331 lines
41 KiB
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331 lines
41 KiB
XML
<document id="10E1B62D41CFFF092141642AE845DBC0" ID-DOI="10.1080/00222930802610535" ID-ISSN="1464-5262" ID-Zenodo-Dep="5216179" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629289220664" checkinUser="felipe" docAuthor="Pfeiler, E., Castrezana, S., Reed, L. K. & Markow, T. A." docDate="2009" docId="81628786FFF56D45FEF1FD63CDD2431F" docLanguage="en" docName="JNatHist.43.15-16.923-938.pdf" docOrigin="Journal of Natural History 43 (15 - 16)" docSource="http://dx.doi.org/10.1080/00222930802610535" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Drosophila mojavensis subsp. mojavensis Patterson and Crow 1940" docType="treatment" docVersion="3" lastPageNumber="932" masterDocId="7D5BFFFEFFFD6D49FFAAFF96CE00423F" masterDocTitle="Genetic, ecological and morphological differences among populations of the cactophilic Drosophila mojavensis from southwestern USA and northwestern Mexico, with descriptions of two new subspecies" masterLastPageNumber="938" masterPageNumber="923" pageNumber="929" updateTime="1699066227865" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A4F917AEE00AD55BE0341F8E6AEDC12A">Genetic, ecological and morphological differences among populations of the cactophilic Drosophila mojavensis from southwestern USA and northwestern Mexico, with descriptions of two new subspecies</mods:title>
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<mods:namePart id="C521FE2914ABE7F1262F41F2EA4A1CFC">Pfeiler, E.</mods:namePart>
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<mods:namePart id="FF981766132C5B6F48560BC9A432D039">Castrezana, S.</mods:namePart>
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<mods:namePart id="4C5B25985AA9DB16096CF36EEB2C0D47">Reed, L. K.</mods:namePart>
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<mods:namePart id="43E9B371B910A91510B818986662B0C7">Markow, T. A.</mods:namePart>
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<mods:title id="B8079E21D9F44D59056C2C83741BCB85">Journal of Natural History</mods:title>
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<mods:part id="E17C8A42DAAC90D04A9E2FA751A9E7B8">
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<mods:date id="E274EDDDA24D8D3D244A7C77A9CE3683">2009</mods:date>
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<mods:number id="798FC8584A6E1560A712130851380D6A">2009-04-30</mods:number>
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<mods:detail id="59C9C978DC88B45665DFFD3EB3B150A9" type="volume">
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<mods:number id="4242F37B212CE535EC27C006EBA68D29">43</mods:number>
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<mods:number id="CB39764871F1E50C62F3BCCA85A9916E">15 - 16</mods:number>
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<paragraph id="09743690FFF56D41FEF1FD63CA0A4132" blockId="8.[213,1078,757,880]" box="[347,1034,757,781]" pageId="8" pageNumber="929">
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<heading id="523C81FCFFF56D41FEF1FD63CA0A4132" box="[347,1034,757,781]" centered="true" fontSize="36" level="2" pageId="8" pageNumber="929" reason="7">
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<taxonomicName id="CECB4D13FFF56D41FEF1FD63CA0A4132" authority="mojavensis Patterson and Crow, 1940" authorityName="Patterson and Crow" authorityYear="1940" box="[347,1034,757,781]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="929" phylum="Basidiomycota" rank="subSpecies" species="mojavensis" subSpecies="mojavensis">
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<emphasis id="3BBFEA82FFF56D41FEF1FD63CCD44132" bold="true" box="[347,724,757,781]" italics="true" pageId="8" pageNumber="929">Drosophila mojavensis mojavensis</emphasis>
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<bibRefCitation id="6D5A4B61FFF56D41FD77FD63CA0A4132" author="Patterson JT & Crow JF" box="[733,1034,757,781]" pageId="8" pageNumber="929" pagination="251 - 6" refId="ref8064" refString="Patterson JT, Crow JF. 1940. Hybridization in the mulleri group of Drosophila. Univ Texas Pub. 4032: 251 - 6." type="journal article" year="1940">Patterson and Crow, 1940</bibRefCitation>
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</heading>
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<subSubSection id="41D1651BFFF56D41FDEDFC80CD1E4111" box="[583,798,790,814]" pageId="8" pageNumber="929" type="description">
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<paragraph id="09743690FFF56D41FDEDFC80CD1E4111" blockId="8.[213,1078,757,880]" box="[583,798,790,814]" pageId="8" pageNumber="929">
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(
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<figureCitation id="91F02A15FFF56D41FDE5FC80CCB74111" box="[591,695,790,814]" captionStart="Figure 2" captionStartId="9.[146,213,1098,1120]" captionTargetBox="[305,944,170,1061]" captionTargetId="figure-197@9.[305,944,165,1061]" captionTargetPageId="9" captionText="Figure 2. Lateral view of abdominal pattern in Drosophila m. mojavensis. (A) Male; (B) female." figureDoi="http://doi.org/10.5281/zenodo.5216183" httpUri="https://zenodo.org/record/5216183/files/figure.png" pageId="8" pageNumber="929">Figures 2</figureCitation>
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and
