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<document id="16EC0A08CB851A1B390300D53C638F16" ENCODING="UTF-8" ID-GBIF-Dataset="b8f58d00-19e5-4cfa-a1f0-4c686286fc5e" ModsDocID="z01603p001" checkinTime="1247319475874" checkinUser="thomas" docAuthor="Juan J. Schmitter-Soto" docDate="2007" docId="131484AFA72F82190797063675C7E124" docLanguage="en" docName="2007_Schmitter-Soto_gg1.xml" docOrigin="Zootaxa 1603" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:AFFCB590-1FC7-4CD0-950C-D1D1A6E59F6C" docTitle="Rocio octofasciata Schmitter-Soto, 2007, n. comb." docType="treatment" docVersion="12" lastPageNumber="59" masterDocId="9F5EADD85808D5BD56FDE2FB757992C0" masterDocTitle="A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species." masterLastPageNumber="78" masterPageNumber="1" pageNumber="57" updateTime="1696279408464" updateUser="admin">
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<mods:title id="FC7F8E604A49BA5EAF7EC7F70E067A41">A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species.</mods:title>
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<mods:namePart id="5A5C90E8EADD576D0E277B10C5616FA8">Juan J. Schmitter-Soto</mods:namePart>
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<treatment id="131484AFA72F82190797063675C7E124" ID-DOI="http://doi.org/10.5281/zenodo.6248797" ID-GBIF-Taxon="100117503" ID-Zenodo-Dep="6248797" LSID="urn:lsid:plazi:treatment:131484AFA72F82190797063675C7E124" httpUri="http://treatment.plazi.org/id/131484AFA72F82190797063675C7E124" lastPageNumber="59" pageNumber="57">
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<paragraph id="7E0609CEC634E2594F7EA26559866969" pageNumber="57">
<taxonomicName id="7A58100B2274CF27CD32B33540DE4021" ID-CoL="4T8T2" ID-ENA="661982" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata" status="n. comb.">
Rocio octofasciata (
<bibRefCitation id="3BCA3066D88E6B06F5AACA6A6370AC6D" author="Regan, C. T." pageNumber="3" pagination="413 - 418" refId="ref36927" refString="Regan, C. T. (1903) Descriptions de poissons nouveaux faisant partie de la collection du Musee d'Histoire Naturelle de Geneve. Revue Suisse de Zoologie, 11, 413 - 418." type="journal article" year="1903">Regan, 1903</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="407B4532E5C12AF0AB55974758F20C1B" pageNumber="3" rank="species">n. comb.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="543BA294D4EF8FB998CF1E1A9EA6F9D6" pageNumber="3" type="description">
<paragraph id="47456171D25A2BDD4B40D4219A146A79" pageNumber="57">Figures 22-24</paragraph>
</subSubSection>
<subSubSection id="FF258A98E5523CA2CA38FB2F44CC59DF" pageNumber="3" type="reference_group">
<paragraph id="B2F7E6D44088A2E834A1AA29599E8D2B" pageNumber="57">
<taxonomicName id="F5C92D38EC613EEBF561911D52C6D6F3" family="Cichlidae" genus="Heros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cyanoguttatus">Heros cyanoguttatus</taxonomicName>
(part. et non Baird &amp; Girard),
<bibRefCitation id="C90149F50082E185873716D8C1653C5E" author="Evermann, B. W. &amp; Goldsborough, E. L." pageNumber="3" pagination="137 - 159" refId="ref35060" refString="Evermann, B. W. &amp; Goldsborough, E. L. (1902) A report on fishes collected in Mexico and Central America, with notes and descriptions of five new species. Bulletin of the United States Fish Commission, 1901, 137 - 159." type="journal article" year="1902">Evermann &amp; Goldsborough 1902: 157</bibRefCitation>
(misidentification).
</paragraph>
<paragraph id="62B58C30FA4600199C251E9E9B96FE04" pageNumber="57">
<taxonomicName id="BFABEA7365E7D5FCCF6E0383BA9313AF" LSID-ZBK="urn:lsid:zoobank.org:act:D76352FE-1F2C-4AFD-9081-678A86D6C797" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros octofasciatus Regan 1903:417" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatus">
Heros octofasciatus
<bibRefCitation id="C40EC35B24236DF24B910FFE51E4B972" author="Regan, C. T." pageNumber="3" pagination="413 - 418" refId="ref36927" refString="Regan, C. T. (1903) Descriptions de poissons nouveaux faisant partie de la collection du Musee d'Histoire Naturelle de Geneve. Revue Suisse de Zoologie, 11, 413 - 418." type="journal article" year="1903">Regan, 1903: 417</bibRefCitation>
</taxonomicName>
(original description).
