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<document id="7A6780393D8FA90900BFAA9269195570" ID-CLB-Dataset="4190" ID-DOI="10.5852/ejt.2021.766.1483" ID-GBIF-Dataset="d2ee0981-75c7-4e5c-9618-2d9dd18ac823" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514587" ID-ZooBank="19110F18-6E35-4C50-B362-7BC742B0B5D5" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1631927040333" checkinUser="felipe" docAuthor="Goharimanesh, Mona, Stöhr, Sabine, Mirshamsi, Omid, Ghassemzadeh, Fereshteh &amp; Adriaens, Dominique" docDate="2021" docId="3B7687F64204624FD67FFA9DFA9AFC6D" docLanguage="en" docName="ejt-766_1-63.pdf" docOrigin="European Journal of Taxonomy 766 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.766.1483" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiolepididae Ljungman 1867" docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="C74FFF8E42226268D457FF89FF82FFB0" masterDocTitle="Interactive identification key to all brittle star families (Echinodermata; Ophiuroidea) leads to revised morphological descriptions" masterLastPageNumber="63" masterPageNumber="1" pageNumber="39" updateTime="1699449478631" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="960D63D8492532550FFBFACACCFF364F">Interactive identification key to all brittle star families (Echinodermata; Ophiuroidea) leads to revised morphological descriptions</mods:title>
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<mods:namePart id="9D49CCA0C349DC73CAFB2D7C7F1E90F7">Goharimanesh, Mona</mods:namePart>
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<mods:affiliation id="6484893103C6D2D8FDFB309A19D6C670">Ferdowsi University of Mashhad, Faculty of Science, Department of Biology, Mashhad, Iran. &amp; Ghent University, Research Group Evolutionary Morphology of Vertebrates, Gent, Belgium. &amp; Mona.</mods:affiliation>
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<mods:affiliation id="D63889539CFDC911534310B804DAC407">Swedish Museum of Natural History, Department of Zoology, Stockholm, Sweden.</mods:affiliation>
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<mods:affiliation id="33851DEF6D5DCA939A945AD708572372">Ferdowsi University of Mashhad, Faculty of Science, Department of Biology, Mashhad, Iran. &amp; Research Department of Zoological Innovations (RDZI), Institute of Applied Zoology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran.</mods:affiliation>
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<mods:affiliation id="E0F5120E0C3FB5D0491473D09E15ED10">Ferdowsi University of Mashhad, Faculty of Science, Department of Biology, Mashhad, Iran</mods:affiliation>
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<mods:date id="E79B821020C5BAD02A4B565B6125B1E4">2021</mods:date>
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<treatment id="3B7687F64204624FD67FFA9DFA9AFC6D" ID-DOI="http://doi.org/10.5281/zenodo.5527109" ID-GBIF-Taxon="188518841" ID-Zenodo-Dep="5527109" LSID="urn:lsid:plazi:treatment:3B7687F64204624FD67FFA9DFA9AFC6D" httpUri="http://treatment.plazi.org/id/3B7687F64204624FD67FFA9DFA9AFC6D" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="FBC5656B4204624ED67FFA9DFB8EFA9E" box="[552,1036,1299,1326]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="B36036E04204624ED67FFA9DFB8EFA9E" blockId="38.[552,1036,1299,1326]" box="[552,1036,1299,1326]" pageId="38" pageNumber="39">
Family
<taxonomicName id="74DF4D634204624ED6D5FA9AFB8EFA9E" ID-CoL="7NK62" authority="Ljungman, 1867" authorityName="Ljungman" authorityYear="1867" box="[642,1036,1299,1326]" class="Ophiuroidea" family="Ophiolepididae" kingdom="Animalia" order="Amphilepidida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="family">
<emphasis id="81ABEAF24204624ED6D5FA9AFCC0FA9D" bold="true" box="[642,834,1299,1325]" pageId="38" pageNumber="39">Ophiolepididae</emphasis>
Ljungman, 1867
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<heading id="E828818C4204624ED4EAFADFFEC8FAC0" bold="true" box="[189,330,1366,1392]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="81ABEAF24204624ED4EAFADFFEC8FAC0" bold="true" box="[189,330,1366,1392]" pageId="38" pageNumber="39">
<typeStatus id="6C6488424204624ED4EAFADFFF78FAC0" box="[189,250,1366,1392]" pageId="38" pageNumber="39">Type</typeStatus>
genus
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<taxonomicName id="74DF4D634204624ED4EAFA0CFDEEFA2F" ID-CoL="7PND6" authority="Muller &amp; Troschel, 1840" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[189,620,1413,1439]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolepis" kingdom="Animalia" order="Amphilepidida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="genus">
<emphasis id="81ABEAF24204624ED4EAFA0CFEBCFA2F" box="[189,318,1413,1439]" italics="true" pageId="38" pageNumber="39">Ophiolepis</emphasis>
Müller &amp; Troschel, 1840
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.
