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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="E14FF08CE22E50808701F5640CC643E5" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Pseudotheopea hsingtzungi Lee &amp; Bezděk 2020, sp. nov." docType="treatment" docUuid="B0707A1E-F90D-45C7-B578-33344BB022D3" docUuidSource="ZooBank" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>124</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
<mods:identifier type="ZooBank">4F361E38B2684B70944DD6B6FCC66542</mods:identifier>
<mods:identifier type="Pensoft-UUID">D1FDF644344C5B5B83E6024904E40A05</mods:identifier>
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<treatment ID-GBIF-Taxon="162147247" LSID="urn:lsid:zoobank.org:act:B0707A1E-F90D-45C7-B578-33344BB022D3" httpUri="http://treatment.plazi.org/id/E14FF08CE22E50808701F5640CC643E5" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="E14FF08C-E22E-5080-8701-F5640CC643E5" authority="Lee &amp; Bezděk, 2020" authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi" status="sp. nov.">Pseudotheopea hsingtzungi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">Figs 15A-C</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">; 16A, B</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">; 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Chujo.Solid" authorityYear="1935" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea sauteri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">Theopea sauteri</taxonomicName>
: Medvedev, 2000: 178 (part, misidentification).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="types">
<paragraph pageId="0" pageNumber="65">Types.</paragraph>
<paragraph pageId="0" pageNumber="65">
Holotype ♂ (NHMUK), LAOS. Hua Phan: Ban Saluei, Phou Pane (Mt.),
<geoCoordinate degrees="20" direction="north" minutes="12" orientation="latitude" precision="925" value="20.2">20°12'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="01" orientation="longitude" precision="925" value="104.01667">104°01'E</geoCoordinate>
, 1300-1900 m, 3-30.IV.2014, leg. C. Holzschuh. Paratypes.
<emphasis bold="true" pageId="0" pageNumber="65">LAOS</emphasis>
. Champassak: 1♀ (HNHM), Dong Hua Xao NBCA, bank of Nam Phak river,
<geoCoordinate degrees="15" direction="north" minutes="59" orientation="latitude" precision="925" value="15.983334">15°59'N</geoCoordinate>
<geoCoordinate degrees="105" direction="east" minutes="55" orientation="longitude" precision="925" value="105.916664">105°55'E</geoCoordinate>
, 280 m, 28-29.III.1998, leg. O. Merkl and G. Csorba (identified as
<taxonomicName authorityName="Chujo.Solid" authorityYear="1935" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea sauteri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea sauteri</emphasis>
</taxonomicName>
by
<bibRefCitation author="Medvedev, LN" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="65" pagination="161 - 182" refId="B13" refString="Medvedev, LN, 2000. Chrysomelidae (Coleoptera) of Laos from the collection of the Hungarian Natural History Museum. Annales Historico-Naturales Musei Nationalis Hungarici 92: 161 - 182" title="Chrysomelidae (Coleoptera) of Laos from the collection of the Hungarian Natural History Museum." volume="92" year="2000">Medvedev (2000)</bibRefCitation>
); Hua Pan: 1♀ (JBCB), Ban Kangpabong env., 25km SE Vieng Xai (by road),
<geoCoordinate degrees="20" direction="north" minutes="19" orientation="latitude" precision="925" value="20.316668">20°19'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="25" orientation="longitude" precision="925" value="104.416664">104°25'E</geoCoordinate>
, 14-18.V.2001, leg. J.