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<figureCitation id="91F02A15FFF56D41FD59FC80CD164111" box="[755,790,790,814]" captionStart="Figure 3" captionStartId="10.[213,280,1285,1307]" captionTargetBox="[435,942,171,1249]" captionTargetId="figure-104@10.[435,947,165,1249]" captionTargetPageId="10" captionText="Figure 3. Photographs showing lateral views of the aedeagus and aedeagal apodeme in the four subspecies of Drosophila mojavensis. (A) D. m. mojavensis; (B) D. m. baja; (C) D. m. sonorensis; (D) D. m. wrigleyi." figureDoi="http://doi.org/10.5281/zenodo.5216185" httpUri="https://zenodo.org/record/5216185/files/figure.png" pageId="8" pageNumber="929">3A</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="41D1651BFFF56D41FF7FFCA1CA36414F" pageId="8" pageNumber="929" type="reference_group">
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<paragraph id="09743690FFF56D41FF7FFCA1CD8A4170" blockId="8.[213,1078,757,880]" box="[213,906,823,847]" pageId="8" pageNumber="929">
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<treatmentCitationGroup id="29DB11BEFFF56D41FF7FFCA1CD8A4170" box="[213,906,823,847]" pageId="8" pageNumber="929">
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<treatmentCitation id="886A1081FFF56D41FF7FFCA1CD8A4170" author="Patterson JT & Crow JF" box="[213,906,823,847]" page="251" pageId="8" pageNumber="929" year="1940">
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<taxonomicName id="CECB4D13FFF56D41FF7FFCA1CD8A4170" authority="Patterson and Crow, 1940: 251" authorityName="Patterson and Crow" authorityPageNumber="251" authorityYear="1940" box="[213,906,823,847]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="929" phylum="Basidiomycota" rank="subSpecies" species="mulleri" subSpecies="mojavensis">
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<emphasis id="3BBFEA82FFF56D41FF7FFCA1CC254170" box="[213,549,823,847]" italics="true" pageId="8" pageNumber="929">Drosophila mulleri mojavensis</emphasis>
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<bibRefCitation id="6D5A4B61FFF56D41FD87FCA1CD8A4170" author="Patterson JT & Crow JF" box="[557,906,823,847]" pageId="8" pageNumber="929" pagination="251 - 6" refId="ref8064" refString="Patterson JT, Crow JF. 1940. Hybridization in the mulleri group of Drosophila. Univ Texas Pub. 4032: 251 - 6." type="journal article" year="1940">Patterson and Crow, 1940:251</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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<paragraph id="09743690FFF56D41FF7FFCCECA36414F" blockId="8.[213,1078,757,880]" box="[213,1078,856,880]" pageId="8" pageNumber="929">
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<treatmentCitationGroup id="29DB11BEFFF56D41FF7FFCCECA36414F" box="[213,1078,856,880]" pageId="8" pageNumber="929">
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<treatmentCitation id="886A1081FFF56D41FF7FFCCECD47414F" author="Patterson JT & Wheeler MR" box="[213,839,856,880]" page="95" pageId="8" pageNumber="929" year="1942">
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<taxonomicName id="CECB4D13FFF56D41FF7FFCCECD47414F" authority="Patterson and Wheeler, 1942: 95" authorityName="Patterson and Wheeler" authorityPageNumber="95" authorityYear="1942" box="[213,839,856,880]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="929" phylum="Basidiomycota" rank="species" species="mojavensis">
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<emphasis id="3BBFEA82FFF56D41FF7FFCCECFCE414F" box="[213,462,856,880]" italics="true" pageId="8" pageNumber="929">Drosophila mojavensis</emphasis>
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<bibRefCitation id="6D5A4B61FFF56D41FE7DFCCECD47414F" author="Patterson JT & Wheeler MR" box="[471,839,856,880]" pageId="8" pageNumber="929" pagination="67 - 109" refId="ref8090" refString="Patterson JT, Wheeler MR. 1942. Description of new species of the subgenera Hirtodrosophila and Drosophila. Univ Texas Pub. 4213: 67 - 109." type="journal article" year="1942">Patterson and Wheeler, 1942:95</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="886A1081FFF56D41FCFFFCCECA36414F" author="Patterson JT" box="[853,1078,856,880]" page="158" pageId="8" pageNumber="929" year="1943">
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<bibRefCitation id="6D5A4B61FFF56D41FCFFFCCECA36414F" author="Patterson JT" box="[853,1078,856,880]" pageId="8" pageNumber="929" pagination="7 - 216" refId="ref8043" refString="Patterson JT. 1943. The Drosophilidae of the Southwest. Univ Texas Pub. 4313: 7 - 216." type="journal article" year="1943">Patterson, 1943:158</bibRefCitation>
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</treatmentCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="41D1651BFFF56D45FF7FFC3FCDD2431F" lastPageId="12" lastPageNumber="933" pageId="8" pageNumber="929" type="diagnosis">
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<paragraph id="09743690FFF56D41FF7FFC3FCF4241FE" blockId="8.[213,322,937,961]" box="[213,322,937,961]" pageId="8" pageNumber="929">
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<emphasis id="3BBFEA82FFF56D41FF7FFC3FCF4241FE" box="[213,322,937,961]" italics="true" pageId="8" pageNumber="929">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="09743690FFF56D41FF7FFC44CF114693" blockId="8.[213,1169,978,1196]" pageId="8" pageNumber="929">
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<emphasis id="3BBFEA82FFF56D41FF7FFC44CFEE41D5" box="[213,494,978,1002]" italics="true" pageId="8" pageNumber="929">
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<taxonomicName id="CECB4D13FFF56D41FF7FFC44CF4F41D5" box="[213,335,978,1002]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="929" phylum="Basidiomycota" rank="genus">Drosophila</taxonomicName>
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m.
