphasis>
</taxonomicName>
Gittenberger, 1969, 
<taxonomicName id="7FFF6F9BB81A610618BB91647B72FB0C" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. densecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="densecostatus">
<emphasis id="D3AF18DB1173E4F173C010CA658B79DD" italics="true" pageId="0" pageNumber="13">S. densecostatus</emphasis>
</taxonomicName>
<normalizedToken id="BECF34B33A5536906BD6496DD8E2C65F" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; A. 
<normalizedToken id="F7A856394254DF4FC05D64AA1F3B240F" originalValue="Reischütz">Reischuetz</normalizedToken>
, sp. n., 
<taxonomicName id="8091D4C0C94164A1A28A6F4F171BAB13" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis id="B975ACC663995B4DA77E8D7950C7278B" italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
(Schmidt, 1855) 
<taxonomicName id="1AA695101D0765F7D367F94E3256E435" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. hunyadii" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hunyadii">
<emphasis id="A2D29BEB4D962C8A4FAE1CDB03D2E466" italics="true" pageId="0" pageNumber="13">S. hunyadii</emphasis>
</taxonomicName>
<normalizedToken id="1C8C807E1FE78493DF6828AC93DB561F" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Deli, sp. n., 
<taxonomicName id="5BD5005C06328F8C190D87D4BAB191D8" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="latecostatus">
<emphasis id="166F1B72141E156D06D0B6AEFB1C11EC" italics="true" pageId="0" pageNumber="13">S. latecostatus</emphasis>
</taxonomicName>
<normalizedToken id="60EBE170ABDCD6E8283D3B4448AEDB1D" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken id="BEFFC2078E5A9E401AF7A6D8A17619C4" originalValue="Erőss">Eross</normalizedToken>
, sp. n., 
<taxonomicName id="209028E8C01AD4F8AC0754913E5DFA54" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. obodensis" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="obodensis">
<emphasis id="593AA8BB4A9230CEFCD1608FD3C5E02A" italics="true" pageId="0" pageNumber="13">S. obodensis</emphasis>
</taxonomicName>
Bole, 1965. 
<taxonomicName id="9717107997F5D0040FB92E21278D13AE" authority="unidentatus" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. unidentatus subsp. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="unidentatus" subSpecies="unidentatus">
<emphasis id="8D1B072D5E02F37B0EDFFE5E44C2ED33" italics="true" pageId="0" pageNumber="13">S. unidentatus unidentatus</emphasis>
</taxonomicName>
Bole, 1961, 
<taxonomicName id="9AB178A84A4BF55D8E1971CAD43A9D6D" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. unidentatus subsp. acutus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="unidentatus" subSpecies="acutus">
<emphasis id="2368225ADF1132C6037948B050D9BD57" italics="true" pageId="0" pageNumber="13">S. unidentatus acutus</emphasis>
</taxonomicName>
<normalizedToken id="CC645D2B4F2D09CD9ABDCBE2646C806E" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken id="13D88D1E0274AF9672A725ABD1AF5FB7" originalValue="Fehér">Feher</normalizedToken>
, ssp. n., 
<taxonomicName id="D2F3B8460F7CE9FC48A6FF9C7A6553D7" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="virpazarioides">
<emphasis id="BA92FE99351B45FB8A6D82EAA8F0A468" italics="true" pageId="0" pageNumber="13">S. virpazarioides</emphasis>
</taxonomicName>
<normalizedToken id="173737A1F5186603019EC5229B19B645" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken id="9144606AEAD7531B69CD180268632903" originalValue="Fehér">Feher</normalizedToken>
, sp. n. For key traits see Table 
<tableCitation id="31F93742E8453294AF102116934900B0" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Number of ribs on the body whorl, shell size, and key traits of Spelaeodiscus species." httpUri="http://table.plazi.org/id/AE7CCB10728A6F0F311DC97F022F808D" pageId="0" pageNumber="13" tableUuid="AE7CCB10728A6F0F311DC97F022F808D">1</tableCitation>
.
