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<document id="70D452C337F321FA8B1E4E862FB5105B" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C589B0FFF67F0789CF0F9F4" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus raziogus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="44" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="44" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="C2D39D316022F09A88F968C8BA4FEC8C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2EC9376EFC25FA8FFA0E197FED1AB65C">
<mods:title id="F3FECB462515FA5C55D81946D24BD9B1">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
<mods:name id="3D332FF8CA7B304A49E9505B4E1B4B3D" type="personal">
<mods:role id="7C8AE1574720597F2EEB6D95A347E195">
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<mods:namePart id="211AB28773ECF8DA8784981DEC868D03">Azevedo, Celso O.</mods:namePart>
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<mods:title id="4828157F171D17E8983FB92B5E0262D6">Zootaxa</mods:title>
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<mods:part id="C398B408657CA76797C0AB639FF4F281">
<mods:date id="08BE6AB848ED9EB46909EDF7332E73F1">2017</mods:date>
<mods:detail id="684D65D29DE76EBE09FF13AAF2D65FE7" type="pubDate">
<mods:number id="1FB6D2C4D4E63FBD88A83EC0B42EC94D">2017-10-13</mods:number>
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<mods:number id="53135A63B48634BAA8785D13DDD294BA">4335</mods:number>
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<mods:number id="A900EFC9593B75234B1DCD718A5B5A99">1</mods:number>
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<mods:classification id="03DF42EB980EFB8DA4CAECCB8E1EC6B5">journal article</mods:classification>
<mods:identifier id="50B16EA7006D0199207580E987544904" type="CLB-Dataset">31852</mods:identifier>
<mods:identifier id="9396766A32663FECEF8A9025C80EAC80" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<mods:identifier id="23C1EA77788C8C34FFBFA4F55CB6221B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3B3687C07C589B0FFF67F0789CF0F9F4" ID-DOI="http://doi.org/10.5281/zenodo.5020431" ID-GBIF-Taxon="135435298" ID-Zenodo-Dep="5020431" LSID="urn:lsid:plazi:treatment:3B3687C07C589B0FFF67F0789CF0F9F4" httpUri="http://treatment.plazi.org/id/3B3687C07C589B0FFF67F0789CF0F9F4" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="FB85655D7C589B0FFF67F0789EA4FD5C" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="B32036D67C589B0FFF67F0789D8BFDBD" blockId="43.[151,750,583,643]" box="[151,750,583,610]" pageId="43" pageNumber="44">
<heading id="E86881BA7C589B0FFF67F0789D8BFDBD" bold="true" box="[151,750,583,610]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="81EBEAC47C589B0FFF67F0789D8BFDBD" bold="true" box="[151,750,583,610]" pageId="43" pageNumber="44">
<taxonomicName id="749F4D557C589B0FFF67F0789DECFDBD" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,649,583,610]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="raziogus" status="sp. nov.">
<emphasis id="81EBEAC47C589B0FFF67F0789ECAFDBE" bold="true" box="[151,431,583,609]" italics="true" pageId="43" pageNumber="44">Dissomphalus raziogus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C589B0FFD60F0779D8BFDBD" box="[656,750,584,610]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C589B0FFF67F0559EA4FD5C" blockId="43.[151,750,583,643]" box="[151,449,618,643]" pageId="43" pageNumber="44">
(
<figureCitation id="2BA42A537C589B0FFF6FF0559F90FD5C" box="[159,245,618,643]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="43" pageNumber="44">Figs 18</figureCitation>
,
<figureCitation id="2BA42A537C589B0FFEF1F0549E45FD5C" box="[257,288,619,643]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="43" pageNumber="44">49</figureCitation>
,
<figureCitation id="2BA42A537C589B0FFEDBF0549E2FFD5C" box="[299,330,619,643]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="43" pageNumber="44">80</figureCitation>
,