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<taxonomicName id="CECB4D13FFF56D41FEDDFC44CFEE41D5" authorityName="Patterson and Wheeler" authorityYear="1942" box="[375,494,978,1002]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="929" phylum="Basidiomycota" rank="species" species="mojavensis">mojavensis</taxonomicName>
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</emphasis>
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can be distinguished from the other subspecies by the shape of the aedeagus (
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<figureCitation id="91F02A15FFF56D41FE18FC64CC284635" box="[434,552,1010,1034]" captionStart="Figure 3" captionStartId="10.[213,280,1285,1307]" captionTargetBox="[435,942,171,1249]" captionTargetId="figure-104@10.[435,947,165,1249]" captionTargetPageId="10" captionText="Figure 3. Photographs showing lateral views of the aedeagus and aedeagal apodeme in the four subspecies of Drosophila mojavensis. (A) D. m. mojavensis; (B) D. m. baja; (C) D. m. sonorensis; (D) D. m. wrigleyi." figureDoi="http://doi.org/10.5281/zenodo.5216185" httpUri="https://zenodo.org/record/5216185/files/figure.png" pageId="8" pageNumber="929">Figure 3A</figureCitation>
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). Internal margin of aedeagus/external margin of aedeagal apodeme index51.0 (0.99–1.01;
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<emphasis id="3BBFEA82FFF56D41FD16FB85CCC94614" box="[700,713,1043,1067]" italics="true" pageId="8" pageNumber="929">n</emphasis>
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530); ventral margin of aedeagus/wide aedeagal apodeme index53.7 (3.65–3.92). Tip of aedeagus with a small pointy protuberance; dorsal and ventral aedeagus margins almost form an isosceles triangle. Aedeagus ventral margin almost straight with a small protuberance in the middle zone.
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</paragraph>
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<paragraph id="09743690FFF56D41FF7FFB73CF6C46C2" blockId="8.[213,364,1253,1277]" box="[213,364,1253,1277]" pageId="8" pageNumber="929">
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<emphasis id="3BBFEA82FFF56D41FF7FFB73CF6C46C2" box="[213,364,1253,1277]" italics="true" pageId="8" pageNumber="929">Redescription</emphasis>
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</paragraph>
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<paragraph id="09743690FFF56D42FF7FFA98CCEA405A" blockId="8.[213,1169,1293,1706]" lastBlockId="11.[146,1103,168,613]" lastPageId="11" lastPageNumber="932" pageId="8" pageNumber="929">
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<emphasis id="3BBFEA82FFF56D41FF7FFA98CF0F4719" box="[213,271,1294,1318]" italics="true" pageId="8" pageNumber="929">Male</emphasis>
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.
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<emphasis id="3BBFEA82FFF56D41FEB7FA98CF594719" box="[285,345,1294,1318]" italics="true" pageId="8" pageNumber="929">Head</emphasis>
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. Front brownish; frontal length 0.32 (0.30–0.34) mm; frontal index50.83 (0.76–0.88). Frontal triangle pale brown, more or less distinct. Ocellar triangle slightly prominent and lighter than front, pollinose. Interfrontal setulae in ‘‘V’’ shape. Frontal vittae brownish; orbital plates wider at the or1 level, almost a pointed shape. Orbital setae black, or2 just slightly posterior to or1; or1/or3 ratio50.90 (0.76–1.07); or2/or1 ratio50.60 (0.56–0.68); vibrissal index50.51 (0.41–0.61). Face brownish. Carina broad below and sulcate, central area similar colour as vittae; distal lateral sections light brown. Cheek index ca. 5.70–9.86. Eye red; eye index51.17 (1.11–1.25). Occiput yellowish, brown above foramen. Antennae tannish brown, pedicel slightly darker, third joint darker; arista with three dorsal, two ventral, and about four small inner branches, plus terminal fork. Proboscis yellowish, clypeus brownish. Palpus pale yellow with about three strong setae along external ventral margin.
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<emphasis id="3BBFEA82FFF56D41FE55F904CC524495" box="[511,594,1682,1706]" italics="true" pageId="8" pageNumber="929">Thorax</emphasis>
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. Length: 1.01 (0.98–1.04)mm. Mesonotum light brownish-yellow; setae arising from brown spots; eight rows of acrostichal setulae. Upper/lower postpronotal setae (h index)51.20 (1.19–1.20); anterior/posterior dorsocentral setae (dc index)50.63 (0.60–0.65). Scutellum dark brownish with margin slightly clear. Basal setae slightly convergent, apical scutellar convergent; basal/apical scutellar setae (scut index)50.82 (0.73–0.90). Pleura brownish, shining, with a narrow, dark brown stripe from upper margin of katepisternum to below procoxa. Faint darker brown stripes, one visible along upper margin, and a second stripe on the middle part of anepisternum; this strip prolonged faintly to the anepimeron. Anterior/posterior katepisternal setae (sterno index)50.72 (0.64–0.80), median katepisternal seta about 52–61% of the anterior one. Haltere yellow with slightly brownish colouration on anterior side of knob. Legs yellow, with a faint brownish postbasal band on tibia which it is darker on hind leg. Procoxa and all fifth tarsal segments slightly darker than the rest of the legs; preapical setae on all tibiae, apical seta on mesotibia.
