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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1166.101444" ID-Pensoft-Pub="1313-2970-1166-103" ID-Pensoft-UUID="1BB6324D5B4B5443A6C1B84EC2844A11" ID-ZooBank="362C54108ABA48A4880C056BF68B23F6" ModsDocID="1313-2970-1166-103" checkinTime="1686259419846" checkinUser="pensoft" docAuthor="Constant, Jerome &amp; Pham, Thai Hong" docDate="2023" docId="FD28F8153B11568E9415B0C3EC6CB181" docLanguage="en" docName="ZooKeys 1166: 103-119" docOrigin="ZooKeys 1166" docPubDate="2023-06-08" docSource="http://dx.doi.org/10.3897/zookeys.1166.101444" docTitle="Pumatiracia Constant &amp; Pham 2023, gen. nov." docType="treatment" docUuid="C389D7E6-5FB1-4D47-8D64-E78E04A1FA41" docUuidSource="ZooBank" docVersion="1" id="1BB6324D5B4B5443A6C1B84EC2844A11" lastPageNumber="103" masterDocId="1BB6324D5B4B5443A6C1B84EC2844A11" masterDocTitle="Parahiraciini planthoppers with elongate head from Vietnam: a new genus and species Pumatiracia venosa gen. et sp. nov. and first record of Laohiracia acuta Constant, 2021 (Hemiptera, Fulgoromorpha, Issidae)" masterLastPageNumber="119" masterPageNumber="103" pageNumber="103" updateTime="1686259419846" updateUser="pensoft">
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<mods:title id="BE672C84D51BDA86E309883E4205A5A9">Parahiraciini planthoppers with elongate head from Vietnam: a new genus and species Pumatiracia venosa gen. et sp. nov. and first record of Laohiracia acuta Constant, 2021 (Hemiptera, Fulgoromorpha, Issidae)</mods:title>
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<mods:namePart id="AD1232C0FDFB106E34028F44B408A2EB">Constant, Jerome</mods:namePart>
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<mods:affiliation id="80C68AB529658F10772926AF1DCF02DD">Royal Belgian Institute of Natural Sciences, O. D. Taxonomy &amp; Phylogeny - Entomology, Vautier street 29, B- 1000 Brussels, Belgium</mods:affiliation>
<mods:nameIdentifier id="4BA496CF76743B6BB25F02A4D51DC384" type="email">jconstant@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="40CDD33219452EF946CC5E9362A4953C">Pham, Thai Hong</mods:namePart>
<mods:nameIdentifier id="96D13C89A75468B8FDAAC9BE8307F753" type="ORCID">https://orcid.org/0000-0002-4763-3679</mods:nameIdentifier>
<mods:affiliation id="5994EA1AB91DCCF0FC296E5609DBDAC3">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="BD4892F7103CCEBB2FE94FF34C5CCD5F" type="email">phamthai1976@yahoo.com</mods:nameIdentifier>
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<mods:date id="09331EC6D7A79601F4677CB872AABF8B">2023</mods:date>
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<subSubSection id="C014CC468F99A6DDF28CA3BDD375275C" pageId="0" pageNumber="103" type="nomenclature">
<paragraph id="ED58705724211245C7CFDA9F387A6D35" pageId="0" pageNumber="103">
Genus
<taxonomicName id="D01D20222262D01B6934A33AEAF66C56" LSID="https://zoobank.org/C389D7E6-5FB1-4D47-8D64-E78E04A1FA41" authority="Constant &amp; Pham, 2023" authorityName="Constant &amp; Pham" authorityYear="2023" family="Issidae" genus="Pumatiracia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pumatiracia" order="Hemiptera" pageId="0" pageNumber="103" rank="genus" status="gen. nov.">Pumatiracia</taxonomicName>
<taxonomicNameLabel id="DADD0D32A43C71DEA41FCABFBC59FD6A" pageId="0" pageNumber="103">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6E21C13FED81F7CDCD3B1CA14BCBDF6E" pageId="0" pageNumber="103" type="type species">
<paragraph id="203DFCF86B27FFBC5593EABE76D38296" pageId="0" pageNumber="103">Type species.</paragraph>
<paragraph id="2F26A8E364939E4DCBF04631FFEA7192" pageId="0" pageNumber="103">
<taxonomicName id="88046C4F128632B75647576873B60CDF" authorityName="gen. et" authorityYear="2023" family="Issidae" genus="Pumatiracia" kingdom="Animalia" lsidName="Pumatiracia venosa" order="Hemiptera" pageId="0" pageNumber="103" rank="species" species="venosa">
<emphasis id="D8B01DC6C74188A83D61B04A6618A5A8" italics="true" pageId="0" pageNumber="103">Pumatiracia venosa</emphasis>
</taxonomicName>
gen. et sp. nov. by present designation.