</paragraph>
<caption id="AE7CCB10728A6F0F311DC97F022F808D" ID-Table-UUID="AE7CCB10728A6F0F311DC97F022F808D" httpUri="http://table.plazi.org/id/AE7CCB10728A6F0F311DC97F022F808D" pageId="0" pageNumber="13" start="Table 1" startId="T1">
<paragraph id="C48427B8BF15775CA0A5F3F772BB5A11" pageId="0" pageNumber="13">
<emphasis id="A81A249395FAD2AC5DE565BB55544BDC" bold="true" pageId="0" pageNumber="13">Table 1.</emphasis>
Number of ribs on the body whorl, shell size, and key traits of 
<taxonomicName id="804818FF0A95AC904866E54DBE2C552F" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="72D83B4ACAFC89718A6D8A1F0CC03E11" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="088ED6A1304CE084675F5CD2925429A8" pageId="0" pageNumber="13">
<table id="46CD79936F913F60EB1156C3C5BEFCDF" pageId="0" pageNumber="13">
<tr id="3D6CA2BFC7F41FB11149B2843FD8A04A" pageId="0" pageNumber="13">
<th id="4911B7CC230294701C1002AC89BD2479" colspan="1" pageId="0" pageNumber="13" rowspan="1">(Sub)species</th>
<th id="8452C986D8EED40776012797813E01EE" colspan="1" pageId="0" pageNumber="13" rowspan="1">No. of ribs</th>
<th id="89B7B21456D30F9121432EB835712325" colspan="1" pageId="0" pageNumber="13" rowspan="1">Shell diameter (in mm)</th>
<th id="BBD7772A47B97E54AA6F64C792B821AB" colspan="1" pageId="0" pageNumber="13" rowspan="1">Key traits</th>
</tr>
<tr id="FAF78761AF762668216562DC5C926870" pageId="0" pageNumber="13">
<td id="44F36A57A922EB45CE090EF3696ACB8E" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<emphasis id="CAC613BFD49594986D3D3ADC2221D373" italics="true" pageId="0" pageNumber="13">
<taxonomicName id="7B000910ACC3B0FA3AAD3FAE1915FEBE" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="albanicus">Spelaeodiscus albanicus</taxonomicName>
<taxonomicName id="F2B8350BEACB1A1A6ED683710A66CD01" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="albanicus">Spelaeodiscus albanicus</taxonomicName>
</emphasis>
</td>
<td id="D12E192F97E2F121486449E5B73665FC" colspan="1" pageId="0" pageNumber="13" rowspan="1">43-93</td>
<td id="87FEB1AC59A1F615C36045E85EFA197C" colspan="1" pageId="0" pageNumber="13" rowspan="1">3.6-4.3</td>
<td id="06349C4FFFEB1861286F6CD1F1C0BEF7" colspan="1" pageId="0" pageNumber="13" rowspan="1">matte protoconch, weak palatal and two weak basal teeth</td>
</tr>
<tr id="9767AFF181FB9F0B1A4E6383BD0A6264" pageId="0" pageNumber="13">
<td id="14EE9C4476B626B0F51A0A2B0CD55599" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="3298C9356195E3ADD2E856BA3B4EA792" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus edentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="edentatus">
<emphasis id="BA276E9AC3B2D226B223E1F3917EE5BE" italics="true" pageId="0" pageNumber="13">Spelaeodiscus albanicus edentatus</emphasis>
</taxonomicName>
ssp. n.
</td>
<td id="F4CF114354A32AC40508F471381DE8BE" colspan="1" pageId="0" pageNumber="13" rowspan="1">35-54</td>
<td id="D8368D95D3477E1D635E88C1F1D91396" colspan="1" pageId="0" pageNumber="13" rowspan="1">3.6-4.2</td>
<td id="099D62A970DE487F7D8AADC97586A993" colspan="1" pageId="0" pageNumber="13" rowspan="1">widely-spaced ribs, glossy protoconch glossy, toothless aperture</td>
</tr>
<tr id="788627E44D9A1FD393D254D9F5AC5B4A" pageId="0" pageNumber="13">
<td id="97A8F7DA9CD26D4556EF341D4CE98A54" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="4957BFAC03EA727098FCAD0935D73CFC" authorityName="Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus dejongi" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="dejongi">
<emphasis id="C6EAE0C6378F3F3411516D03454891D6" italics="true" pageId="0" pageNumber="13">Spelaeodiscus dejongi</emphasis>
</taxonomicName>
</td>
<td id="B972A83330F52DF827C95E105CB396B4" colspan="1" pageId="0" pageNumber="13" rowspan="1">57-112</td>
<td id="AF1AB0E138375B698FE47F2D9A964E97" colspan="1" pageId="0" pageNumber="13" rowspan="1">1.9-3.4</td>
<td id="3E3847BF0BB6F09D6677E82324A98DCD" colspan="1" pageId="0" pageNumber="13" rowspan="1">dense, low ribs, smooth protoconch, toothless aperture</td>
</tr>
<tr id="9AB65B3885824A2A942222BB26F0FB15" pageId="0" pageNumber="13">
<td id="B28592B157500D44BD9723DFC46C629C" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="93DB9F41470062BAF4E6581926D3451A" authorityName="Pall-Gergely &amp; A. Reischuetz" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus densecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="densecostatus">
<emphasis id="83F8B25A6B15319B72D54180A64373B0" italics="true" pageId="0" pageNumber="13">Spelaeodiscus densecostatus</emphasis>
</taxonomicName>
sp. n.
