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<document id="5AE72B6D50DE8D9DBAC700EE179CE914" ID-DOI="10.11646/zootaxa.3884.2.1" ID-GBIF-Dataset="919ca7ac-633d-451b-9e94-3e9a6cb563ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="251041" ID-ZooBank="3FC44983-060E-4AFE-AB51-F0455D217B69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460965288198" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Reip, Hans S., Dražina, Tvrtko, Rađa, Tonći &amp; Makarov, Slobodan E." docDate="2014" docId="476B87CB3239F831B49CBAB313ADFF00" docLanguage="en" docName="zt03884p121.pdf" docOrigin="Zootaxa 3884 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Balkanodesmus biokovensis Antic and Reip, sp. n." docType="treatment" docVersion="10" lastPageNumber="106" masterDocId="BB52FFB33238F834B40BBC2D1704FFF8" masterDocTitle="Three new monotypic genera of Trichopolydesmidae from Croatia, Balkan Peninsula (Diplopoda, Polydesmida)" masterLastPageNumber="121" masterPageNumber="101" pageNumber="102" updateTime="1698557697344" updateUser="plazi">
<mods:mods id="AED5F7E2D974D8EDD13A8F6A8E45542A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="724B123E08694822B4616EAA82D02D58">
<mods:title id="5BA58E333C8BC6312BD9A51991A778BF">Three new monotypic genera of Trichopolydesmidae from Croatia, Balkan Peninsula (Diplopoda, Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="26BE5FA642655025D0F532D7B1486BB1" type="personal">
<mods:role id="0CB821BC9AE35E58C8DF1F1C89ABA895">
<mods:roleTerm id="1A5895C240F63589ADBF3772D6814A66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61CEC592CE32193D4889449779E19E48">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="F815C75623A75522A4306F083E4AEA15" type="personal">
<mods:role id="C9573AC46968576492490B7387FFE6E4">
<mods:roleTerm id="9301BA484C7AA228F0630D8F5F29A6F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD6FA4A2E02C2E96DCB2EE2B4D517661">Reip, Hans S.</mods:namePart>
</mods:name>
<mods:name id="6256D5F96D0CF8348FB9E760820CE6C6" type="personal">
<mods:role id="885D2169C4EFCB76C017083C93F49D6C">
<mods:roleTerm id="7F8B9E8568D41EBB4CB3C759FC4DACC0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="55772859EA8CB12868D56A321CE5396D">Dražina, Tvrtko</mods:namePart>
</mods:name>
<mods:name id="8B5CBDE6796CE711ACF9CD203912EBF3" type="personal">
<mods:role id="9387CE23767F5FED3E5DD6614BE737D4">
<mods:roleTerm id="1EC0DBFA41F30DEEB69D248A0EF6D1AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED9B50CA30B72BD093E75740D63CC5AB">Rađa, Tonći</mods:namePart>
</mods:name>
<mods:name id="ACDAC670BE88C976C188A72A1F11807E" type="personal">
<mods:role id="5F4D8BD99DED4C5237E46066D134C468">
<mods:roleTerm id="FAD6A9963CCC2CCBCA2538FF3A4DF526">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="297F3608B1A06E13F71C6B2ABD885D07">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="92C55B0BCA86CA19FA37F1480A0F0E17">text</mods:typeOfResource>
<mods:relatedItem id="AC8B094024984F8E687F3FED6C94CA68" type="host">
<mods:titleInfo id="D65DF757C7533C305CED607B7937EEDE">
<mods:title id="6CE33603BD0331CA96B25D01DC5FB3ED">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="96BF5368866B809BF31E84679A743846">
<mods:date id="C72DC33D152526195C4A2C376776B15E">2014</mods:date>
<mods:detail id="562402EE2D794B3E9CC23B6B08EACE3E" type="volume">
<mods:number id="18CABB6FAD9B81D687BBF815E59ED117">3884</mods:number>
</mods:detail>
<mods:detail id="9882F49E3D90BC7A8C9F47E0002D2483" type="issue">
<mods:number id="61D9484A5835012C1BFDD1A69A58396D">2</mods:number>
</mods:detail>
<mods:extent id="E827E55C732322CE3B2234A1A88D4B0C" unit="page">
<mods:start id="A5A61DA4485F61D742E6D7FA0E3B3E65">101</mods:start>
<mods:end id="73462B46EA23B1040D77ABB654C03343">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8E97D4D756247EF5C6329241302084A4">journal article</mods:classification>
<mods:identifier id="6778999DB32451DF25C91021ADCFE6F5" type="DOI">10.11646/zootaxa.3884.2.1</mods:identifier>
<mods:identifier id="1A80198040BD36D524494D8A5B240AAB" type="GBIF-Dataset">919ca7ac-633d-451b-9e94-3e9a6cb563ec</mods:identifier>
<mods:identifier id="70317EA0469D024911C36BD80A69AA10" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="547CC8A8FDBEA3C795AEAF321E7B3151" type="Zenodo-Dep">251041</mods:identifier>