<figureCitation id="2BA42A537C589B0FFEA5F0559EDCFD5C" box="[341,441,618,643]" captionStart="FIGURES 163 – 173" captionStartId="41.[151,264,1603,1626]" captionTargetBox="[171,1417,205,1574]" captionTargetId="figure@41.[151,1436,193,1582]" captionTargetPageId="41" captionText="FIGURES 163 – 173. Dissomphalus spp., ♂, genitalia, except fig. 173 Outer lobe of aedeagal dorsal body. 163 – 164. D. curvifoveatus. 163. Ventral view. 162. Apex of aedeagal ventral ramus. 165 – 166. D. arbeius sp. nov. 165. Dorsal view. 166. Ventral view. 167 – 168. D. guttus 167. Dorsal view. 168. Ventral view. 169 – 170. D. raziogus sp. nov. 169. Dorsal view. 170. Ventral view. 171 – 173. D. bilobatus 171. Dorsal view. 172. Ventral view. 173. Apex ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010044/files/figure.png" pageId="43" pageNumber="44">169–170</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C589B0FFF67F08D9CDCFBEC" pageId="43" pageNumber="44" type="description">
<paragraph id="B32036D67C589B0FFF67F08D9BECFBD0" blockId="43.[151,1437,690,1579]" pageId="43" pageNumber="44">
<emphasis id="81EBEAC47C589B0FFF67F08D9E4CFD14" bold="true" box="[151,297,690,715]" pageId="43" pageNumber="44">Description.</emphasis>
Male. Body length
<quantity id="74679B337C589B0FFDE6F08C9DE4FD15" box="[534,641,691,715]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="43" pageNumber="44" unit="mm" value="2.27">2.27 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (
<figureCitation id="2BA42A537C589B0FFF12F0E99E52FD30" box="[226,311,726,751]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="43" pageNumber="44">Fig. 18</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest strongly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C589B0FFA9FF1219FD2FC84" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="43" pageNumber="44">Fig. 49</figureCitation>
). Tergal process absent. Posterior hypopygeal margin weakly concave (
<figureCitation id="2BA42A537C589B0FFBF0F17D9B3CFC84" box="[1024,1113,834,859]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="43" pageNumber="44">Fig. 80</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C589B0FFB1CF17D9AF5FC84" box="[1260,1424,834,859]" captionStart="FIGURES 163 – 173" captionStartId="41.[151,264,1603,1626]" captionTargetBox="[171,1417,205,1574]" captionTargetId="figure@41.[151,1436,193,1582]" captionTargetPageId="41" captionText="FIGURES 163 – 173. Dissomphalus spp., ♂, genitalia, except fig. 173 Outer lobe of aedeagal dorsal body. 163 – 164. D. curvifoveatus. 163. Ventral view. 162. Apex of aedeagal ventral ramus. 165 – 166. D. arbeius sp. nov. 165. Dorsal view. 166. Ventral view. 167 – 168. D. guttus 167. Dorsal view. 168. Ventral view. 169 – 170. D. raziogus sp. nov. 169. Dorsal view. 170. Ventral view. 171 – 173. D. bilobatus 171. Dorsal view. 172. Ventral view. 173. Apex ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010044/files/figure.png" pageId="43" pageNumber="44">Figs 169–170</figureCitation>
): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin excavated; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and narrow, with apical margin acute and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C589B0FFF37F6259CDCFBEC" blockId="43.[151,1437,690,1579]" box="[199,953,1050,1075]" pageId="43" pageNumber="44">
<emphasis id="81EBEAC47C589B0FFF37F6259E22FBEC" bold="true" box="[199,327,1050,1075]" pageId="43" pageNumber="44">Variations.</emphasis>
Mesosoma dark castaneous; median clypeal lobe short.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C589B0FFF37F6019A7FFAD4" pageId="43" pageNumber="44" type="discussion">
<paragraph id="B32036D67C589B0FFF37F6019A7FFAD4" blockId="43.[151,1437,690,1579]" pageId="43" pageNumber="44">
<emphasis id="81EBEAC47C589B0FFF37F6019E5EFB88" bold="true" box="[199,315,1086,1111]" pageId="43" pageNumber="44">Remarks.</emphasis>
This species is similar to
<taxonomicName id="749F4D557C589B0FFD90F6009DAAFB89" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[608,719,1087,1110]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="isortus" status="sp. nov.">