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<emphasis id="3BBFEA82FFF46D40FE1FF9C4CC1D4455" box="[437,541,1618,1642]" italics="true" pageId="9" pageNumber="930">Abdomen</emphasis>
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(
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<figureCitation id="91F02A15FFF46D40FD87F9C4CCA44455" box="[557,676,1618,1642]" captionStart="Figure 2" captionStartId="9.[146,213,1098,1120]" captionTargetBox="[305,944,170,1061]" captionTargetId="figure-197@9.[305,944,165,1061]" captionTargetPageId="9" captionText="Figure 2. Lateral view of abdominal pattern in Drosophila m. mojavensis. (A) Male; (B) female." figureDoi="http://doi.org/10.5281/zenodo.5216183" httpUri="https://zenodo.org/record/5216183/files/figure.png" pageId="9" pageNumber="930">Figure 2A</figureCitation>
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). Pale yellow; apical band on tergites well defined in dorsal area with interruption between sides. On the upper margin of the lateral area, apical band curves slightly and expands as a diffuse extension to almost touch the margin of the anterior tergite. Background colour in lateral area brownish-yellow, diffuse; most of the margin in lateral areas from tergites 1–4 covered by a dark irregular spot; colour of lateral area spots lighter than apical band; colour gradually disappears from ventral to dorsal area; spot almost completely disappears at posterior margin of tergite 5. Testes yellow with 2.5 inner and 3 outer coils.
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<emphasis id="3BBFEA82FFF76D43FEBDF9C6CF5D4457" box="[279,349,1616,1640]" italics="true" pageId="10" pageNumber="931">Wings</emphasis>
|
||
. Hyaline, veins brown. Apex of subcostal break slightly black with two well-developed setae. Third costal section with heavy bristles on its basal third. Crossveins clear. Wing length 1.70 (1.60–1.80)mm, length to width ratio51.82 (1.73–1.88). Indices: Costa (C index)52.91 (2.69–3.06), C-III/C-IV (ac index)52.27 (2.00–2.57), C-III/M-III (4C index)50.92 (0.80–1.05), M-IV/M-III (4v index)51.81 (1.67–1.91), CuA (apical section)/dM-Cu (5x index)51.26 (1.00–1.44), CuA (apical section)/M-III (M index)50.48 (0.43–0.53), Basal R
|
||
<subScript id="954F34D5FFF66D42FD4CFE82CD05431B" attach="right" box="[742,773,275,292]" fontSize="7" pageId="11" pageNumber="932">4+5</subScript>
|
||
/M-III (prox. X index)50.67 (0.64–0.71).
|
||
<emphasis id="3BBFEA82FFF66D42FE89FEBCCF88437D" box="[291,392,298,322]" italics="true" pageId="11" pageNumber="932">Genitalia</emphasis>
|
||
. Cerci with few microtrichose, many long setae, and fused anteriorly to epandrium. Epandrium barely microtrichose on dorsal area; three to five setae on the apical margin in the medial section; ventral lobe enlarged and bare, with filament-like setae, forming a rounded toe that partially overlaps surstylus laterally. Surstylus not microtrichose; 9–11 peg-like prensisetae almost rounded on tip, with the two anterior setae slightly larger and pointed; three to four inner setae larger than prensisetae; seven to eight outer setae. Cerci barely microtrichose; hypandrium slightly longer than epandrium. Aedeagus fused to aedeagal apodeme (
|
||
<figureCitation id="91F02A15FFF66D42FF30FDBBCF10407A" box="[154,272,557,581]" captionStart="Figure 3" captionStartId="10.[213,280,1285,1307]" captionTargetBox="[435,942,171,1249]" captionTargetId="figure-104@10.[435,947,165,1249]" captionTargetPageId="10" captionText="Figure 3. Photographs showing lateral views of the aedeagus and aedeagal apodeme in the four subspecies of Drosophila mojavensis. (A) D. m. mojavensis; (B) D. m. baja; (C) D. m. sonorensis; (D) D. m. wrigleyi." figureDoi="http://doi.org/10.5281/zenodo.5216185" httpUri="https://zenodo.org/record/5216185/files/figure.png" pageId="11" pageNumber="932">Figure 3A</figureCitation>
|
||
). Aedeagal apodeme upper half smaller than aedeagus. Ventral rod almost twice as long as width of aedeagal apodeme.
|
||
</paragraph>
|
||
<caption id="5DB46618FFF46D40FF38FBDCCEDD4642" ID-DOI="http://doi.org/10.5281/zenodo.5216183" ID-Zenodo-Dep="5216183" httpUri="https://zenodo.org/record/5216183/files/figure.png" pageId="9" pageNumber="930" startId="9.[146,213,1098,1120]" targetBox="[305,944,170,1061]" targetPageId="9">
|
||
<paragraph id="09743690FFF46D40FF38FBDCCEDD4642" blockId="9.[146,1102,1098,1149]" pageId="9" pageNumber="930">
|
||
Figure 2. Lateral view of abdominal pattern in
|
||
<emphasis id="3BBFEA82FFF46D40FD35FBDCCDA74660" box="[671,935,1098,1119]" italics="true" pageId="9" pageNumber="930">
|
||
<taxonomicName id="CECB4D13FFF46D40FD35FBDCCD0D4660" box="[671,781,1098,1119]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="930" phylum="Basidiomycota" rank="genus">Drosophila</taxonomicName>
|
||
m.