</paragraph>
<paragraph id="ABF67CC2096AEF350CC751E878FF46B9" pageNumber="57">
<taxonomicName id="2F6B8D7A499DF3942C37B4944CE27FCF" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatum">Cichlasoma octofasciatum</taxonomicName>
,
<bibRefCitation id="2D7AFB740B9F33697100D9323B53488B" author="Meek, S. E." pageNumber="3" pagination="1 - 252" refId="ref36415" refString="Meek, S. E. (1904) The fresh-water fishes of Mexico north of the Isthmus of Tehuantepec. Field Columbian Museum of Natural History Publications, Zoological Series, 93, 1 - 252." type="journal article" year="1904">Meek 1904: 218</bibRefCitation>
(
<taxonomicNameLabel id="BF5693045F0D2638DBE17BA98A81549B" pageNumber="3" rank="species">new combination</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8E11F20B61199CC988D87BAD22421E67" pageNumber="57">
<taxonomicName id="3A5B089077F8F8CA6A30FBE0F0174152" LSID-ZBK="urn:lsid:zoobank.org:act:3A693BD7-FD95-4532-9F14-41B963B40D09" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma hedricki Meek 1904:208" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="hedricki">
Cichlasoma hedricki
<bibRefCitation id="E129A2B756AF148D4E2AD6CAAE336FB2" author="Meek, S. E." pageNumber="3" pagination="1 - 252" refId="ref36415" refString="Meek, S. E. (1904) The fresh-water fishes of Mexico north of the Isthmus of Tehuantepec. Field Columbian Museum of Natural History Publications, Zoological Series, 93, 1 - 252." type="journal article" year="1904">Meek, 1904: 208</bibRefCitation>
</taxonomicName>
(junior synonym).
</paragraph>
<paragraph id="33555ECDDAB24FF9A33630F2E146F120" pageNumber="57">
<taxonomicName id="66411AF07E0C30C2B1E400FF75B80095" LSID-ZBK="urn:lsid:zoobank.org:act:E7A510F9-CF8C-4181-A6CE-DD7BA7F860F2" family="Cichlidae" genus="Cichlosoma" kingdom="Animalia" lsidName-ZBK="Cichlosoma biocellatum Regan 1909:234" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="biocellatum">
Cichlosoma biocellatum
<bibRefCitation id="2E8084256692D0EFB08EEB18B1014DDF" author="Regan, C. T." pageNumber="3" pagination="234 - 235" refId="ref37085" refString="Regan, C. T. (1909) Descriptions of three new freshwater fishes from South America, presented to the British Museum by Herr J. Paul Arnold. Annual Magazine of Natural History, 3, 234 - 235." type="journal article" year="1909">Regan, 1909: 234</bibRefCitation>
</taxonomicName>
(junior synonym).
</paragraph>
<paragraph id="0AE914A7996CEFCC9E8A0DE961B1886F" pageNumber="57">
<taxonomicName id="A3C3257F66ADAB96A1A54302550072E7" family="Cichlidae" genus="Parapetenia" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Parapetenia octofasciata</taxonomicName>
,
<bibRefCitation id="259221342704B0C3B769941B3F35123F" author="Allgayer, R." pageNumber="3" pagination="4 - 30" refId="ref34450" refString="Allgayer, R. (1989) Revision et redescription du genre Theraps Guenther, 1862. Description de deux especes nouvelles du Mexique (Pisces, Perciformes, Cichlidae). Revue Francaise des Cichlidophiles, 90 bis, 4 - 30." type="journal article" year="1989">Allgayer 1989: 26</bibRefCitation>
(
<taxonomicNameLabel id="86C3931BC1E7B32934EF630E2CA138D2" pageNumber="3" rank="species">new combination</taxonomicNameLabel>
).
</paragraph>
<paragraph id="83E5B03F8E2D3CDDC4A0538F13D75D4E" pageNumber="57">
<taxonomicName id="7E6911A5043F9A3375E7227459B77508" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatum">Cichlasoma octofasciatum</taxonomicName>
,
<bibRefCitation id="B5C8427C2E6ED8137B9C2D3A449DD6E0" author="Kullander, S. O." pageNumber="3" pagination="149 - 172" refId="ref35926" refString="Kullander, S. O. (1996) Heroina isonycterina, a new genus and species of cichlid fish from Western Amazonia, with comments on cichlasomine systematics. Ichthyological Exploration of Freshwaters, 7, 149 - 172." type="journal article" year="1996">Kullander 1996: 151</bibRefCitation>
(incertae sedis).
</paragraph>
<paragraph id="0DC1F3B5503155C5B790030D0023C610" pageNumber="57">
<taxonomicName id="7B38AE0A2301E84291E6BF5C4944CD49" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatus">Archocentrus octofasciatus</taxonomicName>
,
<bibRefCitation id="24935C0BC1B47A0C934DE9AF16B2D5F1" author="Schmitter-Soto, J. J." pageNumber="3" refId="ref37257" refString="Schmitter-Soto, J. J. (1998) Catalogo de los peces continentales de Quintana Roo. ECOSUR, San Cristobal de Las Casas, 239 pp." type="book" year="1998">Schmitter-Soto 1998: 154</bibRefCitation>
(
<taxonomicNameLabel id="B14A51C51D24F18DEAC65ABBC86A3668" pageNumber="3" rank="species">new combination</taxonomicNameLabel>
).
</paragraph>
<paragraph id="F1B0CFBF370B22DC32E0E2CEBB6F1385" pageNumber="57">
<taxonomicName id="671E43868C9647003137ABD1B1806146" family="Cichlidae" genus="Nandopsis" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Nandopsis octofasciata</taxonomicName>
,
<bibRefCitation id="D51DC9B7A467EC2AD6465571EDB8EBFC" author="Burgess, W. E." pageNumber="3" pagination="44 - 54" refId="ref34598" refString="Burgess, W. E. (2000) The Cichlasoma story. Herichthys, the break-up. Tropical Fish Hobbyist, 48, 44 - 54." type="journal article" year="2000">Burgess2000: 48</bibRefCitation>
(
<taxonomicNameLabel id="E7BD4037307E15B0E1CC7535C464249A" pageNumber="3" rank="species">new combination</taxonomicNameLabel>
[as
<taxonomicName id="7D4C3C22ECB1BEF09C833393E931AF07" family="Cichlidae" genus="Nandopsis" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatum">octofasciatum</taxonomicName>
]).
</paragraph>
</subSubSection>
<subSubSection id="8803784714A55B8FF9367CF61D78D691" pageNumber="3" type="nomenclature">
<paragraph id="911E00ABCE9AAFAF2602D47175E82E2D" pageNumber="57">
<materialsCitation id="5974908A838BDFBBEF3FE56F29FDD054" ID-GBIF-Occurrence="923904143" collectingDate="1866-02-01" collectionCode="MHNG" collectorName="A. C. Buller" country="Mexico" location="“ Rio de Sarabia ” (Cosamaloapan, according to the original label), Coatzacoalcos drainage" pageNumber="3" specimenCode="MHNG 665.55" stateProvince="Veracruz" typeStatus="holotype">
<typeStatus id="156A9700E252A94F5B3263D71911797C" pageNumber="3" type="holotype">Holotype</typeStatus>
.
<specimenCode id="D8DA426E82473CB72085CCF7472BA67D" pageNumber="3">
<collectionCode id="FE2F004644FFB17E78ABF0B4249F0C65" pageNumber="3">MHNG</collectionCode>
665.55
</specimenCode>
, 39 mm SL (Fig. 23),
<collectorName id="7A53F6DA7ACDED0B7639900FEA078225" pageNumber="3">A. C. Buller</collectorName>
,
<date id="D71745A0B4DBB4025E4480A423321364" day="01" month="02" pageNumber="3" value="1866-02-01" year="1866">
<collectingDate id="DE199658BA1261068156B52951C311E9" day="01" month="02" pageNumber="3" value="1866-02-01" year="1866">Feb. 1, 1866</collectingDate>
</date>
.
<location id="6457DBD0D0902F2F746DF0CA38FE7384" LSID="urn:lsid:plazi:treatment:131484AFA72F82190797063675C7E124:6457DBD0D0902F2F746DF0CA38FE7384" country="Mexico" name="“ Rio de Sarabia ” (Cosamaloapan, according to the original label), Coatzacoalcos drainage" pageNumber="3" stateProvince="Veracruz">
<normalizedToken id="2225A27978DBEC6AA4C25210535A8BD3" originalValue="Río" pageNumber="3">Rio</normalizedToken>
de Sarabia” (Cosamaloapan, according to the original label), Coatzacoalcos drainage
</location>
,
<collectingRegion id="08EBC1E0EDB60D50D09E4C7F687B5A21" pageNumber="3">Veracruz</collectingRegion>
,
<collectingCountry id="7DD1310B24E2451A12149C812AB4AF50" pageNumber="3">Mexico</collectingCountry>
. No paratypes, but originally mixed, unlabeled, in same jar with eight more specimens (see Remarks).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="35E2F4FB0FE15D87A4C8641B435E6919" pageNumber="3" type="diagnosis">
<paragraph id="1810D26ABA7C30446DBBA6AE5135EB6A" lastPageNumber="58" pageNumber="57">
Diagnosis. No unique autapomorphies. Spots on sides smaller than scales, aligned in ca. 15 regular series (vs. not clearly aligned); abdomen predominantly whitish or greyish in life (also in
<taxonomicName id="C8B8DD89E0E6C03690B60EA1CC40A0CA" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="gemmata">R. gemmata</taxonomicName>
, vs. reddish in
<taxonomicName id="3847F12173CB1735D498E4C8C8E13E35" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="ocotal">
<pageBreakToken id="F157964348488D6912814E88E0D9EC8F" pageNumber="3">R</pageBreakToken>
. ocotal
</taxonomicName>
); ventral angle of articular, acute (vs. right); first neural spine oriented rostrad (vs. caudad); circumpeduncular scales as few as 17 (vs. always more than 19); distance from caudal esophageal loop in gut to esophagus always greater than 24% gut length (vs. less than 16%).
</paragraph>
</subSubSection>
<subSubSection id="4FDC3E6CA34D0836E0BAF9E60ABFBEC8" pageNumber="3" type="multiple">
<caption id="052BDE6BADDDB65F778D60B9F80AF125" pageNumber="3">
<paragraph id="694447374F81E9E6460DE0D94A59BC76" pageNumber="58">
FIGURE 23.
<taxonomicName id="D7905973A34A159169DCE47E0D7B4E9D" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Rocio octofasciata</taxonomicName>
, holotype, MHNG 665.55. Photo, C. Ratton.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="588FB0EA492D7BF0415BFA1A58A1EFEB" pageNumber="3" type="description">
<paragraph id="56B364CC481015CECB5396D6043266B9" pageNumber="58">Description. D. XVII-XIX,8-10, modally 10 (7 of 174 with 11 dorsal fin rays); A. VIII-IX,7-9, modally 8 (2 of 177 with 6 or 7 anal-fin spines); pectoral 14-16. Gill rakers rounded, distally expanded or trapezoidal, sometimes bifid and even trifid, sometimes serrated. Scale rows on cheek modally 6 (4-7, 12 of 68 specimens with 7 scale rows); predorsal scales modally 12; pored lateral-line scales (not counting scales overlapping between the two segments of the lateral line) 27-30; scales from lateral line to origin of dorsal fin 4-5.5; scales from lateral line to base of first dorsal-fin ray 3-4 (further meristic data appear in Table 3).</paragraph>
<paragraph id="DEC339BFFDE7509B4A10C19EC5046404" pageNumber="58">
Largest specimen examined, 217 mm SL, but reported to 250 mm SL (
<bibRefCitation id="D40774CE4236084EA6584BBC87B57CCA" author="Page, L. M. &amp; Burr, B. M." pageNumber="3" refId="ref36813" refString="Page, L. M. &amp; Burr, B. M. (1991) A Field Guide to Freshwater Fishes of North America North of Mexico. Houghton Mifflin, Boston, 432 pp." type="book" year="1991">Page &amp; Burr 1991</bibRefCitation>
). Body usually slender, deeper in young (39-51% of SL). Head length 33-42% of SL; orbital diameter usually 21-25% of head length (up to 31% of head length in juveniles -further morphometric data appear in Table 4). Head profile convex, straight, concave on orbits. Maxilla reaching only a horizontal line, not a vertical, from orbit. Corners of lower lip slightly tapering, if at all.
</paragraph>
<paragraph id="AB4762CD5753031315869A549C511901" pageNumber="58">Pectoral and pelvic fins nearly always reaching caudad beyond 1st or 2nd anal-fin spine (not reaching in 2 of 127 specimens). Filamentous rays of dorsal fin to mid-caudal fin. Scales between dorsal and anal fin rays in one row, up to 6 scales long.</paragraph>
<paragraph id="1A9F8E1D692F16F77AA453557424903B" pageNumber="58">Gut simple, anal and anterior esophageal loops adjacent; gut length shorter or subequal to standard length of fish. Genital papilla thick, cylindrical, notched or triangular-tipped, sometimes sunk, longer than broad, medially swollen, somewhat crenulate at tip, sometimes rather vase-shaped; pigmented just in basal half and margins, if at all.</paragraph>
<paragraph id="24DADA87D11854D95CE3DA57ED36D752" pageNumber="58">Stripe from snout to eye diffuse or absent; suborbital streak narrow, with pointed ends (not always visible in preserved specimens). Eye color varied: yellowish, greyish, bluish, coppery, reddish. Bars on sides sometimes diffuse, sporadically dorsally or medially coalescent. Lateral blotch oval to squarish, joining 3rd and 4th bars, sometimes not discernible from longitudinal stripe. About 15 rows of light spots on sides, centered in each scale, not always visible; breast region olive. Axil of pectoral fin with same coloration as breast or dusky; base of pectoral fin paler than surrounding region. For life colors, see Fig. 24.</paragraph>
</subSubSection>
<subSubSection id="934ECC9FE11C286217E99A526CDB713A" pageNumber="3" type="multiple">
<caption id="A63292F581E7FF028EFEAD95A346463F" pageNumber="3">
<paragraph id="EC2645165A40F49FD8F661B35B82922A" pageNumber="59">
<pageBreakToken id="82A71D27CD3D2E324A2F80E4817E76AF" pageNumber="3">FIGURE</pageBreakToken>
24.
<taxonomicName id="EA084D8E7CA2DF9541E5999DC1E93FBE" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Rocio octofasciata</taxonomicName>
live. Locality: Arroyo Tres
<normalizedToken id="23D0F27D60E7DC8DFC7137F2304D7FCE" originalValue="Garantías" pageNumber="3">Garantias</normalizedToken>
, Quintana Roo, Mexico. Photo, H. Bahena.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DF94CAFE05986882D92CF279C4AF849F" pageNumber="3" type="distribution">
<paragraph id="6268891E557E66EE819921C5690FD991" pageNumber="59">
Distribution. From
<normalizedToken id="6B00CFF8FAB7C2DC6FC1AC37A5584F8D" originalValue="Río" pageNumber="3">Rio</normalizedToken>
<normalizedToken id="A78CFC8737C5FA8208C8A31F6ECD1789" originalValue="Ulúa" pageNumber="3">Ulua</normalizedToken>
, Honduras (
<bibRefCitation id="1CAE69E9AAE46405653A5B938F9A1BCC" author="Lee, D. S. &amp; Gilbert, C. R. &amp; Hocutt, C. H. &amp; Jenkins, R. E. &amp; McAllister, D. E. &amp; Stauffer, J. R." pageNumber="3" refId="ref36087" refString="Lee, D. S., Gilbert, C. R., Hocutt, C. H., Jenkins, R. E., McAllister, D. E. &amp; Stauffer, J. R. (1980) Atlas of North American Freshwater Fishes. North Carolina State Museum of Natural History, 867 pp." type="book" year="1980">Lee et al. 1980</bibRefCitation>
), to tributaries of
<normalizedToken id="924216585E8BFBE6C0B3AE31DFDFBC9B" originalValue="Río" pageNumber="3">Rio</normalizedToken>
Actopan, Veracruz, Mexico (
<bibRefCitation id="177C42E20E20A289AFB583D29B26CC4C" author="Greenfield, D. W. &amp; Thomerson, J. E." pageNumber="3" refId="ref35309" refString="Greenfield, D. W. &amp; Thomerson, J. E. (1997) Fishes of the Continental Waters of Belize. University Press of Florida, Gainesville, 311 pp." type="book" year="1997">Greenfield &amp; Thomerson 1997</bibRefCitation>
) (Fig. 22). Introduced in several localities in northern Veracruz (
<bibRefCitation id="63440E16E10E8E506C0E09A5925D48B8" author="Obregon B. &amp; Contreras-Balderas, S. &amp; Lozano-Vilano, M. L." pageNumber="3" pagination="79 - 95" refId="ref36774" refString="Obregon B., H., Contreras-Balderas, S. &amp; Lozano-Vilano, M. L. (1994) The fishes of northern and central Veracruz, Mexico. Hydrobiologia, 286, 79 - 95." type="journal article" year="1994">
<normalizedToken id="C9A085084A49BFA1FC628256DD800A9C" originalValue="Obregón" pageNumber="3">Obregon</normalizedToken>
et al. 1994
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="395F679C817F4EADA6BDE0C4EEE32E6D" pageNumber="3" type="discussion">
<paragraph id="A91BAAA4E4CE204BAE17DB73CCB69CBB" pageNumber="59">
Remarks. Contra Regans (1903) original designation (and S. Fisch -
<normalizedToken id="85B03A400FC7FCFFDEC65BE66CCC8C70" originalValue="Müller" pageNumber="3">Mueller</normalizedToken>
s opinion, pers. com.),
<bibRefCitation id="95B589839B51B624B7FEED911BA483C0" author="Mahnert, V." pageNumber="3" pagination="471 - 496" refId="ref36270" refString="Mahnert, V. (1976) Catalogue des types de poissons, amphibiens, et reptiles du Museum d'Histoire naturelle de Geneve. Revue Suisse de Zoologie, 83, 471 - 496." type="journal article" year="1976">Mahnert(1976: 44)</bibRefCitation>
treated the nine specimens originally in MHNG 665.55 as syntypes. However, there is little doubt that the largest individual is the holotype. The confusion was likely created by
<bibRefCitation id="41551D6505E7011C48B42C5137CB8125" author="Regan, C. T." pageNumber="3" pagination="255 - 259" refId="ref36963" refString="Regan, C. T. (1904) Descriptions of new or little-known fishes from Mexico and British Honduras. Annual Magazine of Natural History, 13, 255 - 259." type="journal article" year="1904">Regan (1904</bibRefCitation>
,
<bibRefCitation id="3998FF616A2F0BF6DDA38A3191A7B774" author="Regan, C. T." pageNumber="3" pagination="433 - 445" refId="ref36996" refString="Regan, C. T. (1905) A revision of the fishes of the American cichlid genus Cichlosoma and of the allied genera. Annual Magazine of Natural History, 16, 60 - 77, 225 - 243, 316 - 340, 433 - 445." type="journal article" year="1905">1905</bibRefCitation>
) himself, when he added material to his successive redescriptions, somewhat more accessible than the original description.
</paragraph>
<paragraph id="FA4CECE9D0B5D39CCFEC29438F3E119E" pageNumber="59">
<taxonomicName id="AE30041A5C4E2D68D774EAC511FB2614" LSID-ZBK="urn:lsid:zoobank.org:act:E7A510F9-CF8C-4181-A6CE-DD7BA7F860F2" family="Cichlidae" genus="Cichlosoma" kingdom="Animalia" lsidName-ZBK="Cichlosoma biocellatum Regan 1909:234" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="biocellatum">Cichlosoma biocellatum</taxonomicName>
, based on an aquarium specimen, is a synonym, its type locality (“
<normalizedToken id="BC2B5FC722682E4F3A47E37B8FD42C7A" originalValue="Río" pageNumber="3">Rio</normalizedToken>
Negro at
<normalizedToken id="B8C5C1AD68984E2C0DB064610FF1BB07" originalValue="Mañaos" pageNumber="3">Manaos</normalizedToken>
[sic], Brazil”) certainly in error.
</paragraph>
<paragraph id="DC1EBD6F0FC4382D7EF731CACAC7BAC4" pageNumber="59">
<taxonomicName id="A487328C29B37D3F763159214977286E" LSID-ZBK="urn:lsid:zoobank.org:act:3A693BD7-FD95-4532-9F14-41B963B40D09" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma hedricki Meek 1904:208" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="hedricki">Cichlasoma hedricki</taxonomicName>
was synonymized by
<bibRefCitation id="C0034C688BE229A2FD1B20BF24FF6704" author="Regan, C. T." pageNumber="3" pagination="433 - 445" refId="ref36996" refString="Regan, C. T. (1905) A revision of the fishes of the American cichlid genus Cichlosoma and of the allied genera. Annual Magazine of Natural History, 16, 60 - 77, 225 - 243, 316 - 340, 433 - 445." type="journal article" year="1905">Regan (1905)</bibRefCitation>
, barely one year after Meeks (1904) description, without explicit reasons. The action was followed by most authors. Topotypes of
<taxonomicName id="86C46D2943840693C93F33F764A803E1" LSID-ZBK="urn:lsid:zoobank.org:act:3A693BD7-FD95-4532-9F14-41B963B40D09" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma hedricki Meek 1904:208" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="hedricki">C. hedricki</taxonomicName>
(from the upper Papaloapan) tend to be deeper-bodied than other
<taxonomicName id="810E4AAD8A21753E74099570E7CD3B1F" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">R. octofasciata</taxonomicName>
, and the presence of a prominent lingual cusp in their symphysial teeth (fig. 15a in Schmitter-Soto, in press) tends to be more constant, but the forms merge imperceptibly towards the type locality of
<taxonomicName id="A5C0FF322DA6DBE08AE25EFADDEE9DE6" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">R. octofasciata</taxonomicName>
in the Coatzacoalcos, the drainage immediately to the south of Papaloapan.
</paragraph>
<paragraph id="F2BC58C7A8BB7F6CDC2CF95ACCD026DC" pageNumber="59">
<materialsCitation id="C66444578FF661ED3B1361538C3449C0" ID-GBIF-Occurrence="923904138" collectionCode="UMMZ" country="Mexico" location="Rio Obispo" pageNumber="3" specimenCode="UMMZ 176671" stateProvince="Oaxaca" typeStatus="paratype">
In print,
<bibRefCitation id="FF76B4D23CDE90054FFB47E0C8E6B5EE" author="Meek, S. E." pageNumber="3" pagination="1 - 252" refId="ref36415" refString="Meek, S. E. (1904) The fresh-water fishes of Mexico north of the Isthmus of Tehuantepec. Field Columbian Museum of Natural History Publications, Zoological Series, 93, 1 - 252." type="journal article" year="1904">Meek (1904)</bibRefCitation>
called the type locality of
<taxonomicName id="0A839D8FD0DD8BF976286F1563A52750" LSID-ZBK="urn:lsid:zoobank.org:act:3A693BD7-FD95-4532-9F14-41B963B40D09" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma hedricki Meek 1904:208" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="hedricki">C. hedricki</taxonomicName>
“Obispo, Veracruz;” however, his manuscript label in the
<typeStatus id="A189B84C7977D9D112314D02BEEA725A" pageNumber="3" type="paratype">paratype</typeStatus>
jar at
<specimenCode id="ABBFCC476DF881156913328462B6A18A" pageNumber="3">
<collectionCode id="2F93456AA8A65A2E374225D2D4F74C91" pageNumber="3">UMMZ</collectionCode>
(176671)
</specimenCode>
reads “
<location id="A1C646617637B64FBDF70B2FD3BA8175" LSID="urn:lsid:plazi:treatment:131484AFA72F82190797063675C7E124:A1C646617637B64FBDF70B2FD3BA8175" country="Mexico" name="Rio Obispo" pageNumber="3" stateProvince="Oaxaca">
<normalizedToken id="A40361A9D4A3EAAE85FB25216224228C" originalValue="Río" pageNumber="3">Rio</normalizedToken>
Obispo
</location>
,
<collectingRegion id="AD05B9D1D944E0CE9B09EA12B21EDB76" pageNumber="3">Oaxaca</collectingRegion>
.”
</materialsCitation>
The eight
<typeStatus id="501B37757CD2235D3BA635DB999C8AD3" pageNumber="3" type="paratype">paratypes</typeStatus>
in this lot (labelled as such by Meek himself) should probably be added to the 44 specimens mentioned by
<bibRefCitation id="FCF88B738602913D97ECBCBAD9D0EBE4" author="Eschmeyer, W. N." pageNumber="3" refId="ref35012" refString="Eschmeyer, W. N. (Ed.) (2005) Catalog of Fishes. California Academy of Sciences, San Francisco. Available from http: // www. calacademy. org / research / ichthyology / Catalog / (accessed 15 January 2005)" type="url" year="2005">Eschmeyer (2005)</bibRefCitation>
; however, on the other hand, the original description mentions only one “type,” thus rendering the literature (and paratype holdings at UMMZ and FMNH) in error.
</paragraph>
</subSubSection>
</treatment>
</document>