<mods:identifier id="FD2AFC4083395B0C4C3A02D96E7E4020" type="ZooBank">3FC44983-060E-4AFE-AB51-F0455D217B69</mods:identifier>
</mods:mods>
<treatment id="476B87CB3239F831B49CBAB313ADFF00" ID-DOI="http://doi.org/10.5281/zenodo.6143426" ID-GBIF-Taxon="119593256" ID-Zenodo-Dep="6143426" LSID="urn:lsid:plazi:treatment:476B87CB3239F831B49CBAB313ADFF00" httpUri="http://treatment.plazi.org/id/476B87CB3239F831B49CBAB313ADFF00" lastPageId="5" lastPageNumber="106" pageId="1" pageNumber="102">
<subSubSection id="87D865563239F835B49CBAB31609F922" pageId="1" pageNumber="102" type="nomenclature">
<paragraph id="CF7D36DD3239F835B49CBAB315FBF940" blockId="1.[151,767,1694,1754]" box="[151,767,1694,1720]" pageId="1" pageNumber="102">
<heading id="943581B13239F835B49CBAB315FBF940" bold="true" box="[151,767,1694,1720]" fontSize="11" level="1" pageId="1" pageNumber="102" reason="1">
<emphasis id="FDB6EACF3239F835B49CBAB315FBF940" bold="true" box="[151,767,1694,1720]" pageId="1" pageNumber="102">
<taxonomicName id="08C24D5E3239F835B49CBAB315B4F940" ID-CoL="KH94" authority="Antic and Reip" authorityName="Antic and Reip" box="[151,688,1694,1720]" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="biokovensis" status="sp. nov.">
<emphasis id="FDB6EACF3239F835B49CBAB316EEF940" bold="true" box="[151,490,1694,1720]" italics="true" pageId="1" pageNumber="102">Balkanodesmus biokovensis</emphasis>
Antić and Reip
</taxonomicName>
,
<taxonomicNameLabel id="E68557B43239F835B6B6BAB315FBF940" box="[701,767,1694,1720]" pageId="1" pageNumber="102" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CF7D36DD3239F835B49CBAEC1609F922" blockId="1.[151,767,1694,1754]" box="[151,269,1729,1754]" pageId="1" pageNumber="102">
<figureCitation id="57F92A583239F835B49CBAEC17DBF922" box="[151,223,1729,1754]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1886,1908]" captionTargetBox="[151,1436,195,1827]" captionTargetId="figure@2.[151,1436,195,1828]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Balkanodesmus biokovensis gen. n., sp. n., holoype male (1 – 5). 1: habitus, lateral view, 2: body segments VIII – X, dorsal view, 3: body segments VI – VIII, lateral view, 4: gonopods, lateral view, 5: gonopods, in situ. Paratype juvenile female. 6: body end, lateral view. Scale line: 1 mm (1 – 3), 0.2 mm (4 – 5) and 0.5 mm (6)." httpUri="https://zenodo.org/record/251042/files/figure.png" pageId="1" pageNumber="102">Figs. 1</figureCitation>
–12
</paragraph>
</subSubSection>
<subSubSection id="87D865563239F835B49CBB241238F892" pageId="1" pageNumber="102" type="materials_examined">
<paragraph id="CF7D36DD3239F835B49CBB241238F892" blockId="1.[151,1437,1801,1934]" pageId="1" pageNumber="102">
<emphasis id="FDB6EACF3239F835B49CBB241688F8DA" bold="true" box="[151,396,1801,1826]" pageId="1" pageNumber="102">Material examined:</emphasis>
<typeStatus id="1079887F3239F835B597BB241500F8DA" box="[412,516,1801,1826]" pageId="1" pageNumber="102" type="holotype">Holotype</typeStatus>
male: Amfora Pit, Saint Jure, Mt. Biokovo,
<collectingCountry id="B7D5764D3239F835B034BB241391F8DA" box="[1087,1173,1801,1826]" name="Croatia" pageId="1" pageNumber="102">Croatia</collectingCountry>
,
<geoCoordinate id="AAF6501A3239F835B0A3BB271213F8DA" box="[1192,1303,1802,1826]" direction="north" orientation="latitude" pageId="1" pageNumber="102" precision="55" value="43.341">43.341°N</geoCoordinate>
,
<geoCoordinate id="AAF6501A3239F835B127BB24129CF8DA" box="[1324,1432,1801,1826]" direction="east" orientation="longitude" pageId="1" pageNumber="102" precision="55" value="17.05">17.050°E</geoCoordinate>
,
<date id="BB7C101D3239F835B49CBB031627F8BE" box="[151,291,1838,1862]" pageId="1" pageNumber="102" value="2008-07-14">14.VII.2008</date>
, leg. J. Bedek (IZB, VHD BB 100-2). Allotype female: same data as
<typeStatus id="1079887F3239F835B036BB0013A6F8BE" box="[1085,1186,1837,1862]" pageId="1" pageNumber="102" type="holotype">holotype</typeStatus>
, but
<date id="BB7C101D3239F835B0D0BB031262F8BE" box="[1243,1382,1838,1862]" pageId="1" pageNumber="102" value="2008-07-17">17.VII.2008</date>
, leg. M. Lukić (IZB, VHD BB 100-3).