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
; 1♀ (NHMB), Phou Pane Mt.,
<geoCoordinate degrees="20" direction="north" minutes="13" orientation="latitude" precision="925" value="20.216667">20°13'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="00" orientation="longitude" precision="925" value="104.0">104°00'E</geoCoordinate>
, 1350-1500 m, 1-16.VI.2009, leg. M. Brancucci; Oudomxai: 1♀ (NHMB), Oudom Xai (17 km NEE),
<geoCoordinate degrees="20" direction="north" minutes="45" orientation="latitude" precision="925" value="20.75">20°45'N</geoCoordinate>
<geoCoordinate degrees="102" direction="east" minutes="09" orientation="longitude" precision="925" value="102.15">102°09'E</geoCoordinate>
, 1100 m, 1-9.V.2002, leg. V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="description">
<paragraph pageId="0" pageNumber="65">Description.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 5.8-6.2 mm, width 2.1-2.4 mm. Body color (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A-C</figureCitation>
) golden green, but mouthparts and legs yellowish brown, antennae dark brown. Frontoclypeus (Figs
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16A-16B</figureCitation>
) with transverse deep groove between eyes in males, concavity 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as interspace between eyes. Antennae filiform in males (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17A</figureCitation>
), antennomere I smaller than others, length ratios of antennomeres I-XI 1.0: 0.3: 1.0: 1.2: 1.2: 1.2: 1.2: 1.1: 1.1: 1.0: 1.1, length to width ratios of antennomeres I-XI 3.3: 1.2: 3.8: 4.8: 5.7: 5.5: 5.5: 5.4: 6.1: 5.6: 6.2; similar but slightly shorter in females (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.8: 0.9: 0.9: 0.9: 0.9: 0.9: 0.9: 0.8: 0.9, length to width ratios of antennomeres I-XI 3.6: 1.7: 4.0: 4.9: 5.3: 5.3: 5.1: 5.4: 5.3: 4.9: 4.7. Elytra elongate, parallel-sided, 1.8-1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one distinct longitudinal ridge between two longitudinal rows of punctures. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17C-E</figureCitation>
) extremely slender, 7.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex with deep notch; tectum elongate, from apical 1/9 to middle; almost straight in lateral view, moderately curved near base, angular at apical 2/5; triangular sclerites elongate; internal sac with elongate, endophallic sclerite complex, 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, composed of two sclerites, apical piece as long as basal piece, one pair of dorsal sclerite hook-like, connected near base of apical piece; ventral sclerites absent. Gonocoxae (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17G</figureCitation>
) elongate, both gonocoxae fused from basal 1/4 to apical 1/4; apices convergent and narrowly rounded, each gonocoxa with eight setae along lateral margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17F</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct shallowly inserted into receptacle, narrow and short.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A-C</figureCitation>
),
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A-C</figureCitation>
),
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
(Medvedev) (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D-F</figureCitation>
), and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Gressitt and Kimoto) (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15D-F</figureCitation>
), are characterized by their golden green coloration. They can be identified based on their distribution:
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. from India,
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
from Vietnam,
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. from Laos, and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
from China.
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A, B</figureCitation>
) is similar to
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A, C</figureCitation>
) and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15D, E</figureCitation>
) based on the shared indistinct longitudinal ridges on the elytra (convex and distinct longitudinal ridges on the elytra in
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D, F</figureCitation>
)), but differs by having the concavity narrower between the eyes in males (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16A, B</figureCitation>
) (concavity wide between eyes in others (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9A, B</figureCitation>
;
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16C, D</figureCitation>
)). Males of
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. are similar to those of
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov. in possessing elongate triangular aedeagal sclerites (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17D, E</figureCitation>
;
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20E, F</figureCitation>
) but differ in the presence of one pair of long hook-like lateral sclerites of the median elongate sclerite, lacking small spines near the apex of the median elongate sclerite, and the median division (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17C, D</figureCitation>
) (lacking hook-like sclerites at sides of median elongate sclerite, with small spines near apex of median elongate sclerite, and undivided in those of
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20D, E</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="etymology">
<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="65">The new species is dedicated to Mr. Hsing-Tzung Cheng, who is a member of the Taiwan Chrysomelid Research Team (TCRT) for inventorying leaf beetles.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">Laos.</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 15.</emphasis>
Habitus of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea hsingtzungi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
, male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
, female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Same, ventral view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 16.</emphasis>
Heads of males of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea hsingtzungi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov., dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, front view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
, dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Same, front view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 17.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea hsingtzungi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea hsingtzungi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>