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<heading id="E828818C4204624ED4EAFA4EFEE6FA51" bold="true" box="[189,356,1479,1505]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="81ABEAF24204624ED4EAFA4EFEE6FA51" bold="true" box="[189,356,1479,1505]" pageId="38" pageNumber="39">Other genera</emphasis>
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<taxonomicName id="74DF4D634204624ED4EAFA7FFDAEF9A0" ID-CoL="7PR26" authority="H. L. Clark, 1938" authorityName="H. L. Clark" authorityYear="1938" box="[189,556,1526,1553]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophioteichus" kingdom="Animalia" order="Amphilepidida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="genus">
<emphasis id="81ABEAF24204624ED4EAFA7FFEDBF9A0" box="[189,345,1526,1552]" italics="true" pageId="38" pageNumber="39">Ophioteichus</emphasis>
H.L. Clark, 1938
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;
<taxonomicName id="74DF4D634204624ED66FFA7FFCE7F9A0" ID-CoL="7PR2N" authority="Koehler, 1897" authorityName="Koehler" authorityYear="1897" box="[568,869,1526,1552]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiotypa" kingdom="Animalia" order="Amphilepidida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="genus">
<emphasis id="81ABEAF24204624ED66FFA7FFD36F9A0" box="[568,692,1526,1552]" italics="true" pageId="38" pageNumber="39">Ophiotypa</emphasis>
Koehler, 1897
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.
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<paragraph id="B36036E04204624ED4EAF9B1FECCF9E2" blockId="38.[189,334,1592,1618]" box="[189,334,1592,1618]" pageId="38" pageNumber="39">
<heading id="E828818C4204624ED4EAF9B1FECCF9E2" bold="true" box="[189,334,1592,1618]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="81ABEAF24204624ED4EAF9B1FECCF9E2" bold="true" box="[189,334,1592,1618]" pageId="38" pageNumber="39">Description</emphasis>
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<paragraph id="B36036E04204624FD4EAF9E1FD33FEDA" blockId="38.[187,1400,1640,2025]" lastBlockId="39.[188,1398,265,363]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
ARM.Arm shape unbranched; length 3× disc diameter; covering without granules/spines/tubercles. Arm combs absent. Dorsal arm plate present. Accessory dorsal arm plate present. Ventral arm plate present. Accessory ventral arm plate absent or present in other genera or species of
<taxonomicName id="74DF4D634204624ED041F926FB14F979" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[1046,1174,1711,1737]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolepis" kingdom="Animalia" order="Amphilepidida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="genus">
<emphasis id="81ABEAF24204624ED041F926FB14F979" box="[1046,1174,1711,1737]" italics="true" pageId="38" pageNumber="39">Ophiolepis</emphasis>
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(
<bibRefCitation id="D74E4B114204624ED0F1F926FAC5F979" author="Koehler R." box="[1190,1351,1711,1737]" pageId="38" pageNumber="39" refId="ref34274" refString="Koehler R. 1922. Ophiurans of the Philippine Seas and Adjacent Waters. US Government Printing Office, Washington. https: // doi. org / 10.5962 / bhl. title. 32917" type="book" year="1922">Koehler 1922</bibRefCitation>
and
<bibRefCitation id="D74E4B114204624ED4ECF95AFE44F95D" author="Pineda-Enriquez T. &amp; Solis-Marin F. A. &amp; Laguarda-Figueras A." box="[187,454,1747,1773]" pageId="38" pageNumber="39" pagination="241 - 250" refId="ref35058" refString="Pineda-Enriquez T., Solis-Marin F. A. &amp; Laguarda-Figueras A. 2014. Revision of the genus Ophioteichus H. L. Clark, 1938 (Ophiuroidea: Ophiolepididae). Zootaxa 3784 (3): 241 - 250. https: // doi. org / 10.11646 / zootaxa. 3784.3.3" type="journal article" year="2014">Pineda-Enriquez 2014</bibRefCitation>
). Girdle of tiny hooks on the arms absent. Spines all similar in shape. Arm spine cross section round. The ratio of thickness of distal ⅓ of arm spine to maximum thickness (in a given cross-sectional plane) 0.60.9. Arm spines pointed; mainly lateral; predominantly parallel to arm axis (adpressed). Longest arm spines shorter than half a segment. Tentacle scales present; pores visible along most of the arm; slightly longer than wide; shorter than the spine nearby; cross section oval. Position of lateral arm plate (LAP) arched (wrapped around the arm). Dorsal and/or ventral edge(s) of LAPs without constriction. Ventral position of LAP projecting ventro-proximalwards. Ventro-distal tip of LAP not projecting ventralwards. Arm spine articulation with dorsal and ventral lobes; without sigmoidal fold; arranged over entire distal LAP edge; larger in middle of arm. Distance between spine articulations dorsalwards increasing. Nerve and muscle openings separated by small ridge if at all. Nerve opening approximately as large as muscle opening.
</paragraph>
<paragraph id="B36036E04205624FD4EAFE1AFB40FDA7" blockId="39.[189,1399,402,536]" pageId="39" pageNumber="40">DORSAL DISC. Disc pentagonal to circular; flat (about same level as arms). Integument not obscuring plates. Dorsal disc without granules; with obvious primary plates; with scales; without spines. Dorsal disc scale/plate without tubercles. Radial shield present; without granules/spines; length between one third and half of the disc radius; isosceles (mirror-symmetric) triangular to pear-shaped.</paragraph>
<paragraph id="B36036E04205624FD4EAFDC9FEDEFC2C" blockId="39.[187,1400,575,924]" pageId="39" pageNumber="40">VENTRAL DISC. Gonads restricted to the disc. Ventral interradii without granules/spines. Scales on ventral interradii absent. Ventral teeth single. Infradental papillae on oral plate present. Buccal scales on OP present. Accessory oral papillae on OP absent. Lateral oral papillae on OP absent. Adoral shield spine present. Lymans ossicle present. Oral shield covering less than one third of interradius; much longer than wide. Madreporite one. Adoral shield meeting in front of the oral shield. Abradial genital plate not twisted. Number of genital slits one per bursa. Genital slit longer than half the length of an interradius. Genital papillae absent. Dental plate of equal width all over, or dorsal half widest; entire (single plate). Dental sockets circular to heart-shaped; equal to or more than 50% of the width. At least one socket on DP as depression or perforation without septum. Teeth block-like (with square tip). Teeth on DP arranged as single row.</paragraph>
<paragraph id="B36036E04205624FD4EAFC4AFA9AFC6D" blockId="39.[189,1304,963,990]" box="[189,1304,963,990]" pageId="39" pageNumber="40">VERTEBRAE. Vertebrae zygospondylous; beyond segment 5 with short keel; without oral bridge.</paragraph>
</subSubSection>
</treatment>
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