</paragraph>
</subSubSection>
<subSubSection id="B7E0229D101D56F24EC4B039BB6A9975" pageId="0" pageNumber="103" type="diagnosis">
<paragraph id="59553C9B204514CBE0B5C04F5CA1A61D" pageId="0" pageNumber="103">Diagnosis.</paragraph>
<paragraph id="271A5B73CE8CA6E57043705757BE8EBB" pageId="0" pageNumber="103">
The genus can be separated from all other
<taxonomicName id="16D73FB7C9263BC8C9BFD367DD6640AB" authorityName="Cheng &amp; Yang" authorityYear="1991" lsidName="" pageId="0" pageNumber="103" rank="tribe" tribe="Parahiraciini">Parahiraciini</taxonomicName>
genera by the following combination of characters:
</paragraph>
<paragraph id="678112E97EA95E289151B53229D76FED" pageId="0" pageNumber="103">
Head with vertex elongate, 1.2
<normalizedToken id="2A324D7BF77647FF4F1B37C6F7C23EBF" originalValue="×">x</normalizedToken>
longer in median line than maximal width, projecting well beyond eyes in dorsal view, but not forming a real cephalic process;
</paragraph>
<paragraph id="75398AB3D2AE19876ECC9273A466ECD3" pageId="0" pageNumber="103">Vertex and frons with a median carina;</paragraph>
<paragraph id="6F7B7F7035A112D34F022F998B9002E7" pageId="0" pageNumber="103">Genae with strong carina under the antennae;</paragraph>
<paragraph id="1B007AA538CF6D8023F9409E5781A7A6" pageId="0" pageNumber="103">Body elongate, more than twice a long as maximum width, with side margins broadly rounded in dorsal view;</paragraph>
<paragraph id="4206B2BFB9B6BA60D4071017D0E33804" pageId="0" pageNumber="103">Metatibiae with three lateral and eight apical spines;</paragraph>
<paragraph id="158389E27B24E30659B57C84BDDD9B3E" pageId="0" pageNumber="103">Anal tube of male dorsoventrally flattened, subtriangular in dorsal view furcate distally;</paragraph>
<paragraph id="91EB591CFEAC9922D2696BF13093B737" pageId="0" pageNumber="103">Posterior margin of pygofer broadly rounded;</paragraph>
<paragraph id="A0017C92B93BFA59096429003BF216B2" pageId="0" pageNumber="103">Gonostyli with capitulum strongly projecting dorsad;</paragraph>
<paragraph id="865DDC6D93CCD28D6CEF97D6CDF5AEC1" pageId="0" pageNumber="103">Aedeagus evenly curved in lateral view, with pair of elongate symmetrical, lateroventral processes projecting cephalad.</paragraph>
</subSubSection>
<subSubSection id="649D1D8A077C879B83BED48A70C9264C" pageId="0" pageNumber="103" type="description">
<paragraph id="DD6D3C420A7B42EEE8DD9801BE69DF82" pageId="0" pageNumber="103">Description.</paragraph>
<paragraph id="C8EA92CF71AD5E8F8CC8B36D6493488B" pageId="0" pageNumber="103">
<emphasis id="C4F4341160E774717A660DC4B146A5E3" bold="true" italics="true" pageId="0" pageNumber="103">Head</emphasis>
: (Figs
<figureCitation id="1311A49FC1E32C8564F1E78D9B204C50" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pumatiracia venosa gen. et sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D right hind wing E metatarsus, detail, ventral view F metatibia and metatarsus, ventral view." figureDoi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103">1A-C</figureCitation>
,