</td>
<td id="68D2CA90B0837C99AF0EF54C814218DC" colspan="1" pageId="0" pageNumber="13" rowspan="1">116</td>
<td id="B1C833ED80939AA4E8CBDE74FB2EA6E2" colspan="1" pageId="0" pageNumber="13" rowspan="1">3.7</td>
<td id="19E048B63CA686F7A81931BB364A2230" colspan="1" pageId="0" pageNumber="13" rowspan="1">very low and dense ribs, toothless aperture</td>
</tr>
<tr id="E3BA1ED3326D8A0B677E03EB1D270B85" pageId="0" pageNumber="13">
<td id="16904BAF9BB748AE5FC9CBBF087DC8EB" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="68147D4F01BA97F39976658FF18BE4B6" baseAuthorityName="Schmidt" baseAuthorityYear="1855" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis id="33BECAC51943FC424B42834E426B7CA1" italics="true" pageId="0" pageNumber="13">Spelaeodiscus hauffeni</emphasis>
</taxonomicName>
</td>
<td id="6E23726256617FE947F22652B13630D7" colspan="1" pageId="0" pageNumber="13" rowspan="1">41-52</td>
<td id="F7F0D4938864DF15EB9DFD4D11287220" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.8-3.5</td>
<td id="F0C86DDDC6C501857F9CBA76BAAD4431" colspan="1" pageId="0" pageNumber="13" rowspan="1">widely spaced, strong ribs, rounded, toothless aperture, finely granular protoconch</td>
</tr>
<tr id="30914E933E49A404D087874BBCB0590D" pageId="0" pageNumber="13">
<td id="047CFD7E8A7D58FF789DD7EBC3110539" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="BAC7B29E08B79A65CE714F5F05F8AADF" authorityName="Pall-Gergely &amp; Eross" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="latecostatus">
<emphasis id="E0CFEDDE6B20C4F25317A550126D6732" italics="true" pageId="0" pageNumber="13">Spelaeodiscus latecostatus</emphasis>
</taxonomicName>
sp. n.
</td>
<td id="429F9B250447C7F43CE6CBEE6B4326BA" colspan="1" pageId="0" pageNumber="13" rowspan="1">42</td>
<td id="F424A43E5627B99EA1C1E545F4C8E405" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.2</td>
<td id="BFE53A52F6BB7ED2F7BD8347B7A61BA8" colspan="1" pageId="0" pageNumber="13" rowspan="1">strong, very widely spaced ribs, glossy protoconch, toothless aperture</td>
</tr>
<tr id="4C5197FD97C0BADE9FEF3E25009D91DB" pageId="0" pageNumber="13">
<td id="7A903CBBFF0C06325C58AFDEE952E70A" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="23EB24387326CBE4E5C5EDCD31704276" authorityName="Pall-Gergely &amp; Eross" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="latecostatus">Spelaeodiscus cf. latecostatus</taxonomicName>
sp. n. (2017/005)
</td>
<td id="7AB713669E8D65722733D7CF8C12496B" colspan="1" pageId="0" pageNumber="13" rowspan="1">47-54</td>
<td id="E73523E364B4FE65645ACFAE46FFF2B8" colspan="1" pageId="0" pageNumber="13" rowspan="1">1.9</td>
<td id="497E6624225D7FC8F037273294C62853" colspan="1" pageId="0" pageNumber="13" rowspan="1">strong, widely spaced ribs, glossy protoconch, toothless aperture</td>
</tr>
<tr id="972807B5DF9B394872D2FDFD8C3B18B9" pageId="0" pageNumber="13">
<td id="6D9BB713933D1723AB4C7AFA8E07793A" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="725C73E362B87DA1FD0FE90A7A58EDFF" authorityName="Pall-Gergely &amp; Deli" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus hunyadii" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hunyadii">
<emphasis id="A4A59B79A77A6D09B8FA0D17481A4996" italics="true" pageId="0" pageNumber="13">Spelaeodiscus hunyadii</emphasis>
</taxonomicName>
sp. n.