<typeStatus id="1079887F3239F835B618BB7F1571F892" box="[531,629,1874,1898]" pageId="1" pageNumber="102" type="paratype">Paratype</typeStatus>
juvenile female: same data as
<typeStatus id="1079887F3239F835B7C6BB7C132BF892" box="[973,1071,1873,1898]" pageId="1" pageNumber="102" type="holotype">holotype</typeStatus>
(IZB, VHD BB 100-4).
</paragraph>
</subSubSection>
<subSubSection id="87D865563239F835B4CCBB5815F6F876" box="[199,754,1909,1934]" pageId="1" pageNumber="102" type="etymology">
<paragraph id="CF7D36DD3239F835B4CCBB5815F6F876" blockId="1.[151,1437,1801,1934]" box="[199,754,1909,1934]" pageId="1" pageNumber="102">
<emphasis id="FDB6EACF3239F835B4CCBB581655F876" bold="true" box="[199,337,1909,1934]" pageId="1" pageNumber="102">Etymology:</emphasis>
After Mt. Biokovo, the
<typeStatus id="1079887F3239F835B66ABB5B1595F876" box="[609,657,1910,1934]" pageId="1" pageNumber="102">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="9BBD6655323AF836B49CBB731312F849" httpUri="https://zenodo.org/record/251042/files/figure.png" pageId="2" pageNumber="103" targetBox="[151,1436,195,1827]" targetPageId="2">
<paragraph id="CF7D36DD323AF836B49CBB731312F849" blockId="2.[151,1436,1886,1969]" pageId="2" pageNumber="103">
<emphasis id="FDB6EACF323AF836B49CBB731639F88C" bold="true" box="[151,317,1886,1908]" pageId="2" pageNumber="103">FIGURES 1–6.</emphasis>
<taxonomicName id="08C24D5E323AF836B54DBB731560F88C" box="[326,612,1886,1908]" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="biokovensis" status="gen. nov.">
<emphasis id="FDB6EACF323AF836B54DBB731560F88C" box="[326,612,1886,1908]" italics="true" pageId="2" pageNumber="103">Balkanodesmus biokovensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E68557B4323AF836B666BB7215B2F88C" box="[621,694,1887,1908]" pageId="2" pageNumber="103" rank="genus">gen. n.</taxonomicNameLabel>
, sp. n., holoype male (1–5). 1: habitus, lateral view, 2: body segments VIII–X, dorsal view, 3: body segments VI–VIII, lateral view, 4: gonopods, lateral view, 5: gonopods,
<emphasis id="FDB6EACF323AF836B094BB5013DAF86A" box="[1183,1246,1917,1938]" italics="true" pageId="2" pageNumber="103">in situ</emphasis>
. Paratype juvenile female. 6: body end, lateral view. Scale line: 1 mm (1–3), 0.2 mm (4–5) and 0.5 mm (6).