<figureCitation id="D9A38F395E563719E73663739C248257" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pumatiracia venosa gen. et sp. nov., holotype A-D detail of head and thorax A dorsal view B perpendicular view of frons C lateral view D anterolateral view E, F venation of wings E right tegmen F right hind wing." figureDoi="10.3897/zookeys.1166.101444.figure2" httpUri="https://binary.pensoft.net/fig/861475" pageId="0" pageNumber="103">2A-D</figureCitation>
) Head narrower than thorax and elongate but without forming a definite cephalic process. Vertex elongate, as long in mid-line as pronotum, ~ 1.2
<normalizedToken id="FAFA10E6AA7B9CD552E9BF52E6CEC02C" originalValue="×">x</normalizedToken>
as long in mid-line as broad basally, with sides subparallel, slightly converging towards rounded apex; strong median carina not reaching apex; lateral margins carinate with carinae merging anteriorly; posterior margin of vertex angularly emarginate and carinate. Frons elongate, mostly flat with median carina, wider above fronto-clypeal suture, slightly tapering towards rounded dorsal margin; lateral margins carinate; oblique lateral carinae on distal
<normalizedToken id="890DA513E9162165A23D834C0FFEC2CE" originalValue="¼">1/4</normalizedToken>
, merging dorsally with anterior carina of vertex. Genae with blunt carina running from under the antenna obliquely to fronto-clypeal suture. Clypeus flat in middle portion, moderately elongate, subtriangular with fronto-clypeal suture rounded; anteclypeus with median blunt carina. Labium elongate and narrow, reaching metacoxae, with apical segment elongate, nearly as long as penultimate. Eyes reniform (not emarginate) protruding laterally; ocelli absent. Antennae rather short with scape ring-shaped and pedicel cylindrical, slightly longer than broad.
</paragraph>
<caption id="CBBC2AB44AB78FC0E89E537E7FA9B7D9" doi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103" start="Figure 1" startId="F1">
<paragraph id="C71B75A72AE24E8BE819585E05773447" pageId="0" pageNumber="103">
<emphasis id="FC2364AFC91610DFB275DBB6E3C5A2FF" bold="true" pageId="0" pageNumber="103">Figure 1.</emphasis>
<taxonomicName id="B5F576111EB8C134D31F4AF20EF53E9E" authorityName="gen. et" authorityYear="2023" family="Issidae" genus="Pumatiracia" kingdom="Animalia" lsidName="Pumatiracia venosa" order="Hemiptera" pageId="0" pageNumber="103" rank="species" species="venosa">
<emphasis id="68198E19A55C598ABC8B7DCDD3FF986C" italics="true" pageId="0" pageNumber="103">Pumatiracia venosa</emphasis>
</taxonomicName>
gen. et sp. nov., holotype ♂
<emphasis id="DFE9F4EE8EE138E9E83DE9CF5A3B86CA" bold="true" pageId="0" pageNumber="103">A</emphasis>
habitus, dorsal view
<emphasis id="A2707120C2C844B75F018A49E458E52C" bold="true" pageId="0" pageNumber="103">B</emphasis>
habitus, ventral view
<emphasis id="BD0588DA4DA0C5177ABF516C163F4592" bold="true" pageId="0" pageNumber="103">C</emphasis>
habitus, lateral view
<emphasis id="849A81E0141769BC6BA3BB24802CF41F" bold="true" pageId="0" pageNumber="103">D</emphasis>
right hind wing
<emphasis id="C7159E35A603BF8A1B2E0FD2C8CFCE92" bold="true" pageId="0" pageNumber="103">E</emphasis>
metatarsus, detail, ventral view
<emphasis id="EE3124A17617E46E7BD60E515FCB567D" bold="true" pageId="0" pageNumber="103">F</emphasis>
metatibia and metatarsus, ventral view.