</td>
<td id="B425CF2263CF0D65DBCEEA971576F9E8" colspan="1" pageId="0" pageNumber="13" rowspan="1">42-48</td>
<td id="7086A062F82524EB86BC628BA76A1884" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.1-2.2</td>
<td id="3AF323D754F209F42B2E33DFE957B038" colspan="1" pageId="0" pageNumber="13" rowspan="1">widely spaced, strong ribs, glossy protoconch, strongly oblique, toothless aperture</td>
</tr>
<tr id="1B26207536B5872352B928C0AE8CD5B4" pageId="0" pageNumber="13">
<td id="1487F552918FDE1D4658F193742951A7" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="DE96E618228950421A6A6BAB3EAC339F" authorityName="Páll-Gergely &amp; Deli &amp; Erőss &amp; Reischütz &amp; Reischütz &amp; Fehér" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus obodensis" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="obodensis">
<emphasis id="59BEC65AFB2B633981C23DDBCC2FDE83" italics="true" pageId="0" pageNumber="13">Spelaeodiscus obodensis</emphasis>
</taxonomicName>
</td>
<td id="EA722CF61BA489F6914F332D7AC533A6" colspan="1" pageId="0" pageNumber="13" rowspan="1">43-76</td>
<td id="131DC69EDD7B5D5F1B659E609B3A403A" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.6-3.0</td>
<td id="8DF0D5C30C152C3252D983D19D3F5F93" colspan="1" pageId="0" pageNumber="13" rowspan="1">elevated spire, roughly sculptured protoconch, strong ribs, toothless aperture</td>
</tr>
<tr id="A345B550DCAB7BBD6FD0D6A418CE8058" pageId="0" pageNumber="13">
<td id="452E77599A78F2040BD226E9BB15CBD9" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<emphasis id="FA13D86FB73A2500E48355024E97F24D" italics="true" pageId="0" pageNumber="13">
<taxonomicName id="E2E6510A1DA7F89AA172743D5CAA2CB9" authorityName="Bole" authorityYear="1961" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">Spelaeodiscus unidentatus</taxonomicName>
<taxonomicName id="BBD9EF437E25FD5A4502F045FD265972" authorityName="Bole" authorityYear="1961" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">Spelaeodiscus unidentatus</taxonomicName>
</emphasis>
</td>
<td id="4533082E25C237898E13384C83D4ECCF" colspan="1" pageId="0" pageNumber="13" rowspan="1">74-118</td>
<td id="055710481F8AC5FB8BD01BFD003EF30E" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.4-3.2</td>
<td id="7B4651501A53BC1AFD0BF2201BEA02B2" colspan="1" pageId="0" pageNumber="13" rowspan="1">low basal tooth; palatal part of peristome with strong incision</td>
</tr>
<tr id="3D28535C24B64DF4524D692A74B62643" pageId="0" pageNumber="13">
<td id="B72E82AACF37527D9A07214FFA05C1C6" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="B4B1E1D43C34A4DA75595A30CCEB8B7A" authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus subsp. acutus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="unidentatus" subSpecies="acutus">
<emphasis id="46AC0FF252A883A2BF5D3F827F46E9CF" italics="true" pageId="0" pageNumber="13">Spelaeodiscus unidentatus acutus</emphasis>
</taxonomicName>
ssp. n.
</td>
<td id="1EF9403AFD639A8A1322F6549C71FD5D" colspan="1" pageId="0" pageNumber="13" rowspan="1">64-91</td>
<td id="6F79C9C0747229E8C43834ECC5CB59E2" colspan="1" pageId="0" pageNumber="13" rowspan="1">2.9-3.5</td>
<td id="D2E44895765E3F1DFAB645981C90CF1D" colspan="1" pageId="0" pageNumber="13" rowspan="1">pointed basal tooth; palatal part of peristome with shallow incision</td>
</tr>
<tr id="00A2F7135DBF851AA1E342733AEFDBBE" pageId="0" pageNumber="13">
<td id="0216FF3D724860E8190B443DF5E003D9" colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName id="CC0D1C7E03F14D49BF83FCED974CCD64" authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="virpazarioides">
<emphasis id="CBAD43B8DEAB1B2960448E451738F071" italics="true" pageId="0" pageNumber="13">Spelaeodiscus virpazarioides</emphasis>
</taxonomicName>
sp. n.