|
||
<taxonomicName id="CECB4D13FFF46D40FC97FBDCCDA74660" authorityName="Patterson and Wheeler" authorityYear="1942" box="[829,935,1098,1119]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="930" phylum="Basidiomycota" rank="species" species="mojavensis">mojavensis</taxonomicName>
|
||
</emphasis>
|
||
. (A) Male; (B) female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5DB46618FFF76D43FF7FFA93CC0B4769" ID-DOI="http://doi.org/10.5281/zenodo.5216185" ID-Zenodo-Dep="5216185" httpUri="https://zenodo.org/record/5216185/files/figure.png" pageId="10" pageNumber="931" startId="10.[213,280,1285,1307]" targetBox="[435,942,171,1249]" targetPageId="10">
|
||
<paragraph id="09743690FFF76D43FF7FFA93CC0B4769" blockId="10.[213,1169,1285,1366]" pageId="10" pageNumber="931">
|
||
Figure 3. Photographs showing lateral views of the aedeagus and aedeagal apodeme in the four subspecies of
|
||
<taxonomicName id="CECB4D13FFF76D43FE37FAB5CC7F4707" authorityName="Patterson and Wheeler" authorityYear="1942" box="[413,639,1315,1336]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="931" phylum="Basidiomycota" rank="species" species="mojavensis">
|
||
<emphasis id="3BBFEA82FFF76D43FE37FAB5CC7F4707" box="[413,639,1315,1336]" italics="true" pageId="10" pageNumber="931">Drosophila mojavensis</emphasis>
|
||
</taxonomicName>
|
||
. (A)
|
||
<emphasis id="3BBFEA82FFF76D43FD17FAB5CD6D4707" box="[701,877,1315,1336]" italics="true" pageId="10" pageNumber="931">
|
||
D. m.
|
||
<taxonomicName id="CECB4D13FFF76D43FCA9FAB5CD6D4707" authorityName="Patterson and Wheeler" authorityYear="1942" box="[771,877,1315,1336]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="931" phylum="Basidiomycota" rank="species" species="mojavensis">mojavensis</taxonomicName>
|
||
</emphasis>
|
||
; (B)
|
||
<emphasis id="3BBFEA82FFF76D43FC02FAB5CA194707" box="[936,1049,1315,1336]" italics="true" pageId="10" pageNumber="931">D. m. baja</emphasis>
|
||
; (C)
|
||
<emphasis id="3BBFEA82FFF76D43FBFFFAB5CF3B476A" italics="true" pageId="10" pageNumber="931">D. m. sonorensis</emphasis>
|
||
; (D)
|
||
<emphasis id="3BBFEA82FFF76D43FEDEFAD6CC05476A" box="[372,517,1344,1365]" italics="true" pageId="10" pageNumber="931">D. m. wrigleyi</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="09743690FFF66D42FF38FD08CF2141E6" blockId="11.[146,1103,670,985]" pageId="11" pageNumber="932">
|
||
<emphasis id="3BBFEA82FFF66D42FF38FD08CEE24089" box="[146,226,670,694]" italics="true" pageId="11" pageNumber="932">Female</emphasis>
|
||
. Identical to male, except as follows. Abdomen similar to male except lateral light area on tergite 5 diffuse, darker than male (
|
||
<figureCitation id="91F02A15FFF66D42FD1EFD28CD2640E9" box="[692,806,702,726]" captionStart="Figure 2" captionStartId="9.[146,213,1098,1120]" captionTargetBox="[305,944,170,1061]" captionTargetId="figure-197@9.[305,944,165,1061]" captionTargetPageId="9" captionText="Figure 2. Lateral view of abdominal pattern in Drosophila m. mojavensis. (A) Male; (B) female." figureDoi="http://doi.org/10.5281/zenodo.5216183" httpUri="https://zenodo.org/record/5216183/files/figure.png" pageId="11" pageNumber="932">Figure 2B</figureCitation>
|
||
).
|
||
<emphasis id="3BBFEA82FFF66D42FC90FD28CD8040E9" box="[826,896,702,726]" italics="true" pageId="11" pageNumber="932">Wings</emphasis>
|
||
. Wing length 1.78 (1.62–1.87)mm, length to width ratio52.01(1.93–2.05). Indices: Costa53.20 (2.45– 3.43), C-III/C-IV52.34 (2.13–2.57), C-III/M-III50.86 (0.77–1.03), M-IV/M- III51.64 (1.51–1.74), CuA (apical section)/dM-Cu51.19 (1.00–1.38), CuA (apical section)/M-III50.52 (0.46–0.58), Basal R
|
||
<subScript id="954F34D5FFF66D42FD23FCDCCCA84165" attach="left" box="[649,680,841,858]" fontSize="7" pageId="11" pageNumber="932">4+5</subScript>
|
||
/M-III50.80 (0.70–1.00).
|
||
<emphasis id="3BBFEA82FFF66D42FC48FCD6CA474167" box="[994,1095,832,856]" italics="true" pageId="11" pageNumber="932">Genitalia</emphasis>
|
||
. Valve of oviscapt distally rounded, ventrally almost straight, with ca. 6 distal and 12–13 marginal, peg-like, mostly roundish-tipped, outer ovisensi1la; inner ovisensi1la: four thin, trichoid-like, distally positioned, and one long, straight, subterminal.
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D42FF38FB84CFCD4615" blockId="11.[146,461,1042,1066]" box="[146,461,1042,1066]" pageId="11" pageNumber="932">
|
||
<emphasis id="3BBFEA82FFF66D42FF38FB84CFCD4615" box="[146,461,1042,1066]" italics="true" pageId="11" pageNumber="932">Distribution and host cactus</emphasis>
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D42FF38FBADCC50464C" blockId="11.[146,1102,1083,1139]" pageId="11" pageNumber="932">
|
||
Mojave Desert (southern
|
||
<collectingRegion id="CB0FF872FFF66D42FE19FBADCC27466C" box="[435,551,1083,1107]" country="United States of America" name="California" pageId="11" pageNumber="932">California</collectingRegion>
|
||
to northwestern
|
||
<collectingRegion id="CB0FF872FFF66D42FD42FBADCD47466C" box="[744,839,1083,1107]" country="United States of America" name="Arizona" pageId="11" pageNumber="932">Arizona</collectingRegion>
|
||
;
|
||
<figureCitation id="91F02A15FFF66D42FCF8FBADCDAE466C" box="[850,942,1083,1107]" captionStart="Figure 1" captionStartId="3.[146,213,1075,1097]" captionTargetBox="[148,1101,165,1039]" captionTargetId="figure-162@3.[148,1101,165,1039]" captionTargetPageId="3" captionText="Figure 1. Map showing approximate geographic distribution of the four subspecies of Drosophila mojavensis in southwestern USA and northwestern Mexico.?5unconfirmed subspecies at San Felipe, Baja California. Numbers show localities where flies used for laboratory cultures were collected: (1) Mojave; (2) Baja; (3) Sonora; (4) Catalina (see ‘Materials and methods’ for details)." figureDoi="http://doi.org/10.5281/zenodo.5216181" httpUri="https://zenodo.org/record/5216181/files/figure.png" pageId="11" pageNumber="932">Figure 1</figureCitation>
|
||
).