</paragraph>
</caption>
<caption id="9BBD6655323BF837B49CBBB91390F81F" httpUri="https://zenodo.org/record/251043/files/figure.png" pageId="3" pageNumber="104" targetBox="[177,1390,186,1938]" targetPageId="3">
<paragraph id="CF7D36DD323BF837B49CBBB91390F81F" blockId="3.[151,1436,1940,2023]" pageId="3" pageNumber="104">
<emphasis id="FDB6EACF323BF837B49CBBB9164CF852" bold="true" box="[151,328,1940,1962]" pageId="3" pageNumber="104">FIGURES 7–10.</emphasis>
<taxonomicName id="08C24D5E323BF837B545BBB9156EF852" box="[334,618,1940,1962]" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="biokovensis" status="gen. nov.">
<emphasis id="FDB6EACF323BF837B545BBB9156EF852" box="[334,618,1940,1962]" italics="true" pageId="3" pageNumber="104">Balkanodesmus biokovensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E68557B4323BF837B67ABBB815B3F852" box="[625,695,1941,1962]" pageId="3" pageNumber="104" rank="genus">gen. n.</taxonomicNameLabel>
, sp. n., holotype male gonopod. 7: ventral view, 8: dorsal view, 9: mesal view, 10: lateral view. Designations:
<emphasis id="FDB6EACF323BF837B617BB9F152DF830" bold="true" box="[540,553,1970,1992]" pageId="3" pageNumber="104">a</emphasis>
: acropodite,
<emphasis id="FDB6EACF323BF837B6BFBB9F15C6F830" bold="true" box="[692,706,1970,1992]" pageId="3" pageNumber="104">d</emphasis>
: denticles,
<emphasis id="FDB6EACF323BF837B736BB9F1454F830" bold="true" box="[829,848,1970,1992]" pageId="3" pageNumber="104">al</emphasis>
: apical lamella,
<emphasis id="FDB6EACF323BF837B00ABB9F1312F830" bold="true" box="[1025,1046,1970,1992]" pageId="3" pageNumber="104">bl</emphasis>
: basal lamella,
<emphasis id="FDB6EACF323BF837B0B5BB9F13C8F830" bold="true" box="[1214,1228,1970,1992]" pageId="3" pageNumber="104">ll</emphasis>
: longitudinal mesal lamella,
<emphasis id="FDB6EACF323BF837B4E5BBFC1606F81F" bold="true" box="[238,258,2001,2023]" pageId="3" pageNumber="104">lo</emphasis>
: lobuses,
<emphasis id="FDB6EACF323BF837B563BBFC1680F81F" bold="true" box="[360,388,2001,2023]" pageId="3" pageNumber="104">ml</emphasis>
: middle lamella,
<emphasis id="FDB6EACF323BF837B63EBBFC154FF81F" bold="true" box="[565,587,2001,2023]" pageId="3" pageNumber="104">pf</emphasis>
: prefemoral part,
<emphasis id="FDB6EACF323BF837B709BBFC141CF81F" bold="true" box="[770,792,2001,2023]" pageId="3" pageNumber="104">tp</emphasis>
: triangular point. Scale line: 0.6 mm.
</paragraph>
</caption>
<subSubSection id="87D86556323CF831B4CCBCBA13ADFF00" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105" type="description">
<paragraph id="CF7D36DD323CF830B4CCBCBA168CFEB8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBCBA165DFF48" bold="true" box="[199,345,151,176]" pageId="4" pageNumber="105">Description.</emphasis>
Body with 19 segments (including telson) in adults (
<typeStatus id="1079887F323CF830B7A2BCB5130FFF48" box="[937,1035,152,176]" pageId="4" pageNumber="105" type="paratype">paratype</typeStatus>
juvenile female with 18 segments).
<emphasis id="FDB6EACF323CF830B49CBC901646FF2C" box="[151,322,189,212]" italics="true" pageId="4" pageNumber="105">Measurements:</emphasis>
<typeStatus id="1079887F323CF830B541BC9116A8FF2D" box="[330,428,188,213]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
male
<quantity id="083A9B38323CF830B5F9BC911553FF2C" box="[498,599,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="105" unit="mm" value="17.8">17.8 mm</quantity>
long, width of midbody pro- and metazonites (segment nine) 1.0 mm and
<quantity id="083A9B38323CF830B49CBCCD17F6FF0F" box="[151,242,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="105" unit="mm" value="1.4">1.4 mm</quantity>
; allotype female
<quantity id="083A9B38323CF830B5BCBCCD1519FF0F" box="[439,541,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="4" pageNumber="105" unit="mm" value="17.9">17.9 mm</quantity>
long, width of midbody pro- and metazonites (segment nine) 1.0 mm and
<quantity id="083A9B38323CF830B172BCCD17C2FEE4" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="105" unit="mm" value="1.4">1.4 mm</quantity>
;
<typeStatus id="1079887F323CF830B4D9BD281637FEE5" box="[210,307,261,285]" pageId="4" pageNumber="105" type="paratype">paratype</typeStatus>
juvenile female
<quantity id="083A9B38323CF830B5FABD291552FEE4" box="[497,598,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="4" pageNumber="105" unit="mm" value="12.9">12.9 mm</quantity>
long, width of midbody pro- and metazonites (segment nine)
<quantity id="083A9B38323CF830B11EBD291268FEE4" box="[1301,1388,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="105" unit="mm" value="0.8">0.8 mm</quantity>
and
<quantity id="083A9B38323CF830B49CBD0517EBFEC7" box="[151,239,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="105" unit="mm" value="1.1">1.1 mm</quantity>
, respectively.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCBD61128AFE9C" blockId="4.[151,1437,151,2013]" box="[199,1422,332,357]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBD61164EFE9D" box="[199,330,332,357]" italics="true" pageId="4" pageNumber="105">Coloration:</emphasis>