<emphasis id="81EBEAC47C589B0FFD90F6009DAAFB89" box="[608,719,1087,1110]" italics="true" pageId="43" pageNumber="44">D. isortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C589B0FFD26F6019C54FB88" box="[726,817,1086,1111]" pageId="43" pageNumber="44" rank="species">
<emphasis id="81EBEAC47C589B0FFD26F6019C54FB88" bold="true" box="[726,817,1086,1111]" pageId="43" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
by having the aedeagal dorsal body narrowing slightly apicad, the apical lobes of the aedeagal dorsal body in conical-shaped. However, this species has the tergal process absent, the paramere narrow, the apex of the apical lobes of the aedeagal dorsal body mesad and the aedeagal ventral ramus with filament inserted in the median part, characteristic diagnosis for
<emphasis id="81EBEAC47C589B0FFB9EF6939B8BFB1C" box="[1134,1262,1196,1219]" italics="true" pageId="43" pageNumber="44">microsticus</emphasis>
species-group, whereas
<taxonomicName id="749F4D557C589B0FFF0AF6F09E0CFB39" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[250,361,1231,1254]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="isortus" status="sp. nov.">
<emphasis id="81EBEAC47C589B0FFF0AF6F09E0CFB39" box="[250,361,1231,1254]" italics="true" pageId="43" pageNumber="44">D. isortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C589B0FFE81F6F19EAEFB38" box="[369,459,1230,1255]" pageId="43" pageNumber="44" rank="species">
<emphasis id="81EBEAC47C589B0FFE81F6F19EAEFB38" bold="true" box="[369,459,1230,1255]" pageId="43" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
has the tergal process with a tubercle, pit with thick setae the paramere wide, the apex of the apical lobes of the aedeagal dorsal body dorsad and the aedeagal ventral ramus without filament.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C589B0FFF37F7299DF9F9D8" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="B32036D67C589B0FFF37F7299DF9F9D8" blockId="43.[151,1437,690,1579]" pageId="43" pageNumber="44">
<materialsCitation id="03F73C8B7C589B0FFF37F7299AFCFA8C" ID-GBIF-Occurrence="1671715861" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr" country="Panama" county="Material" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C589B0FFF37F7299ED6FAF0" bold="true" box="[199,435,1302,1327]" pageId="43" pageNumber="44">
<collectingCounty id="5A414E5A7C589B0FFF37F7299E4AFAF0" box="[199,303,1302,1327]" pageId="43" pageNumber="44">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="6C2488747C589B0FFE4CF7299D42FAF0" box="[444,551,1302,1327]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C589B0FFDC5F7289D29FAF1" box="[565,588,1303,1326]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
,
<collectingRegion id="715BF8347C589B0FFDAAF7289DB4FAF0" box="[602,721,1303,1327]" country="Panama" name="Panama" pageId="43" pageNumber="44">
<collectingCountry id="CB8876467C589B0FFDAAF7289DB4FAF0" box="[602,721,1303,1327]" name="Panama" pageId="43" pageNumber="44">PANAMA</collectingCountry>
</collectingRegion>
,
<collectingRegion id="715BF8347C589B0FFD11F7289CEBFAF1" box="[737,910,1302,1327]" country="Panama" name="Bocas del Toro" pageId="43" pageNumber="44">
<emphasis id="81EBEAC47C589B0FFD11F7289CEBFAF1" box="[737,910,1302,1327]" italics="true" pageId="43" pageNumber="44">Bocas del Toro</emphasis>
</collectingRegion>
:
<collectingMunicipality id="5344ACAC7C589B0FFC6FF7299C8BFAF0" box="[927,1006,1302,1327]" pageId="43" pageNumber="44">Wekso</collectingMunicipality>
,
<location id="B640600D7C589B0FFC0BF7299B20FAF0" LSID="urn:lsid:plazi:treatment:3B3687C07C589B0FFF67F0789CF0F9F4:B640600D7C589B0FFC0BF7299B20FAF0" box="[1019,1093,1302,1327]" country="Panama" county="Material" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Teribe" pageId="43" pageNumber="44" stateProvince="PANAMA">Teribe</location>
, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C589B0FFE57F7059D48FA8C" box="[423,557,1338,1363]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C589B0FFDB0F7059DA8FA8C" box="[576,717,1338,1363]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C589B0FFD16F7049CCEFA8C" box="[742,939,1339,1363]" pageId="43" pageNumber="44" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C589B0FFD16F7049CCEFA8C" box="[742,939,1339,1363]" pageId="43" pageNumber="44" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C589B0FFC4DF7049CBFFA8D" box="[957,986,1339,1362]" pageId="43" pageNumber="44">Tr</collectorName>
[ap] Malaise, A. Santos[col.] (
<collectionCode id="D58EAE137C589B0FFAB4F7049AF5FA8C" box="[1348,1424,1339,1363]" pageId="43" pageNumber="44">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C589B0FFF67F7609D0DFAA8" ID-GBIF-Occurrence="1671715863" box="[151,616,1374,1399]" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr" country="Panama" county="Material" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="43" pageNumber="44" specimenCount="8" specimenCount-male="8" stateProvince="PANAMA" typeStatus="paratype">
<typeStatus id="6C2488747C589B0FFF67F7609E6CFAA8" box="[151,265,1375,1399]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A599FD5F7C589B0FFEEBF7609E25FAA8" box="[283,320,1375,1399]" pageId="43" pageNumber="44" type="male">8♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="03F73C8B7C589B0FFD8BF7619A03FA44" ID-GBIF-Occurrence="1671715864" collectingDate="1930-10-05" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Rodriguez" country="Panama" elevation="1800" latitude="8.9" location="Jurutungo" longLatPrecision="1300" longitude="-82.73333" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui Pr" typeStatus="paratype">
<emphasis id="81EBEAC47C589B0FFD8BF7619C0BFAA9" box="[635,878,1374,1399]" italics="true" pageId="43" pageNumber="44">
<collectingRegion id="715BF8347C589B0FFD8BF7619C60FAA9" box="[635,773,1374,1399]" isEnumeration="true" pageId="43" pageNumber="44">
<collectingRegion id="715BF8347C589B0FFD8BF7619DBFFAA8" box="[635,730,1374,1399]" country="Panama" name="Chiriqui" pageId="43" pageNumber="44">Chiriquí</collectingRegion>
<collectingRegion id="715BF8347C589B0FFD19F7609C60FAA9" box="[745,773,1375,1398]" country="Brazil" name="Para" pageId="43" pageNumber="44">Pr</collectingRegion>
</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C589B0FFC74F7609CCCFAA8" box="[900,937,1375,1399]" pageId="43" pageNumber="44" type="male">1♂</specimenCount>
, P[arque] I[nternacional] L[a] A[mistad],
<location id="B640600D7C589B0FFF67F7BC9E6FFA44" LSID="urn:lsid:plazi:treatment:3B3687C07C589B0FFF67F0789CF0F9F4:B640600D7C589B0FFF67F7BC9E6FFA44" box="[151,266,1411,1435]" country="Panama" latitude="8.9" longLatPrecision="1300" longitude="-82.73333" name="Jurutungo" pageId="43" pageNumber="44" stateProvince="Chiriqui Pr">Jurutungo</location>
,
<elevation id="38B2D1E57C589B0FFEE5F7BC9E08FA45" box="[277,365,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="43" pageNumber="44" unit="m" value="1800.0">
<quantity id="74679B337C589B0FFEE5F7BC9E08FA45" box="[277,365,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="43" pageNumber="44" unit="m" value="1800.0">1800 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C589B0FFE70F7BD9E84FA44" box="[384,481,1410,1435]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="8.9">08°54'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C589B0FFE1EF7BC9D30FA44" box="[494,597,1411,1435]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="-82.73333">82°44'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C589B0FFD98F7BC9DAEFA44" box="[616,715,1410,1435]" pageId="43" pageNumber="44" value="1930-10-05">
<date id="C72110167C589B0FFD98F7BC9DAEFA44" box="[616,715,1410,1435]" pageId="43" pageNumber="44" value="1930-10-05">30 oct. 5</date>
</collectingDate>
nov. 1994,