|
||
<emphasis id="3BBFEA82FFF66D42FC6FFBADCA47466C" box="[965,1095,1083,1107]" italics="true" pageId="11" pageNumber="932">Host cactus</emphasis>
|
||
: barrel cactus (
|
||
<taxonomicName id="CECB4D13FFF66D42FE9DFBCDCC40464C" box="[311,576,1115,1139]" class="Magnoliopsida" family="Cactaceae" genus="Ferocactus" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="932" phylum="Tracheophyta" rank="species" species="cylindraceus">
|
||
<emphasis id="3BBFEA82FFF66D42FE9DFBCDCC40464C" box="[311,576,1115,1139]" italics="true" pageId="11" pageNumber="932">Ferocactus cylindraceus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D42FF38FB3ACEF546FB" blockId="11.[146,245,1196,1220]" box="[146,245,1196,1220]" pageId="11" pageNumber="932">
|
||
<emphasis id="3BBFEA82FFF66D42FF38FB3ACEF546FB" box="[146,245,1196,1220]" italics="true" pageId="11" pageNumber="932">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D42FF38FB43CF944790" blockId="11.[146,1103,1237,1455]" pageId="11" pageNumber="932">
|
||
Referring to many newly described species of
|
||
<taxonomicName id="CECB4D13FFF66D42FD3FFB43CD0F46D2" box="[661,783,1237,1261]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="11" pageNumber="932" phylum="Basidiomycota" rank="genus">
|
||
<emphasis id="3BBFEA82FFF66D42FD3FFB43CD0F46D2" box="[661,783,1237,1261]" italics="true" pageId="11" pageNumber="932">Drosophila</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<emphasis id="3BBFEA82FFF66D42FC26FB43CDCA46D2" box="[908,970,1237,1261]" italics="true" pageId="11" pageNumber="932">D. m.</emphasis>
|
||
<taxonomicName id="CECB4D13FFF66D42FC7BFB43CF524732" authority=", Patterson (1943)" authorityName=", Patterson" authorityYear="1943" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="11" pageNumber="932" phylum="Basidiomycota" rank="species" species="mojavensis">
|
||
<emphasis id="3BBFEA82FFF66D42FC7BFB43CA4846D2" box="[977,1096,1237,1261]" italics="true" pageId="11" pageNumber="932">mojavensis</emphasis>
|
||
,
|
||
<bibRefCitation id="6D5A4B61FFF66D42FF38FB63CF524732" author="Patterson JT" box="[146,338,1269,1293]" pageId="11" pageNumber="932" pagination="7 - 216" refId="ref8043" refString="Patterson JT. 1943. The Drosophilidae of the Southwest. Univ Texas Pub. 4313: 7 - 216." type="journal article" year="1943">Patterson (1943)</bibRefCitation>
|
||
</taxonomicName>
|
||
commented that type specimens are widely scattered in private collections and museums. Whether a
|
||
<typeStatus id="D6708832FFF66D42FD97FA83CCA14712" box="[573,673,1301,1325]" pageId="11" pageNumber="932" type="holotype">holotype</typeStatus>
|
||
was designated is not stated in either
|
||
<bibRefCitation id="6D5A4B61FFF66D42FF38FAA0CFC84771" author="Patterson JT & Crow JF" box="[146,456,1334,1358]" pageId="11" pageNumber="932" pagination="251 - 6" refId="ref8064" refString="Patterson JT, Crow JF. 1940. Hybridization in the mulleri group of Drosophila. Univ Texas Pub. 4032: 251 - 6." type="journal article" year="1940">Patterson and Crow (1940)</bibRefCitation>
|
||
or
|
||
<bibRefCitation id="6D5A4B61FFF66D42FE5BFAA0CCB04771" author="Patterson JT" box="[497,688,1334,1358]" pageId="11" pageNumber="932" pagination="7 - 216" refId="ref8043" refString="Patterson JT. 1943. The Drosophilidae of the Southwest. Univ Texas Pub. 4313: 7 - 216." type="journal article" year="1943">Patterson (1943)</bibRefCitation>
|
||
, but a ‘‘cotype’’ (either a
|
||
<typeStatus id="D6708832FFF66D42FC7DFAA0CA2E4771" box="[983,1070,1334,1358]" pageId="11" pageNumber="932" type="syntype">syntype</typeStatus>
|
||
or
|
||
<typeStatus id="D6708832FFF66D42FF38FAC0CEF74751" box="[146,247,1366,1390]" pageId="11" pageNumber="932" type="paratype">paratype</typeStatus>
|
||
) of
|
||
<emphasis id="3BBFEA82FFF66D42FE81FAC0CFEE4751" box="[299,494,1366,1390]" italics="true" pageId="11" pageNumber="932">