Body whitish, slightly translucent, with a visible gut; head light brownish with whitish antennae.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCBD4215A4FCC0" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBD421609FE70" box="[199,269,367,392]" italics="true" pageId="4" pageNumber="105">Head:</emphasis>
Broader than collum, covered with fine short and some longer setae. Occipital suture distinct. Labrum with only one long, medial, laterally flattened and ventrally curved tooth; two paramedian, short, blunt setae, one on each side of the tooth. Tooth significantly darker than labrum (Fig. 12). Gnathochilarium (in allotype female) densely setose. Setae robust. Promentum almost triangular, with four setae in one medial row. Stipites with more than 30 setae each. Lingual plates with about 20 setae each, with a long, paramedian, subapical seta on each plate and with one long, paramedian, apical, setiform, distally branching process on each plate. External palps with 18+18, medial palps with 15+15, internal palps with 4+4 papillae. Internal palps bearing one long seta crossing medially. Hypopharynx with strongly developed, long and thin fringes (
<figureCitation id="57F92A58323CF830B7CFBE411312FD7C" box="[964,1046,620,645]" captionStart="FIGURES 11 and 12" captionStartId="5.[151,264,1741,1763]" captionTargetBox="[151,1436,287,1717]" captionTargetId="figure@5.[151,1436,283,1717]" captionTargetPageId="5" captionText="FIGURES 11 and 12. Balkanodesmus biokovensis gen. n., sp. n., allotype female. 11: dorsal view of gnathochilarium and hypopharinx, 12: labrum. Scale line: 0.3 mm" httpUri="https://zenodo.org/record/251044/files/figure.png" pageId="4" pageNumber="105">Fig. 11</figureCitation>
). Antennae elongated; in
<typeStatus id="1079887F323CF830B131BE411298FD7D" box="[1338,1436,620,645]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
male
<quantity id="083A9B38323CF830B4EBBEA21645FD5F" box="[224,321,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="105" unit="mm" value="2.5">2.5 mm</quantity>
long. Antennomeres II–VI club-like, elongated; antennomere VII cylindrical and shorter. Antennomeres V and VI with few longer setae; antennomere VI with a distodorsal pit with numerous bacilliform sensilla. Antennomere VII with one longer seta and a distodorsal pad with few setiform setae. Apical part of antennae with four large apical cones. Length/width ratios of antennomeres I–VII: I (0.7), II (3.1), III (3.5), IV (2.5), V (2.1), VI (1.7), VII (1.7), respectively.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCBF6916F8FC78" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBF691626FCA5" box="[199,290,836,861]" italics="true" pageId="4" pageNumber="105">Collum:</emphasis>
Transversely subovoid; one caudal incision on lateral sides, with three rows of setae on small tubercles, setal formula: 8+8+8.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCBFA11582FAAD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBFA11676FC5C" box="[199,370,908,933]" italics="true" pageId="4" pageNumber="105">Body segments</emphasis>
: Segments II–IV with three rows of medium-sized trichoid setae on small tubercles while segments V–XVIII with mainly four, partly five, irregular transverse rows of setae. Pleurosternal carinae with welldeveloped triangular teeth present on each side of segments II–XVII. Apodous (XVIII) segment without pleurosternal carinae. Number of teeth variable not only between specimens and segments of same specimen, but also between right and left sides of same segment. Number of teeth in
<typeStatus id="1079887F323CF830B7A5B8311314FBCD" box="[942,1040,1052,1077]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
male: segments II and III (6/6); IV (5/5); V and VI (7–9); VII (5/7); VIII–XI (8–10); XII–XIV (10–12); XV–XVII (8–9), respectively. Prozonite slightly punctate. Stricture distinct. Paraterga with three dorsolaterally positioned, laterally directed cones; each bearing one seta. Last cone on each poriferous metazonite enlarged and bearing an ozopore and 3–4 setae. These enlarged cones also contain reactivation chambers of the defense mechanism. Pore formula normal: V, VII, IX, X, XII, XIII, XV–XVIII. Epiproct short, but protruding beyond paraprocts, slightly curved caudoventrally; subtriangular in dorsal view, truncate at tip, dorsally with long setae. Tip of epiproct with 2+2 spinnerets. Paraprocts semi-circular, smooth, each with two setae. Hypoproct subtrapeziform, with four long apical setae. Sterna unmodified. Epigynal ridge elevated.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCB9721653FA65" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCB972165BFA80" box="[199,351,1375,1400]" italics="true" pageId="4" pageNumber="105">Walking legs:</emphasis>
Long and slender, without modified setae. Femora and tarsi elongated, claws medium-sized and clearly infuscate.