<collectorName id="1E6A53007C589B0FFCA0F7BC9C09FA45" box="[848,876,1411,1434]" pageId="43" pageNumber="44">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C589B0FFC30F7BD9B5AFA44" box="[960,1087,1410,1435]" pageId="43" pageNumber="44">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C589B0FFBBAF7BD9BDBFA44" box="[1098,1214,1410,1435]" pageId="43" pageNumber="44">Rodriguez</collectorName>
[col.] (
<collectionCode id="D58EAE137C589B0FFAE2F7BC9A3BFA44" box="[1298,1374,1411,1435]" pageId="43" pageNumber="44">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C589B0FFA82F7BC9A00FA60" ID-GBIF-Occurrence="1671715874" collectingDate="1997-05-24" collectingDateMax="1997-05-31" collectingDateMin="1997-05-24" collectionCode="CNCI" country="Panama" elevation="1200" latitude="8.766666" location="Volcano" longLatPrecision="1300" longitude="-82.63333" municipality="Hato" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui Pr" typeStatus="paratype">
<specimenCount id="A599FD5F7C589B0FFA82F7BC9AF3FA44" box="[1394,1430,1411,1435]" pageId="43" pageNumber="44" type="male">1♂</specimenCount>
,
<quantity id="74679B337C589B0FFF67F7999F85FA60" box="[151,224,1446,1471]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="km" value="15.0">15 km</quantity>
NW
<collectingMunicipality id="5344ACAC7C589B0FFED1F7989E32FA61" box="[289,343,1447,1470]" pageId="43" pageNumber="44">Hato</collectingMunicipality>
del
<location id="B640600D7C589B0FFE7DF7999E8EFA60" LSID="urn:lsid:plazi:treatment:3B3687C07C589B0FFF67F0789CF0F9F4:B640600D7C589B0FFE7DF7999E8EFA60" box="[397,491,1446,1471]" country="Panama" latitude="8.766666" longLatPrecision="1300" longitude="-82.63333" municipality="Hato" name="Volcano" pageId="43" pageNumber="44" stateProvince="Chiriqui Pr">Volcano</location>
,
<elevation id="38B2D1E57C589B0FFE08F7989D37FA61" box="[504,594,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="44" unit="m" value="1200.0">
<quantity id="74679B337C589B0FFE08F7989D37FA61" box="[504,594,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="44" unit="m" value="1200.0">1200 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C589B0FFD97F7999DADFA60" box="[615,712,1446,1471]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="8.766666">08°46'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C589B0FFD26F7989C5BFA60" box="[726,830,1447,1471]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="-82.63333">82°38'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C589B0FFCA2F7989C94FA60" box="[850,1009,1447,1471]" pageId="43" pageNumber="44" value="1997-05-24" valueMax="1997-05-31" valueMin="1997-05-24">
<date id="C72110167C589B0FFCA2F7989C94FA60" box="[850,1009,1447,1471]" pageId="43" pageNumber="44" value="1997-05-24" valueMax="1997-05-31" valueMin="1997-05-24">24–31.V.1997</date>
</collectingDate>
, Peck &amp; Howden [col.] (
<collectionCode id="D58EAE137C589B0FFAE4F7989A38FA60" box="[1300,1373,1447,1471]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="43" pageNumber="44">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C589B0FFA82F7989DFDF9D8" ID-GBIF-Occurrence="1671715849" collectingDate="1999-04-08" collectingDateMax="1999-04-11" collectingDateMin="1999-04-08" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, A &amp; Santos" country="Panama" elevation="1080" latitude="8.77125" location="Toro" longLatPrecision="1" longitude="-82.19092" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui Pr" typeStatus="paratype">
<specimenCount id="A599FD5F7C589B0FFA82F7989AF2FA60" box="[1394,1431,1447,1471]" pageId="43" pageNumber="44" type="male">1♂</specimenCount>
,
<taxonomicName id="749F4D557C589B0FFF67F7F49E1BFA3C" authority=": Div. Cont." authorityName="Div. Cont." box="[151,382,1482,1507]" class="Gastropoda" family="Achatinidae" genus="Fortuna" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="genus">Fortuna: Div. Cont.</taxonomicName>
Chiriqui-B[ocas del]
<location id="B640600D7C589B0FFD75F7F49DD9FA3D" LSID="urn:lsid:plazi:treatment:3B3687C07C589B0FFF67F0789CF0F9F4:B640600D7C589B0FFD75F7F49DD9FA3D" box="[645,700,1483,1506]" country="Panama" latitude="8.77125" longLatPrecision="1" longitude="-82.19092" name="Toro" pageId="43" pageNumber="44" stateProvince="Chiriqui Pr">Toro</location>