|
||
D. m.
|
||
<taxonomicName id="CECB4D13FFF66D42FEDDFAC0CFEE4751" authorityName=", Patterson" authorityYear="1943" box="[375,494,1366,1390]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="11" pageNumber="932" phylum="Basidiomycota" rank="species" species="mojavensis">mojavensis</taxonomicName>
|
||
</emphasis>
|
||
from Mesquite Springs, Death Valley,
|
||
<collectingRegion id="CB0FF872FFF66D42FC15FAC0CA334751" box="[959,1075,1366,1390]" country="United States of America" name="California" pageId="11" pageNumber="932">California</collectingRegion>
|
||
is deposited in the American Museum of Natural History,
|
||
<collectingRegion id="CB0FF872FFF66D42FC9DFAE0CDB747B1" box="[823,951,1398,1422]" country="United States of America" name="New York" pageId="11" pageNumber="932">New York</collectingRegion>
|
||
. We did not examine the ‘‘cotype’’.
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D42FF38FA7ECF67443F" blockId="11.[146,359,1512,1536]" box="[146,359,1512,1536]" pageId="11" pageNumber="932">
|
||
<emphasis id="3BBFEA82FFF66D42FF38FA7ECF67443F" box="[146,359,1512,1536]" italics="true" pageId="11" pageNumber="932">Material examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="09743690FFF66D45FF38F987CDD2431F" blockId="11.[146,1103,1553,1706]" lastBlockId="12.[213,1169,168,288]" lastPageId="12" lastPageNumber="933" pageId="11" pageNumber="932">
|
||
External and internal (genitalia) measurements were conducted on
|
||
<materialsCitation id="B9A33CCDFFF66D42FC3FF987CC7B4476" collectionCode="ANZA" pageId="11" pageNumber="932" specimenCode="ANZA406" specimenCount="42" specimenCount-female="12" specimenCount-male="30">
|
||
<specimenCount id="1FCDFD19FFF66D42FC3FF987CDF84416" box="[917,1016,1553,1577]" pageId="11" pageNumber="932" type="male">30 males</specimenCount>
|
||
and
|
||
<specimenCount id="1FCDFD19FFF66D42FB9EF987CEE74476" pageId="11" pageNumber="932" type="female">12 females</specimenCount>
|
||
from laboratory culture
|
||
<specimenCode id="596D9EEBFFF66D42FDA8F9A7CC7B4476" box="[514,635,1585,1609]" collectionCode="ANZA" pageId="11" pageNumber="932">ANZA406</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="B9A33CCDFFF66D42FD2BF9A7CFA944B5" collectingDate="2006-04" collectionCode="L, K" country="United States of America" county="San Diego County" location="Anza-Borrego Desert State Park" municipality="Borrego Springs" pageId="11" pageNumber="932" specimenCount="1" stateProvince="California">
|
||
started with flies from
|
||
<collectingRegion id="CB0FF872FFF66D42FC28F9A7CDF64476" box="[898,1014,1585,1609]" country="United States of America" name="California" pageId="11" pageNumber="932">California</collectingRegion>
|
||
(
|
||
<collectingCountry id="71DC7600FFF66D42FBAFF9A7CA424476" box="[1029,1090,1585,1609]" name="United States of America" pageId="11" pageNumber="932">USA</collectingCountry>
|
||
), near
|
||
<collectingMunicipality id="E910ACEAFFF66D42FF64F9C7CF8D4456" box="[206,397,1617,1641]" pageId="11" pageNumber="932">Borrego Springs</collectingMunicipality>
|
||
,
|
||
<location id="0C14604BFFF66D42FE36F9C7CD194456" LSID="urn:lsid:plazi:treatment:81628786FFF56D45FEF1FD63CDD2431F:0C14604BFFF66D42FE36F9C7CD194456" box="[412,793,1617,1641]" country="United States of America" county="San Diego County" municipality="Borrego Springs" name="Anza-Borrego Desert State Park" pageId="11" pageNumber="932" stateProvince="California">Anza-Borrego Desert State Park</location>
|
||
,
|
||
<collectingCounty id="E0154E1CFFF66D42FC82F9C7CA044456" box="[808,1028,1617,1641]" pageId="11" pageNumber="932">San Diego County</collectingCounty>
|
||
,
|
||
<date id="7D751050FFF66D42FBB9F9C7CECA44B5" pageId="11" pageNumber="932" value="2006-04">
|
||
<collectingDate id="6D31E9B8FFF66D42FBB9F9C7CECA44B5" pageId="11" pageNumber="932" value="2006-04">April 2006</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectionCode id="6FDAAE55FFF66D42FEDDF9E4CF8C44B5" box="[375,396,1650,1674]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="11" pageNumber="932" type="Herbarium">L</collectionCode>
|
||
.
|
||
<collectionCode id="6FDAAE55FFF66D42FE3AF9E4CFA944B5" box="[400,425,1650,1674]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="11" pageNumber="932" type="Herbarium">K</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B9A33CCDFFF66D42FE11F9E4CC6C44B5" box="[443,620,1650,1674]" collectionCode="T, A" country="United States of America" location="Reed" pageId="11" pageNumber="932" specimenCount="1" stateProvince="California">
|
||
<location id="0C14604BFFF66D42FE11F9E4CFF544B5" LSID="urn:lsid:plazi:treatment:81628786FFF56D45FEF1FD63CDD2431F:0C14604BFFF66D42FE11F9E4CFF544B5" box="[443,501,1650,1674]" country="United States of America" name="Reed" pageId="11" pageNumber="932" stateProvince="California">Reed</location>
|
||
and
|
||
<collectionCode id="6FDAAE55FFF66D42FD96F9E4CC5044B5" box="[572,592,1650,1674]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="932">T</collectionCode>
|
||
.