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCB98A121FF8FC" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCB98A1647FA38" box="[199,323,1447,1472]" italics="true" pageId="4" pageNumber="105">Gonopods:</emphasis>
Gonopod aperture large, transverse-oval, taking up
<emphasis id="FDB6EACF323CF830B7AFB98414D9FA38" box="[932,989,1449,1472]" italics="true" pageId="4" pageNumber="105">circa</emphasis>
2/3 of ventral part of metazonite VII. Gonocoxa large, exposed, subglobose, with two distofrontal setae and one distocaudal seta, with neither additional processes nor protruding parts. Prefemoral part (
<emphasis id="FDB6EACF323CF830B6B6B9C215D2F9F0" bold="true" box="[701,726,1519,1544]" pageId="4" pageNumber="105">pf</emphasis>
) slender, held transversely to body axis, curved caudally, with almost straight margins. First 1/3 smooth, second 1/3 only with apical setae, remaining 1/3 densely setose. Acropodite (
<emphasis id="FDB6EACF323CF830B522BA1A1633F9A8" bold="true" box="[297,311,1591,1616]" pageId="4" pageNumber="105">a</emphasis>
) erect, long, unipartite and parallel-sided; directed frontally. Basal part with setae and a narrow, longitudinal, mesal lamella (
<emphasis id="FDB6EACF323CF830B5D5BA7116E9F98D" bold="true" box="[478,493,1628,1653]" pageId="4" pageNumber="105">ll</emphasis>
) reaching the first, most shallow and widest (basal) lamella (
<emphasis id="FDB6EACF323CF830B0BBBA7113C3F98D" bold="true" box="[1200,1223,1628,1653]" pageId="4" pageNumber="105">bl</emphasis>
) in middle part of acropodite. A second, deeper and narrower (middle) lamella (
<emphasis id="FDB6EACF323CF830B75BBA52146BF960" bold="true" box="[848,879,1663,1688]" pageId="4" pageNumber="105">ml</emphasis>
) above
<emphasis id="FDB6EACF323CF830B7C0BA5214E7F960" bold="true" box="[971,995,1663,1688]" pageId="4" pageNumber="105">bl</emphasis>
and the narrowest and deepest (apical) lamella (
<emphasis id="FDB6EACF323CF830B4F0BA891615F945" bold="true" box="[251,273,1700,1725]" pageId="4" pageNumber="105">al</emphasis>
) above
<emphasis id="FDB6EACF323CF830B564BA89168AF945" bold="true" box="[367,398,1700,1725]" pageId="4" pageNumber="105">ml</emphasis>
. Seminal canal probably opening at base of
<emphasis id="FDB6EACF323CF830B787BA8914A6F945" bold="true" box="[908,930,1700,1725]" pageId="4" pageNumber="105">al</emphasis>
. Solenomere absent. Apart from
<emphasis id="FDB6EACF323CF830B114BA89123AF945" bold="true" box="[1311,1342,1700,1725]" pageId="4" pageNumber="105">ml</emphasis>
, middle part of acropodite bearing characteristic, short, triangular denticles (
<emphasis id="FDB6EACF323CF830B7A8BAEA14B6F918" bold="true" box="[931,946,1735,1760]" pageId="4" pageNumber="105">d</emphasis>
) associated with
<emphasis id="FDB6EACF323CF830B077BAEA139FF918" bold="true" box="[1148,1179,1735,1760]" pageId="4" pageNumber="105">ml</emphasis>
and
<emphasis id="FDB6EACF323CF830B0DDBAEA13EFF918" bold="true" box="[1238,1259,1735,1760]" pageId="4" pageNumber="105">al</emphasis>
. Apical part of acropodite long and slender, with two apical lobes (
<emphasis id="FDB6EACF323CF830B6D2BAC115EBF8FD" bold="true" box="[729,751,1772,1797]" pageId="4" pageNumber="105">lo</emphasis>
) and a mesally directed, triangular, acute tip (
<emphasis id="FDB6EACF323CF830B0F9BAC1120FF8FD" bold="true" box="[1266,1291,1772,1797]" pageId="4" pageNumber="105">tp</emphasis>
).