,
<elevation id="38B2D1E57C589B0FFD3EF7F49C14FA3D" box="[718,881,1483,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.09" metricValueMin="1.0699999999999998" pageId="43" pageNumber="44" unit="m" value="1080.0" valueMax="1090.0" valueMin="1070.0">
<quantity id="74679B337C589B0FFD3EF7F49C14FA3D" box="[718,881,1483,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.09" metricValueMin="1.0699999999999998" pageId="43" pageNumber="44" unit="m" value="1080.0" valueMax="1090.0" valueMin="1070.0">1070–1090 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C589B0FFC7BF7F59B4DFA3C" box="[907,1064,1482,1507]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="8.77125">08°46'16.5&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C589B0FFBCAF7F49BB8FA3C" box="[1082,1245,1483,1507]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="-82.19092">82°11'27.3&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C589B0FFB06F7F59AFDFA3C" box="[1270,1432,1482,1507]" pageId="43" pageNumber="44" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">
<date id="C72110167C589B0FFB06F7F59AFDFA3C" box="[1270,1432,1482,1507]" pageId="43" pageNumber="44" value="1999-04-08" valueMax="1999-04-11" valueMin="1999-04-08">8–11.abr.1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C589B0FFF67F7D19E76F9D9" box="[151,275,1518,1543]" pageId="43" pageNumber="44">Malaise, R</collectorName>
.
<collectorName id="1E6A53007C589B0FFEEEF7D19EF9F9D8" box="[286,412,1518,1543]" pageId="43" pageNumber="44">Cambra, A</collectorName>
.
<collectorName id="1E6A53007C589B0FFE57F7D09E97F9D8" box="[423,498,1519,1543]" pageId="43" pageNumber="44">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C589B0FFDB4F7D09DF5F9D8" box="[580,656,1519,1543]" pageId="43" pageNumber="44">MIUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C589B0FFF37F42D9CF0F9F4" box="[199,917,1554,1579]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="B32036D67C589B0FFF37F42D9CF0F9F4" blockId="43.[151,1437,690,1579]" box="[199,917,1554,1579]" pageId="43" pageNumber="44">
<emphasis id="81EBEAC47C589B0FFF37F42D9E3CF9F4" bold="true" box="[199,345,1554,1579]" pageId="43" pageNumber="44">Distribution</emphasis>
(
<figureCitation id="2BA42A537C589B0FFE98F42D9EAEF9F4" box="[360,459,1554,1579]" captionStart="FIGURES 234 – 243" captionStartId="67.[151,264,1856,1879]" captionTargetBox="[174,1439,203,1822]" captionTargetId="figure@67.[162,1447,193,1827]" captionTargetPageId="67" captionText="FIGURES 234 – 243. Known distribution of Dissomphalus in Panama. 234. D. galeatus, D. geniculatus and D. geortus sp. nov.. 235. D. gilvipes, D. ibocius sp. nov. and D. ibrastus sp. nov.. 236. D. guttus and D. ilarius sp. nov.. 237. D. infissus, D. intradentatus and D. iozus sp. nov.. 238. D. isortus sp. nov., D. isteus sp. nov. and D. latus. 239. D. persontaus, D. pilus and D. pronus. 240. D. plaumanni and D. politus. 241. D. piscicercus, D. ramosus and D. raziogus sp. nov.. 242. D. punctatus, D. strepsus and D. subdeformis. 243. D. racarus sp. nov., D. strabus and D. tisinus sp. nov.." httpUri="https://zenodo.org/record/1010060/files/figure.png" pageId="43" pageNumber="44">Fig. 241</figureCitation>
).
<collectingRegion id="715BF8347C589B0FFE2EF42C9D52F9F5" box="[478,567,1555,1578]" country="Panama" name="Panama" pageId="43" pageNumber="44">
<collectingCountry id="CB8876467C589B0FFE2EF42C9D52F9F5" box="[478,567,1555,1578]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C589B0FFDB6F42C9D95F9F5" box="[582,752,1554,1579]" country="Panama" name="Bocas del Toro" pageId="43" pageNumber="44">Bocas del Toro</collectingRegion>
and
<collectingRegion id="715BF8347C589B0FFCD6F42D9CECF9F4" box="[806,905,1554,1579]" country="Panama" name="Chiriqui" pageId="43" pageNumber="44">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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