treatments-xml/data/0A/6A/87/0A6A87F2FFEE800AFF10FBB9FA77FEF9.xml
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<document id="9078D9E328DA5C2A07E3C730A84CE057" ID-CLB-Dataset="29933" ID-DOI="10.11646/zootaxa.4429.3.1" ID-GBIF-Dataset="249faa66-1f6a-44a0-b7f2-f12acaf62e5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1285294" ID-ZooBank="004C0DCB-9B63-4A44-A1BB-2D98ADFB1CEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528403104686" checkinUser="plazi" docAuthor="Ferro, Gustavo Borges &amp; Marshall, Stephen A." docDate="2018" docId="0A6A87F2FFEE800AFF10FBB9FA77FEF9" docLanguage="en" docName="zootaxa.4429.3.1.pdf" docOrigin="Zootaxa 4429 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiacephala schildi Ferro &amp; Marshall 2018, new combination" docType="treatment" docVersion="9" lastPageNumber="447" masterDocId="F653FF8AFFC38024FF87FF98FFFDFFB9" masterDocTitle="A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)" masterLastPageNumber="458" masterPageNumber="401" pageNumber="446" updateTime="1698548227473" updateUser="ExternalLinkService">
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<mods:title id="801AA8739928D6E7488D3A64C84A2C3E">A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)</mods:title>
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<mods:namePart id="DE328A04F7B51C093E8502D292EF3746">Ferro, Gustavo Borges</mods:namePart>
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<mods:namePart id="A350A490488CB67228A3AC587EFD446E">Marshall, Stephen A.</mods:namePart>
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<mods:title id="D29B5271779799871D3C974BFF28CEDD">Zootaxa</mods:title>
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<mods:date id="D7E708C63643840D681AA77512F6E9F0">2018</mods:date>
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<mods:number id="1C3D0F71F77B60B1418DB86B3D34BF41">2018-06-07</mods:number>
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<treatment id="0A6A87F2FFEE800AFF10FBB9FA77FEF9" ID-DOI="http://doi.org/10.5281/zenodo.5997545" ID-GBIF-Taxon="145927993" ID-Zenodo-Dep="5997545" LSID="urn:lsid:plazi:treatment:0A6A87F2FFEE800AFF10FBB9FA77FEF9" httpUri="http://treatment.plazi.org/id/0A6A87F2FFEE800AFF10FBB9FA77FEF9" lastPageId="46" lastPageNumber="447" pageId="45" pageNumber="446">
<subSubSection id="CAD9656FFFEE8009FF10FBB9FED3FBE4" pageId="45" pageNumber="446" type="nomenclature">
<paragraph id="827C36E4FFEE8009FF10FBB9FCB8FB82" blockId="45.[151,837,1057,1117]" box="[151,837,1057,1083]" pageId="45" pageNumber="446">
<heading id="D9348188FFEE8009FF10FBB9FCB8FB82" bold="true" box="[151,837,1057,1083]" fontSize="11" level="1" pageId="45" pageNumber="446" reason="1">
<emphasis id="B0B7EAF6FFEE8009FF10FBB9FCB8FB82" bold="true" box="[151,837,1057,1083]" pageId="45" pageNumber="446">
<taxonomicName id="45C34D67FFEE8009FF10FBB9FD95FB82" authority="(Cresson, 1930)" authorityName="Ferro &amp; Marshall" authorityYear="2018" baseAuthorityName="Cresson" baseAuthorityYear="1930" box="[151,616,1057,1083]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="446" phylum="Arthropoda" rank="species" species="schildi" status="comb. nov.">
<emphasis id="B0B7EAF6FFEE8009FF10FBB9FE62FB82" bold="true" box="[151,415,1057,1083]" italics="true" pageId="45" pageNumber="446">Cardiacephala schildi</emphasis>
(
<bibRefCitation id="E6524B15FFEE8009FE28FBB9FD9DFB82" author="Cresson, E. T. Jr." box="[431,608,1057,1083]" journalOrPublisher="Transactions of the American Entomological Society" pageId="45" pageNumber="446" pagination="307 - 362" part="56" refId="ref43804" refString="Cresson, E. T. Jr. (1930) Notes on and descriptions of some Neotropical Neriidae and Micropezidae (Diptera). Transactions of the American Entomological Society, 56, 307 - 362." title="Notes on and descriptions of some Neotropical Neriidae and Micropezidae (Diptera)" type="journal article" year="1930">Cresson, 1930</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="AB84578DFFEE8009FDE8FBB9FCB8FB82" box="[623,837,1057,1083]" pageId="45" pageNumber="446">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="827C36E4FFEE8009FF10FBDCFED3FBE4" blockId="45.[151,837,1057,1117]" box="[151,302,1092,1117]" pageId="45" pageNumber="446">
(
<figureCitation id="1AF82A61FFEE8009FF18FBDCFED8FBE4" box="[159,293,1092,1117]" captionStart="FIGURES 8081" captionStartId="46.[151,264,1151,1174]" captionTargetBox="[158,1436,375,1130]" captionTargetId="figure@46.[151,1436,375,1130]" captionTargetPageId="46" captionText="FIGURES 8081. Cardiacephala schildi. 80, female, lateral view; 81, female genitalia, ventral view." httpUri="https://zenodo.org/record/1285344/files/figure.png" pageId="45" pageNumber="446">Figs. 8081</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFEE8009FF10FB14FDD6FB1B" box="[151,555,1164,1186]" pageId="45" pageNumber="446" type="reference_group">
<paragraph id="827C36E4FFEE8009FF10FB14FDD6FB1B" blockId="45.[151,555,1164,1186]" box="[151,555,1164,1186]" pageId="45" pageNumber="446">
<treatmentCitationGroup id="A2D311CAFFEE8009FF10FB14FDD6FB1B" box="[151,555,1164,1186]" pageId="45" pageNumber="446">
<taxonomicName id="45C34D67FFEE8009FF10FB14FDDAFB1B" authority="Cresson, 1930: 334" authorityName="Cresson" authorityPageNumber="334" authorityYear="1930" box="[151,551,1164,1186]" class="Insecta" family="Micropezidae" genus="Plocoscelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="446" phylum="Arthropoda" rank="species" species="schildi">
<emphasis id="B0B7EAF6FFEE8009FF10FB14FEA7FB1B" box="[151,346,1164,1186]" italics="true" pageId="45" pageNumber="446">Plocoscelus schildi</emphasis>
<treatmentCitation id="036210F5FFEE8009FEE7FB14FDDAFB1B" author="Cresson, E. T. Jr." box="[352,551,1164,1186]" page="334" pageId="45" pageNumber="446" year="1930">
<bibRefCitation id="E6524B15FFEE8009FEE7FB14FE09FB1B" author="Cresson, E. T. Jr." box="[352,500,1164,1186]" journalOrPublisher="Transactions of the American Entomological Society" pageId="45" pageNumber="446" pagination="307 - 362" part="56" refId="ref43804" refString="Cresson, E. T. Jr. (1930) Notes on and descriptions of some Neotropical Neriidae and Micropezidae (Diptera). Transactions of the American Entomological Society, 56, 307 - 362." title="Notes on and descriptions of some Neotropical Neriidae and Micropezidae (Diptera)" type="journal article" year="1930">Cresson, 1930</bibRefCitation>
: 334
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFEE8009FF10FB56FB6AF894" pageId="45" pageNumber="446" type="description">
<paragraph id="827C36E4FFEE8009FF10FB56FCCEFACE" blockId="45.[151,1437,1230,2017]" pageId="45" pageNumber="446">
<emphasis id="B0B7EAF6FFEE8009FF10FB56FED6FB5E" bold="true" box="[151,299,1230,1255]" pageId="45" pageNumber="446">Description:</emphasis>
Body length
<quantity id="453B9B01FFEE8009FE44FB48FDFBFB5E" box="[451,518,1232,1255]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="45" pageNumber="446" unit="mm" value="7.0">7 mm</quantity>
, wing
<quantity id="453B9B01FFEE8009FDD6FB57FD68FB5E" box="[593,661,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="45" pageNumber="446" unit="mm" value="4.0">4 mm</quantity>
. Palpus mostly dark brown, outer face slightly white. Clypeus brown with pale microsetulae. First flagellomere black covered with golden setulae. Fronto-orbital plates brown with a black spot posteriorly. Postpronotal lobe and scutellum brown. Row of katepisternal setae brownish. Mid and hind femora brown, slightly darkened on medial portion. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape dark brown, apex black.
</paragraph>
<paragraph id="827C36E4FFEE8009FF40FA1BFD3BF9B1" blockId="45.[151,1437,1230,2017]" pageId="45" pageNumber="446">Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Frontal vitta with a narrowed depression centrally, reaching ocellar triangle. Inner vertical seta insertion flat, without tubercle. Paracephalon smooth, without swelling (not extending beyond the vertex). Vertex rounded. Line of black supracervical setae strong.</paragraph>
<paragraph id="827C36E4FFEE8009FF40F98BFA60F923" blockId="45.[151,1437,1230,2017]" pageId="45" pageNumber="446">Thorax: Mesonotum with white pruinose and irregularly darkened. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing with stigmatal and discal bands, apex clear with a dark and narrow preapical pigmentation. Crossvein r-m with small dark spot. R4+5 and M1+2 meeting at the wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling.</paragraph>
<paragraph id="827C36E4FFEE8009FF40F930FB6AF894" blockId="45.[151,1437,1230,2017]" pageId="45" pageNumber="446">
Abdomen: Tergites with brownish setulae. Female abdomen: Oviscape with pale microsetulae. Single spermathecal duct apically broad, spermatheca cylindrical and elongated. Paired spermathecal duct uniformly thick with an apical swelling before splitting into stems. Paired spermathecal duct stems thin and smooth in basal third, apical two thirds broad and convoluted. Paired spermathecae striate and spherical (
<figureCitation id="1AF82A61FFEE8009FBB1F88CFB77F894" box="[1078,1162,1812,1837]" captionStart="FIGURES 8081" captionStartId="46.[151,264,1151,1174]" captionTargetBox="[158,1436,375,1130]" captionTargetId="figure@46.[151,1436,375,1130]" captionTargetPageId="46" captionText="FIGURES 8081. Cardiacephala schildi. 80, female, lateral view; 81, female genitalia, ventral view." httpUri="https://zenodo.org/record/1285344/files/figure.png" pageId="45" pageNumber="446">Fig. 81</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFEE800AFF40F8A0FAC3FF6D" lastPageId="46" lastPageNumber="447" pageId="45" pageNumber="446" type="materials_examined">
<paragraph id="827C36E4FFEE8009FF40F8A0FCFEF8CC" blockId="45.[151,1437,1230,2017]" pageId="45" pageNumber="446">
<emphasis id="B0B7EAF6FFEE8009FF40F8A0FE8AF8E8" bold="true" box="[199,375,1848,1873]" pageId="45" pageNumber="446">
<typeStatus id="5D788846FFEE8009FF40F8A0FEFFF8E8" box="[199,258,1848,1873]" pageId="45" pageNumber="446">Type</typeStatus>
material:
</emphasis>
<typeStatus id="5D788846FFEE8009FEF9F8A0FE1BF8E8" box="[382,486,1848,1873]" pageId="45" pageNumber="446" type="holotype">Holotype</typeStatus>
(
<specimenCount id="94C5FD6DFFEE8009FE7FF8A1FDDEF8E9" box="[504,547,1849,1873]" pageId="45" pageNumber="446" type="male">1 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFEE8009FDA9F8A1FD78F8E8" box="[558,645,1849,1873]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="446">USNM</collectionCode>
):
<collectingCountry id="FAD47674FFEE8009FD10F8A1FCC2F8E8" box="[663,831,1849,1873]" name="Costa Rica" pageId="45" pageNumber="446">COSTA RICA</collectingCountry>
. Higuito, San Mateo. Coll. Pablo Schild (white label).
<typeStatus id="5D788846FFEE8009FF10F8C5FF32F8CC" box="[151,207,1885,1909]" pageId="45" pageNumber="446">Type</typeStatus>
No. 27264 (orange label). USNMENT 0 0 677015.
</paragraph>
<paragraph id="827C36E4FFEE800AFF40F818FAC3FF6D" blockId="45.[151,1437,1230,2017]" lastBlockId="46.[151,1436,151,320]" lastPageId="46" lastPageNumber="447" pageId="45" pageNumber="446">
<emphasis id="B0B7EAF6FFEE8009FF40F818FE4EF820" bold="true" box="[199,435,1920,1945]" pageId="45" pageNumber="446">Material examined:</emphasis>
<collectingCountry id="FAD47674FFEE8009FE3DF819FD9FF820" box="[442,610,1921,1945]" name="Costa Rica" pageId="45" pageNumber="446">COSTA RICA</collectingCountry>
.
<collectingRegion id="4007F806FFEE8009FDEAF819FD16F821" box="[621,747,1921,1944]" country="Costa Rica" name="Puntarenas" pageId="45" pageNumber="446">Puntarenas</collectingRegion>
, Osa Peninsula
<quantity id="453B9B01FFEE8009FC1AF818FC16F820" box="[925,1003,1920,1945]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="45" pageNumber="446" unit="km" value="2.5">2.5 km</quantity>
S Rincon, -
<quantity id="453B9B01FFEE8009FBF6F818FB53F821" box="[1137,1198,1920,1945]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="45" pageNumber="446" unit="m" value="50.0">50 m</quantity>
, 8º 42 1 N 83º3050 W, trail nr. station, human dung,
<date id="F67D1024FFEE8009FD96F83DFD33F804" box="[529,718,1957,1981]" pageId="45" pageNumber="446" value="2001-08-10" valueMax="2001-08-11" valueMin="2001-08-10">1011.VIII.2001</date>
, S. A. Marshall (
<specimenCount id="94C5FD6DFFEE8009FC13F83DFC3DF805" box="[916,960,1957,1981]" pageId="45" pageNumber="446" type="female">2 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFEE8009FC57F83DFBE1F804" box="[976,1052,1957,1981]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="45" pageNumber="446">DEBU</collectionCode>
00 168243,
<collectionCode id="E4D2AE21FFEE8009FB1BF83CFB17F804" box="[1180,1258,1956,1981]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="45" pageNumber="446">DEBU</collectionCode>
00168270);
<collectingRegion id="4007F806FFEE8009FAF5F83DFF37F859" country="Costa Rica" name="San Jose" pageId="45" pageNumber="446">San Jose</collectingRegion>
, San Carlos, Riosparaiso Res. Pecari Stn.,
<quantity id="453B9B01FFEE8009FD21F850FD10F858" box="[678,749,1992,2017]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="45" pageNumber="446" unit="km" value="16.0">16 km</quantity>
NNE Quepos,
<quantity id="453B9B01FFEE8009FC10F851FC1DF859" box="[919,992,1993,2017]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="45" pageNumber="446" unit="m" value="400.0">400 m</quantity>
,
<geoCoordinate id="E7F75023FFEE8009FC6BF850FB96F859" box="[1004,1131,1992,2017]" direction="north" orientation="latitude" pageId="45" pageNumber="446" precision="15" value="9.564722">9º3353” N</geoCoordinate>
<geoCoordinate id="E7F75023FFEE8009FBF5F851FB05F859" box="[1138,1272,1993,2017]" direction="west" orientation="longitude" pageId="45" pageNumber="446" precision="15" value="-84.12556">84º732” W</geoCoordinate>
,
<date id="F67D1024FFEE8009FA84F850FA87F858" box="[1283,1402,1992,2017]" pageId="45" pageNumber="446" value="2003-02-15">15.II.2003</date>
, S. A. Marshall (
<specimenCount id="94C5FD6DFFED800AFEA8FF00FEA5FF16" box="[303,344,152,176]" pageId="46" pageNumber="447" type="female">1 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFED800AFEE1FF0FFE49FF09" box="[358,436,151,176]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="46" pageNumber="447">DEBU</collectionCode>
00205738);
<collectingRegion id="4007F806FFED800AFDBCFF00FD62FF16" box="[571,671,152,176]" country="Costa Rica" name="San Jose" pageId="46" pageNumber="447">San Jose</collectingRegion>
, San Carlos, Riosparaiso Res. Pecari Stn.,
<quantity id="453B9B01FFED800AFBF8FF0FFB3BFF09" box="[1151,1222,151,176]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="46" pageNumber="447" unit="km" value="16.0">16 km</quantity>
NNE Quepos,
<quantity id="453B9B01FFED800AFAF5FF00FF52FF6D" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="46" pageNumber="447" unit="m" value="400.0">400 m</quantity>
,
<geoCoordinate id="E7F75023FFED800AFF3CFF24FEC7FF6D" box="[187,314,188,213]" direction="north" orientation="latitude" pageId="46" pageNumber="447" precision="15" value="9.564722">9º3353” N</geoCoordinate>
<geoCoordinate id="E7F75023FFED800AFEC6FF24FE35FF6D" box="[321,456,188,212]" direction="west" orientation="longitude" pageId="46" pageNumber="447" precision="15" value="-84.12556">84º732” W</geoCoordinate>
,
<date id="F67D1024FFED800AFE54FF24FDB4FF6C" box="[467,585,188,213]" pageId="46" pageNumber="447" value="2003-02-15">15.II.2003</date>
, S. A. Marshall (
<specimenCount id="94C5FD6DFFED800AFC8CFF24FCCEFF6D" box="[779,819,188,212]" pageId="46" pageNumber="447" type="female">1 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFED800AFCC6FF24FC73FF6C" box="[833,910,188,213]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="46" pageNumber="447">DEBU</collectionCode>
00205735, sequence
<collectionCode id="E4D2AE21FFED800AFBFEFF24FB22FF6D" box="[1145,1247,188,212]" pageId="46" pageNumber="447">DEBUA</collectionCode>
28817).
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFED800AFF40FF47FE16FF41" box="[199,491,223,248]" pageId="46" pageNumber="447" type="distribution">
<paragraph id="827C36E4FFED800AFF40FF47FE16FF41" blockId="46.[151,1436,151,320]" box="[199,491,223,248]" pageId="46" pageNumber="447">
<emphasis id="B0B7EAF6FFED800AFF40FF47FE9FFF41" bold="true" box="[199,354,223,248]" pageId="46" pageNumber="447">Distribution:</emphasis>
<collectingCountry id="FAD47674FFED800AFEEEFF78FE1AFF41" box="[361,487,223,248]" name="Costa Rica" pageId="46" pageNumber="447">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFED800AFF40FE9CFA77FEF9" pageId="46" pageNumber="447" type="discussion">
<paragraph id="827C36E4FFED800AFF40FE9CFA77FEF9" blockId="46.[151,1436,151,320]" pageId="46" pageNumber="447">
<emphasis id="B0B7EAF6FFED800AFF40FE9CFEAFFEA4" bold="true" box="[199,338,260,285]" pageId="46" pageNumber="447">Comments:</emphasis>
This species is probably closely related to
<taxonomicName id="45C34D67FFED800AFCB8FE9DFC34FEA4" authorityName="Ferro &amp; Marshall" authorityYear="2018" baseAuthorityName="Cresson" baseAuthorityYear="1930" box="[831,969,260,285]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="447" phylum="Arthropoda" rank="species" species="harenosa">
<emphasis id="B0B7EAF6FFED800AFCB8FE9DFC34FEA4" box="[831,969,260,285]" italics="true" pageId="46" pageNumber="447">C. harenosa</emphasis>
</taxonomicName>
, although the female genitalia are unlike any other
<taxonomicName id="45C34D67FFED800AFE80FEBFFE53FEF9" box="[263,430,295,320]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="447" phylum="Arthropoda" rank="genus">
<emphasis id="B0B7EAF6FFED800AFE80FEBFFE53FEF9" box="[263,430,295,320]" italics="true" pageId="46" pageNumber="447">Cardiacephala</emphasis>
</taxonomicName>
. Male genitalia dissections were not done because the
<typeStatus id="5D788846FFED800AFB93FEBFFB8BFEF9" box="[1044,1142,295,320]" pageId="46" pageNumber="447" type="holotype">holotype</typeStatus>
is the only known male.
</paragraph>
</subSubSection>
</treatment>
</document>