</paragraph>
<paragraph id="CF7D36DD323CF830B4CCBB221298F824" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="105">
<emphasis id="FDB6EACF323CF830B4CCBB221628F8D0" bold="true" box="[199,300,1807,1832]" pageId="4" pageNumber="105">Habitat:</emphasis>
Amfora Pit, where
<taxonomicName id="08C24D5E323CF830B604BB22144EF8D0" box="[527,842,1807,1832]" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="FDB6EACF323CF830B604BB22144EF8D0" box="[527,842,1807,1832]" italics="true" pageId="4" pageNumber="105">Balkanodesmus biokovensis</emphasis>
</taxonomicName>
was collected, is a 788-meter deep. The entrance of the pit is situated at an altitude of
<quantity id="083A9B38323CF830B621BB191588F8B4" box="[554,652,1844,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="4" pageNumber="105" unit="m" value="1620.0">1,620 m</quantity>
. All three specimens were collected between a depth of 100 to 400 meters. The almost stable air temperature ranged from 5.1 to 5.4°C (
<bibRefCitation id="AB534B2C323CF830B7B1BB7A138EF888" author="Lukic" box="[954,1162,1879,1904]" pageId="4" pageNumber="105" refString="Lukic, M., Houssin, C. &amp; Deharveng, L. (2010) A new relictual and highly troglomorphic species of Tomoceridae (Collembola) from a deep Croatian cave. ZooKeys, 69, 1 - 16. http: // dx. doi. org / 10.3897 / zookeys. 69.739" type="journal article" year="2010">
Lukić
<emphasis id="FDB6EACF323CF830B00CBB741338F888" box="[1031,1084,1879,1904]" italics="true" pageId="4" pageNumber="105">et al</emphasis>
. 2010
</bibRefCitation>
). This pit hosts another endemic troglobitic diplopod,
<taxonomicName id="08C24D5E323CF830B5F0BB511482F86C" authority="Mrsic, 1992" authorityName="Mrsic" authorityYear="1992" box="[507,902,1916,1941]" class="Diplopoda" family="Biokoviellidae" genus="Biokoviella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="mauriesi">
<emphasis id="FDB6EACF323CF830B5F0BB5115EDF86C" box="[507,745,1916,1941]" italics="true" pageId="4" pageNumber="105">Biokoviella mauriesi</emphasis>
<bibRefCitation id="AB534B2C323CF830B6FDBB511482F86C" author="Mrsic" box="[758,902,1916,1941]" pageId="4" pageNumber="105" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subfam. nov. (Anthogonidae) and superfamily Cleidogonoidea (Craspedosomidea, Diplopoda) of the western Balkans. Razprave IV." type="book" year="1992">Mršić, 1992</bibRefCitation>
</taxonomicName>
. Amfora Pit is also the
<typeStatus id="1079887F323CF830B0A4BB5013DBF86D" box="[1199,1247,1917,1941]" pageId="4" pageNumber="105">type</typeStatus>
locality for the interesting, highly troglomorphic collembolan,
<taxonomicName id="08C24D5E323CF830B6A5BB8C17D8F824" authority="Lukic, Houssin &amp; Deharveng, 2010 (Lukic et al. 2010)" authorityName="Lukic, Houssin &amp; Deharveng, 2010 (Lukic et al." authorityYear="2010" class="Entognatha" family="Tomoceridae" genus="Tritomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="veles">
<emphasis id="FDB6EACF323CF830B6A5BB8C146FF841" box="[686,875,1952,1977]" italics="true" pageId="4" pageNumber="105">Tritomurus veles</emphasis>
<bibRefCitation id="AB534B2C323CF830B778BB8D1203F840" author="Lukic" box="[883,1287,1952,1977]" pageId="4" pageNumber="105" refString="Lukic, M., Houssin, C. &amp; Deharveng, L. (2010) A new relictual and highly troglomorphic species of Tomoceridae (Collembola) from a deep Croatian cave. ZooKeys, 69, 1 - 16. http: // dx. doi. org / 10.3897 / zookeys. 69.739" type="journal article" year="2010">Lukić, Houssin &amp; Deharveng, 2010</bibRefCitation>
(
<bibRefCitation id="AB534B2C323CF830B11CBB8D17D0F824" author="Lukic" pageId="4" pageNumber="105" refString="Lukic, M., Houssin, C. &amp; Deharveng, L. (2010) A new relictual and highly troglomorphic species of Tomoceridae (Collembola) from a deep Croatian cave. ZooKeys, 69, 1 - 16. http: // dx. doi. org / 10.3897 / zookeys. 69.739" type="journal article" year="2010">
Lukić
<emphasis id="FDB6EACF323CF830B168BB8C1291F841" box="[1379,1429,1952,1977]" italics="true" pageId="4" pageNumber="105">et al</emphasis>
. 2010
</bibRefCitation>
)