</td>
<td id="BB26086E6B8C3487A46A290034F4FEDB" colspan="1" pageId="0" pageNumber="13" rowspan="1">40-70</td>
<td id="8FCC9CEBE9E244699904F73EC5A16A99" colspan="1" pageId="0" pageNumber="13" rowspan="1">3.3-3.6</td>
<td id="7898F880996FFD102E4CCD22C6E1F569" colspan="1" pageId="0" pageNumber="13" rowspan="1">spiral sculpture, thickened callus</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="2413D85E017009753C924394B57B24CF" pageId="0" pageNumber="13" type="delimitation of this genus">
<paragraph id="1630473AF19562DFC274860260107629" pageId="0" pageNumber="13">Delimitation of this genus.</paragraph>
<paragraph id="D5A6144048F392B91424508EE1A228ED" pageId="0" pageNumber="13">
The reproductive anatomy of 
<taxonomicName id="9A1C56688DE616415168BBCF4B05BDB6" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="F66E659EEE2E14989A28BC431A884049" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C57356CE4740F87B3AE48E824B86F7F8" authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="C3425DB82FC9DF55B72CB98EB320E8DC" italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
is characterized by a short penial caecum, a well-developed penial appendix, sometimes an epiphallic caecum, and a bursa copulatrix without a diverticulum. The retractor muscle is divided into two bounds, one inserting on the penial appendix, whereas the other at the base of the penial caecum. Examining the anatomical descriptions and drawings of 
<taxonomicName id="75DAD6E96EAE7FD0705AB7CB9D8BDE69" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EAE8B01D21709F14EDA8A0476630E668" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
(
<bibRefCitation id="D02C137D0E054FA4D63219862E9AF986" DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
) and 
<taxonomicName id="2CA9DE577455C2D6B56F42D746D8AB5C" authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="908EFC6D7013B5DBACA318E4B2C0D931" italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
(
<bibRefCitation id="8F8B086E5685077593CD82AD768435C8" author="Hudec, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="13" pagination="157 - 165" refId="B19" refString="Hudec, V, 1965. Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna. Archiv fuer Molluskenkunde 94: 157 - 165" title="Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna." volume="94" year="1965">Hudec 1965</bibRefCitation>
, 
<bibRefCitation id="A715B2DA07871B2548C91B50D20E9782" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="13" pagination="263 - 277" refId="B14" refString="Gittenberger, E, 1975. Beitraege zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277" title="Beitraege zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag." volume="48" year="1975">Gittenberger 1975</bibRefCitation>
, 
<bibRefCitation id="94655EF3B39B8FBCD4433E70575D1E73" author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="13" pagination="1 - 127" refId="B38" refString="Schileyko, AA, 1998. Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. Ruthenica, Supplement 2 (1): 1 - 127" title="Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae." volume="2" year="1998">Schileyko 1998</bibRefCitation>
, 
<bibRefCitation id="8EAC835E78C7B8EEF0CDB7D198AD0BB0" author="Subai, P" journalOrPublisher="Basteria" pageId="0" pageNumber="13" pagination="111 - 118" refId="B44" refString="Subai, P, Dedov, I, 2008. A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov. Basteria 72: 111 - 118" title="A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov." volume="72" year="2008">Subai and Dedov 2008</bibRefCitation>
), we were unable to find characters that would constantly differ between the two groups. For example, the penial caecum was long and slender in 
<taxonomicName id="8A57C903C8286219210DD47C9B647709" lsidName="A. tatrica" pageId="0" pageNumber="13" rank="species" species="tatrica">
<emphasis id="EB540D5DCA6402B3333000ED770454CE" italics="true" pageId="0" pageNumber="13">A. tatrica</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="1D1458FD483AF1E9BB9559CA057FA74E" author="Hudec, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="13" pagination="157 - 165" refId="B19" refString="Hudec, V, 1965. Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna. Archiv fuer Molluskenkunde 94: 157 - 165" title="Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna." volume="94" year="1965">Hudec 1965</bibRefCitation>
) and 
<taxonomicName id="F869D607E0D3708D49649B2989AE1159" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis id="B813C174F33AA766C8607C15A560331B" italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="2A20FEEF1D669E95910252769E9D35B2" DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
), but was short and conical in 
<taxonomicName id="4A61D7C152F8964A2C644A5A73FE4593" lsidName="A. triaria" pageId="0" pageNumber="13" rank="species" species="triaria">
<emphasis id="E669FE80DB2290F6F6F4C228250FCE9C" italics="true" pageId="0" pageNumber="13">A. triaria</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="E6A9763FB24DE67224ECE4C74996AE60" author="Subai, P" journalOrPublisher="Basteria" pageId="0" pageNumber="13" pagination="111 - 118" refId="B44" refString="Subai, P, Dedov, I, 2008. A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov. Basteria 72: 111 - 118" title="A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov." volume="72" year="2008">Subai and Dedov 2008</bibRefCitation>
) and 
<taxonomicName id="D4EF62D714F79DE9AAE75120A02E2E03" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">
<emphasis id="DC578FD6C7E6B0299A76B9DCFCFC4128" italics="true" pageId="0" pageNumber="13">S. unidentatus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="0B8C4D6AE7E9EC28BAA7B627959C58BF" DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
). Also, the shape of the bursa and the position of the starting point of the penial appendix was greatly variable across genera. Clear epiphallic caecum was only found in 
<taxonomicName id="A403156DA8EA6E3168B573D8E0CACC57" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis id="55202D446818203E987DBDD396BF90EA" italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
, but some thickening was visible in 
<taxonomicName id="28620200420164ECA3924509DEB5C3A0" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">
<emphasis id="89A5A03F6B4B0FB841FD27F6D1071E3D" italics="true" pageId="0" pageNumber="13">S. unidentatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="B27B75B4825AB78582E86DACD0ABA702" lsidName="A. triaria" pageId="0" pageNumber="13" rank="species" species="triaria">
<emphasis id="0EEADB185B9EAEC61703EECCD400C321" italics="true" pageId="0" pageNumber="13">A. triaria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0C7B00893E9B35010E802F0F3F682107" pageId="0" pageNumber="13">
As for shell characters, 
<taxonomicName id="2FE0B54A487D4A8A89056F449CFCFE96" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="C2CEC1A5CFB6531AFA928E45B2670CCA" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
is characterized by a mostly colourless shell that is smaller than 4.3 mm (majority of species are even smaller than 3.5 mm), the spire is relatively low (height of body whorl at least two third of the height of the entire shell), the body whorl is evenly rounded, the edge of the parietal callus is straight, and the peristome is only slightly expanded. In contrast, 
<taxonomicName id="E9504A2900CBE51D68F9ED694011C4BC" authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="29F3EDBDCD894646B8C1A092EB271C5F" italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
shells are brownish, larger than 4.3 mm, they have higher spire (height of body whorl is approximately half of the height of the entire shell), the shell is shape reverse trapezoid from standard apertural view, the callus is heart-shaped, and the basal part of the peristome is strongly expanded.
</paragraph>
<paragraph id="6D0FE4AB69C5EF4D4FCFB2CA2343BB2D" pageId="0" pageNumber="13">
The habitat was the only 
<normalizedToken id="4AD770AF35EFA62FE0726DA46DBED032" originalValue="“trait”">&quot;trait&quot;</normalizedToken>
mentioned by 
<bibRefCitation id="E4F43E5B29ADFED2C28B90F15DAB5056" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="13" pagination="287 - 306" refId="B13" refString="Gittenberger, E, 1969. Beitraege zur Kenntnis der Pupillacea I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306" title="Beitraege zur Kenntnis der Pupillacea I. Die Spelaeodiscinae." volume="43" year="1969">Gittenberger (1969)</bibRefCitation>
as difference between the two groups. Namely, 
<taxonomicName id="E4675219138C79771A973533FCB95977" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="279E421E97C590ACAA665DA616D54911" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
is subterranean, whereas 
<taxonomicName id="542DD33265CD58EE3D52606FCE91346B" authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="C8131D90D98D145B69267B391B4FE791" italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
can be found on rock surfaces and among leaf litter at the base of limestone rocks. In the lack of sound molecular data it is difficult to infer their relationship, but based only on ecological, conchological, and biogeographical differences it seems reasonable to keep 
<taxonomicName id="02CB64188F318C2B5F71BC944EBB59E2" authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="BCB93B7C49669A61124FD1372E1502A5" italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
and 
<taxonomicName id="8E97E464F3E9F63A5137FE05C6D68F2A" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="9005B3DF76E84ED433F4CBA820FEAB2B" italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
as distinct genera.
</paragraph>
</subSubSection>
</treatment>
</document>