|
||
<collectionCode id="6FDAAE55FFF66D42FDFEF9E4CC6C44B5" box="[596,620,1650,1674]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="11" pageNumber="932" type="Herbarium">A</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B9A33CCDFFF66D42FDD5F9E4CC2E4495" collectionCode="ANZA" country="United States of America" pageId="11" pageNumber="932" specimenCode="ANZA406-4" specimenCount="1" stateProvince="California">
|
||
Markow. Voucher specimens (all from isofemale line culture
|
||
<specimenCode id="596D9EEBFFF66D42FE38F904CC264495" box="[402,550,1682,1706]" collectionCode="ANZA" pageId="11" pageNumber="932">ANZA406-4</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="B9A33CCDFFF66D45FD91F904CA6742DF" collectorName="San Diego & University & La Jolla" country="United States of America" lastPageId="12" lastPageNumber="933" location="California" pageId="11" pageNumber="932" specimenCount="22" specimenCount-female="12" specimenCount-male="10" stateProvince="California">
|
||
<specimenCount id="1FCDFD19FFF66D42FD91F904CC9F4495" box="[571,671,1682,1706]" pageId="11" pageNumber="932" type="male">10 males</specimenCount>
|
||
and
|
||
<specimenCount id="1FCDFD19FFF66D42FD74F904CD574495" box="[734,855,1682,1706]" pageId="11" pageNumber="932" type="female">12 females</specimenCount>
|
||
deposited at the
|
||
<collectorName id="A43E5346FFF66D45FB8EF904CF1A42FF" lastPageId="12" lastPageNumber="933" pageId="11" pageNumber="932">San Diego</collectorName>
|
||
<taxonomicName id="CECB4D13FFF16D45FE8AFF3ECC3442FF" authority="Stock Center" authorityName="Stock Center" box="[288,564,168,192]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="12" pageNumber="933" phylum="Basidiomycota" rank="genus">
|
||
<emphasis id="3BBFEA82FFF16D45FE8AFF3ECF9A42FF" box="[288,410,168,192]" italics="true" pageId="12" pageNumber="933">Drosophila</emphasis>
|
||
Stock Center
|
||
</taxonomicName>
|
||
collection at the
|
||
<collectorName id="A43E5346FFF16D45FD5CFF3ECD6D42FF" box="[758,877,168,192]" pageId="12" pageNumber="933">University</collectorName>
|
||
of
|
||
<collectingRegion id="CB0FF872FFF16D45FC38FF3ECA0942FF" box="[914,1033,168,192]" country="United States of America" name="California" pageId="12" pageNumber="933">California</collectingRegion>
|
||
,
|
||
<collectorName id="A43E5346FFF16D45FBBEFF3ECA8D42FF" box="[1044,1165,168,192]" pageId="12" pageNumber="933">San Diego</collectorName>
|
||
,
|
||
<collectorName id="A43E5346FFF16D45FF7FFF5ECF3842DF" box="[213,312,200,224]" pageId="12" pageNumber="933">La Jolla</collectorName>
|
||
,
|
||
<collectingRegion id="CB0FF872FFF16D45FEEDFF5ECFBB42DF" box="[327,443,200,224]" country="United States of America" name="California" pageId="12" pageNumber="933">California</collectingRegion>
|
||
(acquisition nos. 461–470 (males) and 471–482 (females))
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B9A33CCDFFF16D45FBDCFF5ECDCE431F" collectionCode="USNM" country="United States of America" location="National Museum of Natural History" municipality="Smithsonian Diptera Collection" pageId="12" pageNumber="933" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Washington">
|
||
<specimenCount id="1FCDFD19FFF16D45FBDCFF5ECF15433F" pageId="12" pageNumber="933" type="male">10 males</specimenCount>
|
||
and
|
||
<specimenCount id="1FCDFD19FFF16D45FEE6FF7ECFC3433F" box="[332,451,232,256]" pageId="12" pageNumber="933" type="female">10 females</specimenCount>
|
||
deposited at the
|
||
<collectingMunicipality id="E910ACEAFFF16D45FD28FF7ECDED433F" box="[642,1005,232,256]" pageId="12" pageNumber="933">Smithsonian Diptera Collection</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="71DC7600FFF16D45FC52FF7ECA90433F" box="[1016,1168,232,256]" name="United States of America" pageId="12" pageNumber="933">United States</collectingCountry>
|
||
<location id="0C14604BFFF16D45FF7FFE9ECC88431F" LSID="urn:lsid:plazi:treatment:81628786FFF56D45FEF1FD63CDD2431F:0C14604BFFF16D45FF7FFE9ECC88431F" box="[213,648,264,288]" country="United States of America" municipality="Smithsonian Diptera Collection" name="National Museum of Natural History" pageId="12" pageNumber="933" stateProvince="Washington">National Museum of Natural History</location>
|
||
(
|
||
<collectionCode id="6FDAAE55FFF16D45FD33FE9ECCF2431F" box="[665,754,264,288]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="933" type="Museum">USNM</collectionCode>
|
||
),
|
||
<collectingRegion id="CB0FF872FFF16D45FCA2FE9ECD95431F" box="[776,917,264,288]" country="United States of America" name="Washington" pageId="12" pageNumber="933">Washington</collectingRegion>
|
||
, DC
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |