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<document id="28054221ACF31DC78770748DDCB81115" ID-DOI="10.11646/zootaxa.5371.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10144644" ID-ZooBank="D60B50D1-280B-4403-9E5B-25C0704A43A1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700160671039" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2023" docId="3E380C57FFE04A5427ACB34AFB90E7DB" docLanguage="en" docName="zootaxa.5371.1.1.pdf" docOrigin="Zootaxa 5371 (1)" docSource="https://mapress.com/zt/article/download/zootaxa.5371.1.1/52271" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scydmaenus (Choleropsis) princeps" docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="C201742FFFF54A43273BB26CFFDFE652" masterDocTitle="Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="22" updateTime="1700161323971" updateUser="ExternalLinkService">
<mods:mods id="9C9AD3413C43C722234CE17464A4A793" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="521E73887B9AE500687445CF52578459">Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="F9F89CCCB0150C71944503F88F1A18C6" type="personal">
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</mods:role>
<mods:namePart id="04DC2EE9A22C8168CEB582D8FBFF56FA">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="F9DB3CC4D7D8FD92410F0098584F48C1">text</mods:typeOfResource>
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<mods:title id="3F8A5BCC83304F1DC6411C5A25F6AAB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B869A29075E27B9BCCC2959F55DC5257">
<mods:date id="0AF18017EB836EA4E2490AD796377EC2">2023</mods:date>
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<mods:number id="FC07BCF01542D800EFD0FA4AD380E09F">2023-11-14</mods:number>
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<mods:detail id="5710207192E5EFEC879C172922194BCF" type="volume">
<mods:number id="D586FBAD308ABB0C2EA497F0DF8BA29F">5371</mods:number>
</mods:detail>
<mods:detail id="0B17A1FDBD6743357966B280DC328C63" type="issue">
<mods:number id="405FA28A69DED7C646F7397B606128B3">1</mods:number>
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<mods:start id="6057049DE1EF2378138D61C33AA0F119">1</mods:start>
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</mods:location>
<mods:classification id="5BF272A852CCB540D59FAC90941C6700">journal article</mods:classification>
<mods:identifier id="34178EC1E408FE9E202E6D12B03A41DD" type="DOI">10.11646/zootaxa.5371.1.1</mods:identifier>
<mods:identifier id="799E40710803F06AAB720E14C55551A4" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="681399D31B6D747DB962CC5213366AB3" type="Zenodo-Dep">10144644</mods:identifier>
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</mods:mods>
<treatment id="3E380C57FFE04A5427ACB34AFB90E7DB" LSID="urn:lsid:plazi:treatment:3E380C57FFE04A5427ACB34AFB90E7DB" httpUri="http://treatment.plazi.org/id/3E380C57FFE04A5427ACB34AFB90E7DB" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="FE8BEECAFFE04A5627ACB34AFD57E712" box="[151,648,294,321]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="B62EBD41FFE04A5627ACB34AFD57E712" blockId="21.[151,648,294,321]" box="[151,648,294,321]" pageId="21" pageNumber="22">
<heading id="ED660A2DFFE04A5627ACB34AFD57E712" bold="true" box="[151,648,294,321]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<taxonomicName id="7191C6C2FFE04A5627ACB34AFD57E712" ID-CoL="4W3QW" authority="(King)" baseAuthorityName="King" baseAuthorityYear="1864" box="[151,648,294,321]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="princeps" subGenus="Choleropsis">
<emphasis id="84E56153FFE04A5627ACB34AFD57E712" bold="true" box="[151,648,294,321]" pageId="21" pageNumber="22">
<emphasis id="84E56153FFE04A5627ACB34AFEF7E713" bold="true" box="[151,296,294,321]" italics="true" pageId="21" pageNumber="22">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE04A562603B34AFE1EE713" bold="true" box="[312,449,294,321]" italics="true" pageId="21" pageNumber="22">Choleropsis</emphasis>
)
<emphasis id="84E56153FFE04A5626EAB34BFDECE713" bold="true" box="[465,563,295,321]" italics="true" pageId="21" pageNumber="22">princeps</emphasis>
(King)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE04A5627ACB307FDD8E7B7" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="B62EBD41FFE04A5627ACB307FDEFE7D1" blockId="21.[151,813,363,485]" box="[151,560,363,387]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="96819A6FFFE04A5627ACB307FDEFE7D1" box="[151,560,363,387]" pageId="21" pageNumber="22">
<treatmentCitation id="37309B50FFE04A5627ACB307FDF3E7D1" author="King, R. L." box="[151,556,363,387]" page="98" pageId="21" pageNumber="22" year="1864">
<taxonomicName id="7191C6C2FFE04A5627ACB307FDF3E7D1" authority="King, 1864: 98" authorityName="King" authorityPageNumber="98" authorityYear="1864" box="[151,556,363,387]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE04A5627ACB307FE55E7D1" box="[151,394,363,387]" italics="true" pageId="21" pageNumber="22">Heterognathus princeps</emphasis>
<bibRefCitation id="D200C0B0FFE04A5626AAB307FDF3E7D1" author="King, R. L." box="[401,556,363,387]" pageId="21" pageNumber="22" pagination="91 - 99" refId="ref51270" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99, pls. 6 - 7." type="journal article" year="1864">King, 1864: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFE04A5627ACB3E6FD3FE7F0" blockId="21.[151,813,363,485]" box="[151,736,394,418]" pageId="21" pageNumber="22">
<taxonomicName id="7191C6C2FFE04A5627ACB3E6FDECE7F0" ID-CoL="4W3QW" authority="(King)" baseAuthorityName="King" baseAuthorityYear="1864" box="[151,563,394,418]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="princeps" subGenus="Cholerus">
<emphasis id="84E56153FFE04A5627ACB3E6FECBE7F0" box="[151,276,394,418]" italics="true" pageId="21" pageNumber="22">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE04A562618B3E6FE5FE7F0" box="[291,384,394,418]" italics="true" pageId="21" pageNumber="22">Cholerus</emphasis>
)
<emphasis id="84E56153FFE04A5626B4B3E6FE39E7F3" box="[399,486,394,417]" italics="true" pageId="21" pageNumber="22">princeps</emphasis>
(King)
</taxonomicName>
; Csiki, 1819: 73.
</paragraph>
<paragraph id="B62EBD41FFE04A5627ACB3C4FCF2E793" blockId="21.[151,813,363,485]" box="[151,813,424,449]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="96819A6FFFE04A5627ACB3C4FCF2E793" box="[151,813,424,449]" pageId="21" pageNumber="22">
<treatmentCitation id="37309B50FFE04A5627ACB3C4FD05E793" author="Franz, H." box="[151,730,424,449]" page="291" pageId="21" pageNumber="22" year="1975">
<taxonomicName id="7191C6C2FFE04A5627ACB3C4FD05E793" ID-CoL="4W3QL" authority="Franz, 1975: 291" authorityName="Franz" authorityPageNumber="291" authorityYear="1975" box="[151,730,424,449]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="marwillumbali" subGenus="Cholerus">
<emphasis id="84E56153FFE04A5627ACB3C4FECBE792" box="[151,276,424,448]" italics="true" pageId="21" pageNumber="22">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE04A562618B3C4FE5FE792" box="[291,384,424,448]" italics="true" pageId="21" pageNumber="22">Cholerus</emphasis>
)
<emphasis id="84E56153FFE04A5626B4B3C4FDFBE792" box="[399,548,424,448]" italics="true" pageId="21" pageNumber="22">marwillumbali</emphasis>
<bibRefCitation id="D200C0B0FFE04A562511B3C5FD05E793" author="Franz, H." box="[554,730,424,449]" pageId="21" pageNumber="22" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 291</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="84E56153FFE04A5625D8B3C4FCF2E793" bold="true" box="[739,813,424,449]" pageId="21" pageNumber="22">Syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFE04A5627ACB3A7FDD8E7B7" blockId="21.[151,813,363,485]" box="[151,519,458,485]" pageId="21" pageNumber="22">
(
<figureCitation id="2EAAA1C4FFE04A5627A4B3A7FEFEE7B7" box="[159,289,458,485]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="21" pageNumber="22">Figs 35–40</figureCitation>
,
<figureCitation id="2EAAA1C4FFE04A562617B3A6FE95E7B7" box="[300,330,458,485]" captionStart="FIGURES 51–57" captionStartId="26.[151,264,1248,1272]" captionTargetBox="[178,1413,192,1220]" captionTargetId="figure-21@26.[164,1423,181,1224]" captionTargetPageId="26" captionText="FIGURES 51–57. Antennae of Choleropsis in dorsal view, males (51, 53–57) and female (52). Scydmaenus beechmonti Franz (51–52); S. geniculatus (King) (53); S. kurandae Franz (54); S. princeps (King) (55); S. curviclavatus sp. n. (56); S. kroombitanus sp. n. (57)." figureDoi="http://doi.org/10.5281/zenodo.10144668" httpUri="https://zenodo.org/record/10144668/files/figure.png" pageId="21" pageNumber="22">55</figureCitation>
,
<figureCitation id="2EAAA1C4FFE04A56266DB3A6FE41E7B7" box="[342,414,458,485]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="21" pageNumber="22">64–65</figureCitation>
,
<figureCitation id="2EAAA1C4FFE04A562691B3A7FE17E7B7" box="[426,456,459,485]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="21" pageNumber="22">73</figureCitation>
,
<figureCitation id="2EAAA1C4FFE04A5626EFB3A7FE20E7B7" box="[468,511,459,485]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="21" pageNumber="22">193</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE04A5527ACB07FFA9BE74F" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="B62EBD41FFE04A5627ACB07FFD53E4CB" blockId="21.[151,1436,530,665]" pageId="21" pageNumber="22">
<emphasis id="84E56153FFE04A5627ACB07FFE72E47E" bold="true" box="[151,429,530,557]" pageId="21" pageNumber="22">Type material studied.</emphasis>
<materialsCitation id="06F9B71CFFE04A562683B07FFDD7E427" collectionCode="AMS" country="Australia" location="Het" pageId="21" pageNumber="22" specimenCount="2" typeStatus="lectotype">
<emphasis id="84E56153FFE04A562683B07FFC08E47E" bold="true" box="[440,983,530,557]" pageId="21" pageNumber="22">
<typeStatus id="692A03E3FFE04A562683B07FFDF1E47F" box="[440,558,531,557]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
(here designated) (
<collectingCountry id="CE86FDD1FFE04A56241BB07EFC15E47E" box="[800,970,530,556]" name="Australia" pageId="21" pageNumber="22">AUSTRALIA</collectingCountry>
).
</emphasis>
♁, with labels illustrated in
<figureCitation id="2EAAA1C4FFE04A562209B07FFA47E47F" box="[1330,1432,531,557]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="21" pageNumber="22">Fig. 193</figureCitation>
: “K23219” [brownish, handwritten], “
<location id="B34EEB9AFFE04A56250CB05BFDBDE403" LSID="urn:lsid:plazi:treatment:3E380C57FFE04A5427ACB34AFB90E7DB:B34EEB9AFFE04A56250CB05BFDBDE403" box="[567,610,567,593]" country="Australia" name="Het" pageId="21" pageNumber="22">Het</location>
/ princeps” [brownish, handwritten], “K 197786” [white, printed], “
<typeStatus id="692A03E3FFE04A562262B05BFA43E403" box="[1369,1436,567,593]" pageId="21" pageNumber="22" type="syntype">SYN-</typeStatus>
TYPE” [yellow, printed] (
<collectionCode id="D0802584FFE04A562685B037FDDFE427" box="[446,512,603,629]" pageId="21" pageNumber="22">AMS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="06F9B71CFFE04A56252FB037FD57E4CB" collectionCode="AMS" country="Australia" location="Het" pageId="21" pageNumber="22" specimenCount="2" typeStatus="paralectotype">
<emphasis id="84E56153FFE04A56252FB037FD1FE427" bold="true" box="[532,704,603,629]" pageId="21" pageNumber="22">
<typeStatus id="692A03E3FFE04A56252FB037FD64E427" box="[532,699,603,629]" pageId="21" pageNumber="22" type="paralectotype">Paralectotype</typeStatus>
.
</emphasis>
♁ (
<figureCitation id="2EAAA1C4FFE04A5625D6B037FC9DE427" box="[749,834,603,629]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="21" pageNumber="22">Fig. 36</figureCitation>
), on separated mounting card, on the same pin under
<typeStatus id="692A03E3FFE04A5627ACB013FEDEE4CB" box="[151,257,639,665]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
, same data as for lectotype (
<collectionCode id="D0802584FFE04A562505B013FD5FE4CB" box="[574,640,639,665]" pageId="21" pageNumber="22">AMS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="E2EEEDC9FFE04A5627ACB401FF06E09F" ID-DOI="http://doi.org/10.5281/zenodo.10144662" ID-Zenodo-Dep="10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,264,1645,1669]" targetBox="[170,1417,695,1616]" targetPageId="21" targetType="figure">
<paragraph id="B62EBD41FFE04A5627ACB401FF06E09F" blockId="21.[151,1437,1645,1741]" pageId="21" pageNumber="22">
<emphasis id="84E56153FFE04A5627ACB401FE89E0D7" bold="true" box="[151,342,1645,1669]" pageId="21" pageNumber="22">FIGURES 35–40.</emphasis>
<taxonomicName id="7191C6C2FFE04A562666B401FCC6E0D7" authority="(King)" baseAuthorityName="King" baseAuthorityYear="1864" box="[349,793,1645,1670]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="princeps" subGenus="Choleropsis">
<emphasis id="84E56153FFE04A562666B401FE05E0D7" box="[349,474,1645,1669]" italics="true" pageId="21" pageNumber="22">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE04A5626D0B401FDBAE0D7" box="[491,613,1645,1669]" italics="true" pageId="21" pageNumber="22">Choleropsis</emphasis>
)
<emphasis id="84E56153FFE04A56254EB402FD13E0D7" box="[629,716,1646,1669]" italics="true" pageId="21" pageNumber="22">princeps</emphasis>
(King)
</taxonomicName>
, males. Lateral habitus of holotype of
<taxonomicName id="7191C6C2FFE04A562391B402FA42E0D7" authority="Franz" authorityName="Franz" authorityYear="1975" box="[1194,1437,1645,1669]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="marwillumbali">
<emphasis id="84E56153FFE04A562391B402FA85E0D7" box="[1194,1370,1645,1669]" italics="true" pageId="21" pageNumber="22">S. marwillumbali</emphasis>
Franz
</taxonomicName>
(
<emphasis id="84E56153FFE04A56279BB4FDFF66E0FB" bold="true" box="[160,185,1681,1705]" pageId="21" pageNumber="22">35</emphasis>
) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of
<taxonomicName id="7191C6C2FFE04A562423B4FEFC57E0FB" baseAuthorityName="King" baseAuthorityYear="1864" box="[792,904,1682,1705]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE04A562423B4FEFC57E0FB" box="[792,904,1682,1705]" italics="true" pageId="21" pageNumber="22">S. princeps</emphasis>
</taxonomicName>
(
<emphasis id="84E56153FFE04A5624ACB4FDFC6FE0FB" bold="true" box="[919,944,1681,1705]" pageId="21" pageNumber="22">36</emphasis>
); aedeagus in dorsal (
<emphasis id="84E56153FFE04A5623B4B4FDFB12E0FB" bold="true" box="[1167,1229,1681,1705]" pageId="21" pageNumber="22">37, 39</emphasis>
) and lateral (
<emphasis id="84E56153FFE04A56226EB4FDFA4CE0FB" bold="true" box="[1365,1427,1681,1705]" pageId="21" pageNumber="22">38, 40</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="B62EBD41FFE04A5627FCB497FBEEE1D0" blockId="21.[151,1437,1787,2030]" pageId="21" pageNumber="22">
<emphasis id="84E56153FFE04A5627FCB497FCD1E144" bold="true" box="[199,782,1787,1814]" pageId="21" pageNumber="22">Specimens studied as high-quality photographs.</emphasis>
<materialsCitation id="06F9B71CFFE04A562423B490FBEEE1D0" collectionCode="BM" collectorName="N. S. Wales &amp; Coll. F. Muir" country="Australia" determinerName="H. Franz" location="NEW SOUTH WALES" pageId="21" pageNumber="22" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="84E56153FFE04A562423B490FDE4E16B" bold="true" pageId="21" pageNumber="22">
<typeStatus id="692A03E3FFE04A562423B490FC5AE144" box="[792,901,1788,1814]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="7191C6C2FFE04A56248AB497FA97E144" authority="Franz" authorityName="Franz" authorityYear="1975" box="[945,1352,1787,1814]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="marwillumbali">
<emphasis id="84E56153FFE04A56248AB497FB29E144" bold="true" box="[945,1270,1787,1814]" italics="true" pageId="21" pageNumber="22">Scydmaenus marwillumbali</emphasis>
Franz
</taxonomicName>
(
<collectingCountry id="CE86FDD1FFE04A562262B497FED7E16B" name="Australia" pageId="21" pageNumber="22">AUSTRALIA</collectingCountry>
:
<collectingRegion id="745573A3FFE04A56262FB54CFDF0E16B" box="[276,559,1823,1850]" country="Australia" name="New South Wales" pageId="21" pageNumber="22">NEW SOUTH WALES</collectingRegion>
):
</emphasis>
♁ (
<figureCitation id="2EAAA1C4FFE04A56255CB573FD65E168" box="[615,698,1823,1850]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="21" pageNumber="22">Fig. 35</figureCitation>
), with labels illustrated in
<figureCitation id="2EAAA1C4FFE04A5624DFB573FB9AE168" box="[996,1093,1823,1850]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="21" pageNumber="22">Fig. 194</figureCitation>
: “Murwillumbah /
<collectorName id="1B64D897FFE04A562221B573FA4EE16B" box="[1306,1425,1823,1849]" pageId="21" pageNumber="22">N.S.Wales</collectorName>
” [white, handwritten], “
<collectorName id="1B64D897FFE04A5626AFB52FFDF9E10F" box="[404,550,1859,1886]" pageId="21" pageNumber="22">Coll. F. Muir</collectorName>
/ 801919” [white, printed and handwritten], “
<taxonomicName id="7191C6C2FFE04A56231AB52FFB6EE10F" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1057,1201,1859,1885]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ marwillumbali / m. / det.
<determinerName id="43F25185FFE04A5627F4B50BFEF3E1D3" box="[207,300,1895,1921]" pageId="21" pageNumber="22">H.Franz</determinerName>
” [white, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="D0802584FFE04A5624C8B504FBFBE1D0" box="[1011,1060,1896,1922]" country="United Kingdom" name="Bristol Museum" pageId="21" pageNumber="22">BM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B62EBD41FFE04A5527FCB5E7FA9BE74F" blockId="21.[151,1437,1787,2030]" lastBlockId="22.[151,1437,151,2013]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="84E56153FFE04A5627FCB5E7FDA6E1F7" bold="true" box="[199,633,1931,1958]" pageId="21" pageNumber="22">Revised diagnosis (based on males).</emphasis>
Antennomere 9 only slightly shorter and narrower than 10, with conspicuously deep and abrupt lateral concavity (
<figureCitation id="2EAAA1C4FFE04A56259EB5C3FD23E19B" box="[677,764,1967,1993]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="21" pageNumber="22">Fig. 64</figureCitation>
); antennomere 10 about as long as broad and only slightly impressed (
<figureCitation id="2EAAA1C4FFE04A562622B5BFFE41E1BC" box="[281,414,2003,2030]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="21" pageNumber="22">Figs 64–65</figureCitation>
); aedeagus in dorsal view (
<figureCitation id="2EAAA1C4FFE04A5625E9B5BFFC8BE1BC" box="[722,852,2003,2030]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="21" pageNumber="22">Figs 37, 39</figureCitation>
) with apical margin distinctly concave and as wide as about half of total width of median lobe, lateral subapical lobes prominent, subtriangular, in lateral view (
<figureCitation id="2EAAA1C4FFE34A552251B2FBFF00E687" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Figs 38, 40</figureCitation>
) distal region of median lobe strongly broadened, with dorsal margin strongly rounded in proximal half and weakly concave in distal half; metaventrite (
<figureCitation id="2EAAA1C4FFE34A5525BDB2B3FD07E6AB" box="[646,728,223,249]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="22" pageNumber="23">Fig. 73</figureCitation>
) with shallow and diffuse median impression about as broad as half width of metaventrite and filled with short recumbent setae not differing from those on lateral regions.
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE34A5527FCB34AFE7DE11F" pageId="22" pageNumber="23" type="description">
<paragraph id="B62EBD41FFE34A5527FCB34AFB1AE7DB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="84E56153FFE34A5527FCB34AFEAAE712" bold="true" box="[199,373,294,320]" pageId="22" pageNumber="23">Redescription.</emphasis>
Body of male (
<figureCitation id="2EAAA1C4FFE34A552519B34BFD7EE713" box="[546,673,294,321]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Figs 35–36</figureCitation>
) slightly flattened, elongate and slender, BL
<quantity id="716910A4FFE34A5523ACB34AFB22E713" box="[1175,1277,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="22" pageNumber="23" unit="mm" value="1.75">1.75 mm</quantity>
; pigmentation uniformly light to dark brown, appendages (especially antennae and palps) indistinctly lighter (difference best visible in darkest specimens); cuticle moderately glossy, covered with vestiture of yellowish setae.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB3FFFD3AE5F7" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Head (
<figureCitation id="2EAAA1C4FFE34A55262AB3FFFEBDE7FF" box="[273,354,403,429]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Fig. 36</figureCitation>
) in dorsal view transversely subrectangular, broadest at eyes, HL
<quantity id="716910A4FFE34A552304B3FFFB7AE7FF" box="[1087,1189,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="716910A4FFE34A5523DEB3FEFA94E7FF" box="[1253,1355,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="22" pageNumber="23" unit="mm" value="0.35">0.35 mm</quantity>
; vertex and frons confluent and weakly convex, posterior margin of vertex distinctly concave, arcuate; tempora slightly longer than 3 × length of eye in dorsal view; supraantennal tubercles indistinct; frons over antennal fossae broadly subtrapezoidal and with straight anterior margin. Eyes conspicuously small, nearly semicircular in shape, indistinctly emarginate posteriorly and oblique in relation to long axis of head. Punctures on frons and vertex fine, inconspicuous; setae short, sparse, suberect, those on tempora as sparse as those on head dorsum. Genae as sparsely setose as frons and vertex. Anterior (exposed) region of head capsule demarcated from neck region by narrow abrupt impression around occipital constriction, anterior margin of gular plate on neck region accentuated by minute rounded anteriorly-directed projection. Antennae (
<figureCitation id="2EAAA1C4FFE34A552556B0DFFD2FE49F" box="[621,752,690,717]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Figs 35–36</figureCitation>
,
<figureCitation id="2EAAA1C4FFE34A5525C6B0DEFCC4E49F" box="[765,795,690,717]" captionStart="FIGURES 51–57" captionStartId="26.[151,264,1248,1272]" captionTargetBox="[178,1413,192,1220]" captionTargetId="figure-21@26.[164,1423,181,1224]" captionTargetPageId="26" captionText="FIGURES 51–57. Antennae of Choleropsis in dorsal view, males (51, 53–57) and female (52). Scydmaenus beechmonti Franz (51–52); S. geniculatus (King) (53); S. kurandae Franz (54); S. princeps (King) (55); S. curviclavatus sp. n. (56); S. kroombitanus sp. n. (57)." figureDoi="http://doi.org/10.5281/zenodo.10144668" httpUri="https://zenodo.org/record/10144668/files/figure.png" pageId="22" pageNumber="23">55</figureCitation>
,
<figureCitation id="2EAAA1C4FFE34A552412B0DEFCAEE49F" box="[809,881,690,717]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="22" pageNumber="23">64–65</figureCitation>
) long and slender, AnL
<quantity id="716910A4FFE34A5523B3B0DEFB2EE49F" box="[1160,1265,690,717]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="22" pageNumber="23" unit="mm" value="0.95">0.95 mm</quantity>
; three terminal antennomeres forming sharply delimited club; scape about 3 times as long as broad, distinctly broadening distally; pedicel twice as long as broad; antennomeres 3–4 each nearly 1.5 × as long as broad, 5 twice as long as broad, 6 nearly symmetrical, about as long as broad, 7 and 8 each distinctly transverse and strongly asymmetrical, 9 strongly asymmetrical, with deep and abrupt lateral concavity, 10 slightly broader and indistinctly longer than 9, weakly asymmetrical, with shallow longitudinal (slightly oblique) impression; 11 much shorter than 9 and 10 combined, nearly twice as long as broad, slightly asymmetrical.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB1C3FAEDE22F" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Pronotum in dorsal view (
<figureCitation id="2EAAA1C4FFE34A5526CCB1C3FD94E59B" box="[503,587,943,969]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Fig. 36</figureCitation>
) elongate, broadest near anterior third, PL
<quantity id="716910A4FFE34A55237AB1C3FB75E59B" box="[1089,1194,943,969]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="22" pageNumber="23" unit="mm" value="0.48">0.48 mm</quantity>
, PW
<quantity id="716910A4FFE34A5523D7B1C3FA89E59B" box="[1260,1366,943,969]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="22" pageNumber="23" unit="mm" value="0.43">0.43 mm</quantity>
; anterior and lateral margins confluent and rounded; posterior corners obtuse-angled and blunt; posterior margin nearly straight; base with narrow but distinct posterior marginal carina and with two pairs of small, distinct, sharply marked and relatively deep antebasal pits, each distinctly transverse. Pronotal disc covered with fine and inconspicuous punctures; setae similar to those on frons and vertex, short, moderately dense, suberect. Ventrally prothorax with nearly asetose and impunctate hypomera; basisternal region sparsely covered with short recumbent setae.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB6EBFAFBE2BB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Elytra (
<figureCitation id="2EAAA1C4FFE34A552627B6EBFEB0E2F3" box="[284,367,1159,1185]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Fig. 36</figureCitation>
) oval, broadest slightly in front of middle, EL
<quantity id="716910A4FFE34A5524BFB6EBFC34E2F3" box="[900,1003,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="22" pageNumber="23" unit="mm" value="0.98">0.98 mm</quantity>
, EW
<quantity id="716910A4FFE34A552310B6EBFB4DE2F3" box="[1067,1170,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="22" pageNumber="23" unit="mm" value="0.68">0.68 mm</quantity>
, EI 1.44. Humeral calli weakly elevated; basal impression on each elytron barely marked; basal elytral foveae lacking; apices separately rounded. Punctures fine and inconspicuous; setae similar to those on pronotum. Hind wings not studied.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB69FFBCEE363" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Metaventrite (
<figureCitation id="2EAAA1C4FFE34A55265EB69FFE6AE35F" box="[357,437,1267,1293]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="22" pageNumber="23">Fig. 73</figureCitation>
) with shallow and diffuse median impression about as broad as half width of metaventrite and filled with short recumbent setae not differing from those on lateral regions
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB757FDF7E003" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Legs (
<figureCitation id="2EAAA1C4FFE34A552635B757FEBFE307" box="[270,352,1339,1365]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Fig. 36</figureCitation>
) long and slender; unmodified, except for distal penicillus on mesotibiae and weakly broadened protarsomere 1 and ventral tenent setae distributed on protarsomeres 1–3; protarsi particularly short, tarsomere 1 about 1.5 × as long as broad, 2–4 each about as long as broad, tarsomere 5 3 × as long as broad and slightly shorter than tarsomeres 2–4 combined; mesotarsi long, tarsomere 1 nearly 4 × as long as broad and subequal in length to tarsomeres 2–4 combined, tarsomeres 2–4 each about 1.5 × as long as broad, tarsomere 5 nearly 3 × as long as broad, subequal in length to tarsomeres 3–4 combined; metatarsi as long as mesotarsi, tarsomere 1 about 3. × as long as broad, tarsomeres 2–4 each elongate but decreasing in length distally, tarsomere 5 3 × as long as broad, subequal in length to tarsomeres 3–4 combined.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB437FB75E17B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Aedeagus (
<figureCitation id="2EAAA1C4FFE34A55267EB437FE1AE027" box="[325,453,1627,1653]" captionStart="FIGURES 35–40" captionStartId="21.[151,264,1645,1669]" captionTargetBox="[170,1417,695,1616]" captionTargetId="figure-135@21.[164,1423,689,1622]" captionTargetPageId="21" captionText="FIGURES 35–40. Scydmaenus (Choleropsis) princeps (King), males. Lateral habitus of holotype of S. marwillumbali Franz (35) [© Lisha Jasper, (BM)]; dorsal habitus of paralectotype of S. princeps (36); aedeagus in dorsal (37, 39) and lateral (38, 40) views." figureDoi="http://doi.org/10.5281/zenodo.10144662" httpUri="https://zenodo.org/record/10144662/files/figure.png" pageId="22" pageNumber="23">Figs 37–40</figureCitation>
) elongate but not very slender, AeL
<quantity id="716910A4FFE34A552466B437FC1CE027" box="[861,963,1627,1653]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="22" pageNumber="23" unit="mm" value="0.28">0.28 mm</quantity>
, in dorsal view median lobe broadest near distal third; apical region subtrapezoidal, strongly tapering and with strongly concave apical margin about as wide as half of total width of median lobe; lateral subapical lobes prominent and sharply demarcated from median lobe, each subtriangular, about as long as wide, strongly projecting laterally, flagellum with abruptly broadened and asymmetrically coiled vesicular structure; median lobe in lateral view with distinctly broadened distal portion, its dorsal surface relatively strongly rounded in proximal half and weakly concave in distal half.
</paragraph>
<paragraph id="B62EBD41FFE34A5527FCB55FFE7DE11F" blockId="22.[151,1437,151,2013]" box="[199,418,1843,1869]" pageId="22" pageNumber="23">
<emphasis id="84E56153FFE34A5527FCB55FFEFAE11F" bold="true" box="[199,293,1843,1869]" pageId="22" pageNumber="23">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE34A5527FCB53AFF0FE1C7" pageId="22" pageNumber="23" type="distribution">
<paragraph id="B62EBD41FFE34A5527FCB53AFF0FE1C7" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="84E56153FFE34A5527FCB53AFEBEE122" bold="true" box="[199,353,1878,1904]" pageId="22" pageNumber="23">Distribution.</emphasis>
CE and
<collectingRegion id="745573A3FFE34A5526FAB53BFD8DE123" box="[449,594,1879,1905]" country="Australia" name="Western Australia" pageId="22" pageNumber="23">SE Australia</collectingRegion>
: eastern
<collectingRegion id="745573A3FFE34A552588B53BFCA5E123" box="[691,890,1879,1905]" country="Australia" name="New South Wales" pageId="22" pageNumber="23">New South Wales</collectingRegion>
(known only from vicinity of Brisbane and Sydney).
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE34A5427FCB5F3FB90E7DB" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="discussion">
<paragraph id="B62EBD41FFE34A5527FCB5F3FE09E18F" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="84E56153FFE34A5527FCB5F3FEE4E1EB" bold="true" box="[199,315,1951,1977]" pageId="22" pageNumber="23">Remarks.</emphasis>
The species redescribed by
<bibRefCitation id="D200C0B0FFE34A552547B5F3FCD1E1EB" author="Franz, H." box="[636,782,1950,1977]" pageId="22" pageNumber="23" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
as
<taxonomicName id="7191C6C2FFE34A55240DB5F3FC6CE1EA" baseAuthorityName="King" baseAuthorityYear="1864" box="[822,947,1951,1977]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE34A55240DB5F3FC6CE1EA" box="[822,947,1951,1977]" italics="true" pageId="22" pageNumber="23">S. princeps</emphasis>
</taxonomicName>
on p. 288 is in fact
<taxonomicName id="7191C6C2FFE34A5523A2B5F3FAE8E1EB" box="[1177,1335,1951,1977]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE34A5523A2B5F3FAE8E1EB" box="[1177,1335,1951,1977]" italics="true" pageId="22" pageNumber="23">S. geniculatus</emphasis>
</taxonomicName>
(see Remarks for the latter species).
</paragraph>
<paragraph id="B62EBD41FFE24A5427FCB2FBFBA2E6AB" blockId="23.[151,1437,151,393]" pageId="23" pageNumber="24">
As many Franz’s names, also
<taxonomicName id="7191C6C2FFE24A542522B2FBFD05E6E3" authorityName="Franz" authorityYear="1975" box="[537,730,151,177]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="marwillumbali">
<emphasis id="84E56153FFE24A542522B2FBFD05E6E3" box="[537,730,151,177]" italics="true" pageId="23" pageNumber="24">S. marwillumbali</emphasis>
</taxonomicName>
was based on a misspelled locality, in this case the town Murwillumbah in the Northern Rivers. The antennal modification in
<taxonomicName id="7191C6C2FFE24A542449B2D7FC30E686" baseAuthorityName="King" baseAuthorityYear="1864" box="[882,1007,187,213]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE24A542449B2D7FC30E686" box="[882,1007,187,213]" italics="true" pageId="23" pageNumber="24">S. princeps</emphasis>
</taxonomicName>
and
<taxonomicName id="7191C6C2FFE24A54231CB2D7FB36E687" authorityName="Franz" authorityYear="1975" box="[1063,1257,187,213]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="marwillumbali">
<emphasis id="84E56153FFE24A54231CB2D7FB36E687" box="[1063,1257,187,213]" italics="true" pageId="23" pageNumber="24">S. marwillumbali</emphasis>
</taxonomicName>
is identical and unique among all
<taxonomicName id="7191C6C2FFE24A54265AB2B3FE38E6AB" box="[353,487,223,249]" pageId="23" pageNumber="21" rank="subGenus" subGenus="Choleropsis">
<emphasis id="84E56153FFE24A54265AB2B3FE38E6AB" box="[353,487,223,249]" italics="true" pageId="23" pageNumber="24">Choleropsis</emphasis>
</taxonomicName>
species, and these names are unambiguously synonymous.
</paragraph>
<paragraph id="B62EBD41FFE24A5427FCB36FFB90E7DB" blockId="23.[151,1437,151,393]" pageId="23" pageNumber="24">
<taxonomicName id="7191C6C2FFE24A5427FCB36FFE66E74E" baseAuthorityName="King" baseAuthorityYear="1864" box="[199,441,259,285]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE24A5427FCB36FFE66E74E" box="[199,441,259,285]" italics="true" pageId="23" pageNumber="24">Scydmaenus princeps</emphasis>
</taxonomicName>
is distinct within
<taxonomicName id="7191C6C2FFE24A5425B9B36FFCD7E74F" box="[642,776,259,285]" pageId="23" pageNumber="21" rank="subGenus" subGenus="Choleropsis">
<emphasis id="84E56153FFE24A5425B9B36FFCD7E74F" box="[642,776,259,285]" italics="true" pageId="23" pageNumber="24">Choleropsis</emphasis>
</taxonomicName>
and can be easily distinguished from all remaining species by a deep lateral cavity of the antennomere 9. Only
<taxonomicName id="7191C6C2FFE24A5425EBB34BFC5BE713" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[720,900,295,321]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="kroombitanus" status="sp. n.">
<emphasis id="84E56153FFE24A5425EBB34BFC5BE713" box="[720,900,295,321]" italics="true" pageId="23" pageNumber="24">S. kroombitanus</emphasis>
</taxonomicName>
<emphasis id="84E56153FFE24A5424B1B34BFC17E713" bold="true" box="[906,968,295,321]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="9FD6DC28FFE24A5424B1B34BFC17E713" box="[906,968,295,321]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a somewhat similarly modified antennal club, although the shape of both the antennomere 9 and 10 is different, and in
<taxonomicName id="7191C6C2FFE24A542314B327FB74E736" baseAuthorityName="King" baseAuthorityYear="1864" box="[1071,1195,331,357]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE24A542314B327FB74E736" box="[1071,1195,331,357]" italics="true" pageId="23" pageNumber="24">S. princeps</emphasis>
</taxonomicName>
the antennomere 5 is strongly elongate, whereas in
<taxonomicName id="7191C6C2FFE24A5426DDB303FD44E7DB" authorityName="Jałoszyński" authorityYear="2023" box="[486,667,367,393]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="kroombitanus">
<emphasis id="84E56153FFE24A5426DDB303FD44E7DB" box="[486,667,367,393]" italics="true" pageId="23" pageNumber="24">S. kroombitanus</emphasis>
</taxonomicName>
the antennomere 5 is as long as broad.
</paragraph>
</subSubSection>
</treatment>
</document>
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