</taxonomicName>
. Furthermore, other notable troglobitic taxa are present in the pit (
<bibRefCitation id="AB534B2C323CF830B7C5BBE91393F824" author="Lukic" box="[974,1175,1988,2013]" pageId="4" pageNumber="105" refString="Lukic, M., Houssin, C. &amp; Deharveng, L. (2010) A new relictual and highly troglomorphic species of Tomoceridae (Collembola) from a deep Croatian cave. ZooKeys, 69, 1 - 16. http: // dx. doi. org / 10.3897 / zookeys. 69.739" type="journal article" year="2010">
Lukić
<emphasis id="FDB6EACF323CF830B012BBE8134FF825" box="[1049,1099,1988,2013]" italics="true" pageId="4" pageNumber="105">et al</emphasis>
. 2010
</bibRefCitation>
), such as:
<taxonomicName id="08C24D5E323CF830B105BBE81277F824" box="[1294,1395,1989,2012]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="4" pageNumber="105" phylum="Mollusca" rank="genus">
<emphasis id="FDB6EACF323CF830B105BBE81277F824" box="[1294,1395,1989,2012]" italics="true" pageId="4" pageNumber="105">Zospeum</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph id="CF7D36DD323DF831B49CBCBA13ADFF00" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="106">
(Gastropoda),
<taxonomicName id="08C24D5E323DF831B55BBCBA147DFF48" authority="Muller, 1931" authorityName="Muller" authorityYear="1931" box="[336,889,151,176]" class="Arachnida" family="Neobisiidae" genus="Protoneobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="biocovense">
<emphasis id="FDB6EACF323DF831B55BBCBA1599FF48" box="[336,669,151,176]" italics="true" pageId="5" pageNumber="106">Protoneobisium biocovense</emphasis>
(Müller, 1931)
</taxonomicName>
,
<taxonomicName id="08C24D5E323DF831B797BCBA1246FF48" authority="Curcic, 1988" authorityName="Curcic" authorityYear="1988" box="[924,1346,151,176]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="peruni">
<emphasis id="FDB6EACF323DF831B797BCBA1378FF48" box="[924,1148,151,176]" italics="true" pageId="5" pageNumber="106">Neobisium peruni</emphasis>
Ćurčić, 1988
</taxonomicName>
(both
<taxonomicName id="08C24D5E323DF831B49CBC911660FF2D" box="[151,356,188,213]" class="Arachnida" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="106" phylum="Arthropoda" rank="order">Pseudoscorpiones</taxonomicName>
),
<taxonomicName id="08C24D5E323DF831B570BC9114A0FF2C" authority="Muller, 1934" authorityName="Muller" authorityYear="1934" box="[379,932,188,213]" class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis id="FDB6EACF323DF831B570BC911405FF2D" box="[379,769,188,213]" italics="true" pageId="5" pageNumber="106">Speoplanes giganteus biocovensis</emphasis>
Müller, 1934
</taxonomicName>
,
<taxonomicName id="08C24D5E323DF831B7BFBC911251FF2D" authority="Casale &amp; Jalzic,1989" authorityName="Casale &amp; Jalzic" authorityYear="1989" box="[948,1365,188,213]" class="Insecta" family="Leiodidae" genus="Radziella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="styx">
<emphasis id="FDB6EACF323DF831B7BFBC911351FF2C" box="[948,1109,188,213]" italics="true" pageId="5" pageNumber="106">Radziella styx</emphasis>
Casale &amp; Jalžić,1989
</taxonomicName>
(both
<taxonomicName id="08C24D5E323DF831B49CBCF21611FF00" box="[151,277,223,248]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="08C24D5E323DF831B52BBCF21696FF00" box="[288,402,223,248]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
) and
<taxonomicName id="08C24D5E323DF831B5C4BCF2148AFF00" authority="Jalzic, 1993" authorityName="Jalzic" authorityYear="1993" box="[463,910,223,248]" class="Insecta" family="Carabidae" genus="Biokovoaphaenopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="radici">
<emphasis id="FDB6EACF323DF831B5C4BCF21405FF00" box="[463,769,223,248]" italics="true" pageId="5" pageNumber="106">Biokovoaphaenopsis radici</emphasis>
Jalžić, 1993
</taxonomicName>
(
<taxonomicName id="08C24D5E323DF831B797BCF2131FFF00" box="[924,1051,223,248]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="08C24D5E323DF831B02CBCF2139AFF00" box="[1063,1182,223,248]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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