</paragraph>
</caption>
<caption id="87C88A9CC5FDBEF6933DB3F158DA1F99" doi="10.3897/zookeys.1166.101444.figure2" httpUri="https://binary.pensoft.net/fig/861475" pageId="0" pageNumber="103" start="Figure 2" startId="F2">
<paragraph id="7B8750474EA9A93F36CEEB542B3ACA83" pageId="0" pageNumber="103">
<emphasis id="43B6B23882DF1DEBCA8234982DB01887" bold="true" pageId="0" pageNumber="103">Figure 2.</emphasis>
<taxonomicName id="D1A28E4B5F309ED7E5B09B49949FF800" authorityName="gen. et" authorityYear="2023" family="Issidae" genus="Pumatiracia" kingdom="Animalia" lsidName="Pumatiracia venosa" order="Hemiptera" pageId="0" pageNumber="103" rank="species" species="venosa">
<emphasis id="15893862B88E4ED91D4FC0C769658724" italics="true" pageId="0" pageNumber="103">Pumatiracia venosa</emphasis>
</taxonomicName>
gen. et sp. nov., holotype
<emphasis id="5A657341BC41FAB862CBB8E41A954E70" bold="true" pageId="0" pageNumber="103">A-D</emphasis>
detail of head and thorax
<emphasis id="716EF7C7C244BA63C6DFA4881154D8CB" bold="true" pageId="0" pageNumber="103">A</emphasis>
dorsal view
<emphasis id="D0EC225B9AB645006352A0E90FED42C8" bold="true" pageId="0" pageNumber="103">B</emphasis>
perpendicular view of frons
<emphasis id="9F5D10378A3D0ECE18016B18D603CDEA" bold="true" pageId="0" pageNumber="103">C</emphasis>
lateral view
<emphasis id="D0443D34BFAFE129306693FF3A7D6829" bold="true" pageId="0" pageNumber="103">D</emphasis>
anterolateral view
<emphasis id="DFB92A3DAC5553917D83F62679F6CB56" bold="true" pageId="0" pageNumber="103">E, F</emphasis>
venation of wings
<emphasis id="A71BAB0DC15621CE448FC332CFEFB1BC" bold="true" pageId="0" pageNumber="103">E</emphasis>
right tegmen
<emphasis id="2C4944CB7E010CC0490E61340097CD39" bold="true" pageId="0" pageNumber="103">F</emphasis>
right hind wing.
</paragraph>
</caption>
<paragraph id="4923EE51B1024CBBB9126B409355DB4A" pageId="0" pageNumber="103">
<emphasis id="4D35121D4CBBDD900ABF6BF5A07A45BB" bold="true" italics="true" pageId="0" pageNumber="103">Thorax</emphasis>
: (Figs
<figureCitation id="7CDB25E0FD0A0A0730E640A9CEFF1E6B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pumatiracia venosa gen. et sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D right hind wing E metatarsus, detail, ventral view F metatibia and metatarsus, ventral view." figureDoi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103">1A, C</figureCitation>
,
<figureCitation id="A3756ADD9A553D9079A0E647A0D045BA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pumatiracia venosa gen. et sp. nov., holotype A-D detail of head and thorax A dorsal view B perpendicular view of frons C lateral view D anterolateral view E, F venation of wings E right tegmen F right hind wing." figureDoi="10.3897/zookeys.1166.101444.figure2" httpUri="https://binary.pensoft.net/fig/861475" pageId="0" pageNumber="103">2A-D</figureCitation>
) Pronotum slightly shorter than mesonotum in mid-line; anterior margin carinate in middle portion, strongly sinuate and strongly, roundly protruding anteriorly between eyes; posterior margin nearly truncate; median carina obsolete anteriorly with impressed point on each side; blunt tubercles along anterior margin and irregularly on disc and sides; paranotal lobes (lateral view) broad, with tubercles, sometimes turning into short, blunt, longitudinal carinae along posterior margin, with blunt carina behind level of antenna, and with posteroventral angle rounded. Mesonotum subtriangular with obsolete median carina and transverse anterior carina; disc smooth with additional, more or less undulate, longitudinal carina; weak, blunt slightly rounded longitudinal carina on each side of disc; some blunt tubercles in lateral fields of mesonotum. Tegulae moderately developed.
</paragraph>
<paragraph id="F0AD3E0EC033F4705909CC71A8CD1EB7" pageId="0" pageNumber="103">
Tegmina: (Figs
<figureCitation id="9E34F8EBA91556326329A0BC50C868E7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pumatiracia venosa gen. et sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D right hind wing E metatarsus, detail, ventral view F metatibia and metatarsus, ventral view." figureDoi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103">1A-C</figureCitation>
,
<figureCitation id="900DBB81D18A33B22B330E91B835247B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pumatiracia venosa gen. et sp. nov., holotype A-D detail of head and thorax A dorsal view B perpendicular view of frons C lateral view D anterolateral view E, F venation of wings E right tegmen F right hind wing." figureDoi="10.3897/zookeys.1166.101444.figure2" httpUri="https://binary.pensoft.net/fig/861475" pageId="0" pageNumber="103">2E</figureCitation>
) Tegmina subcoriaceous with longitudinal veins elevated and with a dense reticulum of veinlets, elongate with sides broadly rounded, ~ 2.7
<normalizedToken id="EBC196A40D902ABFBE37AD0994E19951" originalValue="×">x</normalizedToken>
longer than broad, convex. Apex narrowly rounded. Postclaval margin weakly rounded on distal half and slightly notched at apex of clavus. Clavus closed, reaching ca. mid-length of tegmen.
</paragraph>
<paragraph id="F9108C5156F2E27224D9EAD1177C8D05" pageId="0" pageNumber="103">
Venation: ScP+R moderately developed, forking into subparallel ScP+RA and RP; MP forking at ~
<normalizedToken id="12008B82BF2547CE204B8417DA796755" originalValue="¾">3/4</normalizedToken>
of tegmen length; CuA forked slightly after half of clavus length; CuA1+2 forked close to its base; CuA1, CuA2 and CuA3 subparallel, with CuA1 merging distally with MP2; Pcu fused with A1 at
<normalizedToken id="C656179FBF39F489971220E0795CD592" originalValue="¾">3/4</normalizedToken>
of clavus length; Pcu+A1 fused with CuP slightly before apex of clavus.
</paragraph>
<paragraph id="62359F356C661F74D87803AC5A7ADBA3" pageId="0" pageNumber="103">
Hind wings: (Figs
<figureCitation id="44CA348C9E5B54E22B5C90A670D147A7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pumatiracia venosa gen. et sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D right hind wing E metatarsus, detail, ventral view F metatibia and metatarsus, ventral view." figureDoi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103">1 D</figureCitation>
,
<figureCitation id="7C1AA1406BBEEF1595CC11EE44FBE1E0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pumatiracia venosa gen. et sp. nov., holotype A-D detail of head and thorax A dorsal view B perpendicular view of frons C lateral view D anterolateral view E, F venation of wings E right tegmen F right hind wing." figureDoi="10.3897/zookeys.1166.101444.figure2" httpUri="https://binary.pensoft.net/fig/861475" pageId="0" pageNumber="103">2 F</figureCitation>
) Broader than tegmina and deeply bilobed, strongly notched at CuP; costal margin rather weakly sinuate; CuP-Pcu-A1 lobe ~ 1/3 wider than Sc-R-MP-CuA lobe, the two lobes almost the same length; both lobes rounded apically; postclaval margin broadly rounded; A2 lobe reduced and narrow, with A2 vein obsolete.
</paragraph>
<paragraph id="2F24D7651198A6EB008E696AC5237419" pageId="0" pageNumber="103">Venation: main veins present; ScP+R, MP and CuA running more or less parallel, slightly diverging towards apex, with numerous cross-veins; Pcu strongly curved around basal third of wing towards CuP but not reaching the latter; A1 curved, more or less parallel to postclaval margin; CuP-Pcu-A1 lobe with numerous cross-veins.</paragraph>
<paragraph id="775D36834466C876AC151B32EB468371" pageId="0" pageNumber="103">
Legs: (Fig.
<figureCitation id="F5B181D4F8BD8660F391EE543C27F193" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pumatiracia venosa gen. et sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D right hind wing E metatarsus, detail, ventral view F metatibia and metatarsus, ventral view." figureDoi="10.3897/zookeys.1166.101444.figure1" httpUri="https://binary.pensoft.net/fig/861474" pageId="0" pageNumber="103">1A-C, E, F</figureCitation>
) Strongly elongate and slender. Pro- and mesofemora and protibiae slightly flattened dorsoventrally. Tibiae longer than corresponding femora. Metatibiae with one lateral tooth near base, two lateral teeth placed on distal half and eight apical teeth. Tarsi elongate; first metatarsomere elongate and slender, with a strong spine at each side and a row of six smaller spines in between ventrally along posterior margin; second metatarsomere short with one tooth at each side. Metatibiotarsal formula: (3) 8 / 8 / 2.
</paragraph>
<paragraph id="BB339A014AF05BA1F6849A14C7057E03" pageId="0" pageNumber="103">
<emphasis id="6727FF1D0BB9566733A7C7124A2F95FE" bold="true" italics="true" pageId="0" pageNumber="103">Terminalia</emphasis>
♂: (Fig.
<figureCitation id="1C31AB1E199380BD96D490B058D7A255" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pumatiracia venosa gen. et sp. nov., holotype, terminalia ♂ A-E pygofer, gonostyli and anal tube A lateral view B posterolateral view C dorsal view D ventral view E posterior view F-L aedeagus, phallobase and connective F left lateral view G left lateroventral view H dorsal view I apical portion, anterodorsal view J ventral view K posteroventral view L anteroventral view." figureDoi="10.3897/zookeys.1166.101444.figure3" httpUri="https://binary.pensoft.net/fig/861476" pageId="0" pageNumber="103">3</figureCitation>
) Pygofer higher than long in lateral view, with anterior margin weakly concave and posterior margin broadly rounded. Gonostyli (in lateral view) elongate and broad, projecting posteriorly, with capitulum strongly developed dorsad, with lateral laminate projection and with moderately developed neck with outer margin strongly concave in posterior view; in ventral view, gonostyli abruptly narrowing after basal
<normalizedToken id="B61A74BC3F50AF0C7E2F79F9B279594E" originalValue="¼">1/4</normalizedToken>
. Anal tube dorsoventrally flattened, broadly subtriangular. Aedeagus moderately curved dorsad (in lateral view), with symmetrical pair of elongate medioventral processes directed cephalad. Connective elongate.
</paragraph>
<caption id="D0CDB4D12DC0D445853B63E82C0B4590" doi="10.3897/zookeys.1166.101444.figure3" httpUri="https://binary.pensoft.net/fig/861476" pageId="0" pageNumber="103" start="Figure 3" startId="F3">
<paragraph id="B930D0BD0F76C9C171FE421549160FBB" pageId="0" pageNumber="103">
<emphasis id="C663E508013DE7C261B79F4CB75DCC40" bold="true" pageId="0" pageNumber="103">Figure 3.</emphasis>
<taxonomicName id="22404654B5BBA6B2137E312938242CC9" authorityName="gen. et" authorityYear="2023" family="Issidae" genus="Pumatiracia" kingdom="Animalia" lsidName="Pumatiracia venosa" order="Hemiptera" pageId="0" pageNumber="103" rank="species" species="venosa">
<emphasis id="0725460004A9EDF0C248DFF7E481FB2A" italics="true" pageId="0" pageNumber="103">Pumatiracia venosa</emphasis>
</taxonomicName>
gen. et sp. nov., holotype, terminalia ♂
<emphasis id="3C6EA6D10E09D36DCDB537E8FC3AEA19" bold="true" pageId="0" pageNumber="103">A-E</emphasis>
pygofer, gonostyli and anal tube
<emphasis id="729FEC517A375D5D65BAD818457ABE4D" bold="true" pageId="0" pageNumber="103">A</emphasis>
lateral view
<emphasis id="D19F8266AE81A41C526AB6327230B1AD" bold="true" pageId="0" pageNumber="103">B</emphasis>
posterolateral view
<emphasis id="C65798F01E0116C9F68342BE33295C8D" bold="true" pageId="0" pageNumber="103">C</emphasis>
dorsal view
<emphasis id="6D66613690B5AEC0451AA6E6DD6CB630" bold="true" pageId="0" pageNumber="103">D</emphasis>
ventral view
<emphasis id="4743ECB9E67AAC33DCAEABA601DA9A45" bold="true" pageId="0" pageNumber="103">E</emphasis>
posterior view
<emphasis id="BEDD7AF43133D39362AB657ADDC70EA0" bold="true" pageId="0" pageNumber="103">F-L</emphasis>
aedeagus, phallobase and connective
<emphasis id="ACA77E08CE591BDDB67523404EFDE0D5" bold="true" pageId="0" pageNumber="103">F</emphasis>
left lateral view
<emphasis id="8C52D5AB2E6FE727A3448A2A1FBAB0B9" bold="true" pageId="0" pageNumber="103">G</emphasis>
left lateroventral view
<emphasis id="EF959ED08729F004906C5BFCB19C6203" bold="true" pageId="0" pageNumber="103">H</emphasis>
dorsal view
<emphasis id="D4C17DE9C33E0F9E4FD51FED488D3C83" bold="true" pageId="0" pageNumber="103">I</emphasis>
apical portion, anterodorsal view
<emphasis id="EC0D78B8A12FE19B92CAC9E1791BB7B0" bold="true" pageId="0" pageNumber="103">J</emphasis>
ventral view
<emphasis id="82D14C0C1879142CBD5A86E587E10B7A" bold="true" pageId="0" pageNumber="103">K</emphasis>
posteroventral view
<emphasis id="C01AA8E7CB02E3DCC8EF0910D9D725AA" bold="true" pageId="0" pageNumber="103">L</emphasis>
anteroventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1A55EBD67EBCDBF7A6E6083BEFB0903F" pageId="0" pageNumber="103" type="etymology">
<paragraph id="71F5B09159BD75AD1C323039C8B22A88" pageId="0" pageNumber="103">Etymology.</paragraph>
<paragraph id="09531B22E15599405F726072F39A53EC" pageId="0" pageNumber="103">
The new genus name is formed by the combination of
<emphasis id="A62DCF248ADD55DADAEFF411067B4CA6" italics="true" pageId="0" pageNumber="103">Pu Mat</emphasis>
, referring to the National Park where the new genus was found, and
<emphasis id="858A54DC034A5DEB5471C531B0AD1DEC" italics="true" pageId="0" pageNumber="103">hiracia</emphasis>
, which is the same ending as in
<taxonomicName id="1F855D28578B16539A232B7ADE7C1374" authorityName="Ouchi" authorityYear="1940" class="Insecta" family="Issidae" genus="Parahiracia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parahiracia" order="Hemiptera" pageId="0" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="6ED51B3F30BAF72064E0C0AB1C3E502C" italics="true" pageId="0" pageNumber="103">Parahiracia</emphasis>
</taxonomicName>
Ouchi, 1940 (synonymized by
<bibRefCitation id="742963B7854927757AB3D3608AAEDB8B" DOI="https://doi.org/10.31610/zsr/2003.12.2.217" author="Gnezdilov, VM" journalOrPublisher="Arthropod Systematics &amp; Phylogeny" pageId="0" pageNumber="103" refId="B14" refString="Gnezdilov, VM, Drosopoulos, S, Wilson, MR, 2004. New data on taxonomy and distribution of some Fulgoroidea (Homoptera, Cicadina). Zoosystematica Rossica 12(2): (2003) 217-223. https://doi.org/10.31610/zsr/2003.12.2.217" title="New data on taxonomy and distribution of some Fulgoroidea (Homoptera, Cicadina). Zoosystematica Rossica 12 (2): (2003) 217 - 223." url="https://doi.org/10.31610/zsr/2003.12.2.217" year="2004">Gnezdilov et al. (2004)</bibRefCitation>
under
<taxonomicName id="A5CAB15BE9692D7041155E22FFDE14A1" authorityName="Distant" authorityYear="1909" class="Insecta" family="Issidae" genus="Fortunia" kingdom="Animalia" lsidName="Fortunia" order="Hemiptera" pageId="0" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="3B60FED5C9D9EABBC6DC7352A52C0CCB" italics="true" pageId="0" pageNumber="103">Fortunia</emphasis>
</taxonomicName>
Distant, 1909), the type genus of the tribe
<taxonomicName id="85C715C4A6239755B8A7F3C978EFA8C4" authorityName="Cheng &amp; Yang" authorityYear="1991" lsidName="" pageId="0" pageNumber="103" rank="tribe" tribe="Parahiraciini">Parahiraciini</taxonomicName>
and a reminder of the placement of the new genus in this group. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="97A78A7E11C21D541A983D3181B9CD61" pageId="0" pageNumber="103" type="distribution">
<paragraph id="4D32364D06A16E13312BE0880203DB32" pageId="0" pageNumber="103">Distribution.</paragraph>
<paragraph id="56B7C01195894E2F621337862620BB9C" pageId="0" pageNumber="103">Vietnam.</paragraph>
</subSubSection>
</treatment>
</document>