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<document id="C8E21B5E58724D77313E731A43DF309D" ID-DOI="10.1111/zoj.12362" ID-ISSN="0024-4082" ID-Zenodo-Dep="10109299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235092894" checkinUser="marcus" docAuthor="Dellapé, Pablo M., Melo, María C. &amp; Henry, Thomas J." docDate="2016" docId="69250127E932505AFEAD33D0FB31EDD1" docLanguage="en" docName="zoj12362.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (1)" docSource="http://dx.doi.org/10.1111/zoj.12362" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus costalis Dellapé &amp; Melo &amp; Henry 2016, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="47" masterDocId="951C795FE9235048FF9E3541FFCCEE5A" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="46" updateTime="1700019240479" updateUser="ExternalLinkService">
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<mods:title id="A000874FC02DC0EC345C9E63F1948DC7">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="2FE924987A32353EC48B47A102EEE935">Dellapé, Pablo M.</mods:namePart>
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<mods:namePart id="77E81E69BCA1010B8A653DD24021A8AF">Melo, María C.</mods:namePart>
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<mods:namePart id="15109FCC6196588D8AF4CBD60613E348">Henry, Thomas J.</mods:namePart>
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<mods:title id="319F4E404712D6D4B71F2E67832903F0">Zoological Journal of the Linnean Society</mods:title>
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<treatment id="69250127E932505AFEAD33D0FB31EDD1" LSID="urn:lsid:plazi:treatment:69250127E932505AFEAD33D0FB31EDD1" httpUri="http://treatment.plazi.org/id/69250127E932505AFEAD33D0FB31EDD1" lastPageId="18" lastPageNumber="47" pageId="17" pageNumber="46">
<subSubSection id="A996E3BAE9325059FEAD33D0FDB0E8FF" box="[307,636,1680,1702]" pageId="17" pageNumber="46" type="nomenclature">
<paragraph id="E133B031E9325059FEAD33D0FDB0E8FF" blockId="17.[307,636,1680,1702]" box="[307,636,1680,1702]" pageId="17" pageNumber="46">
<heading id="BA7B075DE9325059FEAD33D0FDB0E8FF" box="[307,636,1680,1702]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="46" reason="4">
<taxonomicName id="268CCBB2E9325059FEAD33D0FDD5E8FF" ID-CoL="3KXM4" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[307,537,1681,1702]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="costalis" status="sp. nov.">HERAEUS COSTALIS</taxonomicName>
<emphasis id="D3F86C23E9325059FDBE33D5FDB0E8FF" bold="true" box="[544,636,1680,1701]" pageId="17" pageNumber="46">
<taxonomicNameLabel id="C8CBD158E9325059FDBE33D5FDB0E8FF" box="[544,636,1680,1701]" pageId="17" pageNumber="46" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="A996E3BAE932505AFEA133F6FB31EDD1" lastPageId="18" lastPageNumber="47" pageId="17" pageNumber="46" type="description">
<paragraph id="E133B031E9325059FEA133F6FDBCE897" blockId="17.[319,624,1719,1741]" box="[319,624,1719,1741]" pageId="17" pageNumber="46">
(
<figureCitation id="79B7ACB4E9325059FED933F6FE6AE896" box="[327,422,1719,1740]" captionStart="Figure 3" captionStartId="12.[144,224,1458,1477]" captionTargetBox="[145,1424,195,1428]" captionTargetId="figure-137@12.[144,1424,195,1429]" captionTargetPageId="12" captionText="Figure 3. Habitus dorsal: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus guttatus (Dallas, 1852). G, Heraeus hollyae Baranowski, 2005. H, Heraeus triguttatus (Guérin-Méneville, 1857); I, Heraeus steineri sp. nov.; J, Heraeus caliginosus Slater &amp; Baranowski, 1994; K, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128581" httpUri="https://zenodo.org/record/10128581/files/figure.png" pageId="17" pageNumber="46">FIGS 3C</figureCitation>
,
<figureCitation id="79B7ACB4E9325059FE2D33F6FE1AE897" box="[435,470,1719,1741]" captionStart="Figure 5" captionStartId="14.[144,222,1347,1366]" captionTargetBox="[305,1261,197,1315]" captionTargetId="figure-232@14.[303,1263,195,1317]" captionTargetPageId="14" captionText="Figure 5. Legs: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus caliginosus Slater &amp; Baranowski, 1994; G, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128585" httpUri="https://zenodo.org/record/10128585/files/figure.png" pageId="17" pageNumber="46">5C</figureCitation>
,
<figureCitation id="79B7ACB4E9325059FE7D33F6FDD1E897" box="[483,541,1719,1741]" captionStart="Figure 6" captionStartId="15.[164,243,1703,1722]" captionTargetBox="[243,1361,199,1673]" captionTargetId="figure-0@15.[234,1372,192,1684]" captionTargetPageId="15" captionText="Figure 6. Male genitalia. Heraeus cinnamomeus Barber, 1948: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus coquilletti Barber, 1914: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus costalis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus itzelae sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128587" httpUri="https://zenodo.org/record/10128587/files/figure.png" pageId="17" pageNumber="46">6I–L</figureCitation>
,
<figureCitation id="79B7ACB4E9325059FDB733F6FDF5E896" box="[553,569,1719,1740]" captionStart="Figure 7" captionStartId="16.[144,225,1240,1259]" captionTargetBox="[147,1421,199,1208]" captionTargetId="figure-380@16.[144,1424,196,1211]" captionTargetPageId="16" captionText="Figure 7. Distributional map of Heraeus cinnamomeus Barber, 1948, Heraeus coquilletti Barber, 1914, Heraeus costalis sp. nov., Heraeus itzelae sp. nov., and Heraeus setosus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128589" httpUri="https://zenodo.org/record/10128589/files/figure.png" pageId="17" pageNumber="46">7</figureCitation>
,
<figureCitation id="79B7ACB4E9325059FDD933F6FDABE897" box="[583,615,1719,1741]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="17" pageNumber="46">8B</figureCitation>
)
</paragraph>
<paragraph id="E133B031E9325059FF3A339FFED8E8A9" blockId="17.[164,779,1758,1902]" box="[164,276,1758,1779]" pageId="17" pageNumber="46">
<emphasis id="D3F86C23E9325059FF3A339FFED8E8A9" box="[164,276,1758,1779]" italics="true" pageId="17" pageNumber="46">Diagnosis</emphasis>
</paragraph>
<paragraph id="E133B031E9325059FF3A33BDFEFEE934" blockId="17.[164,779,1758,1902]" pageId="17" pageNumber="46">Head shiny, contrasting with dull pronotum and hemelytra. Distiflagellomeres mostly white, with apex darker. Scutellum without erect setae. Profemur mostly dark brown.</paragraph>
<paragraph id="E133B031E9325059FCCD3094FBD7E885" blockId="17.[826,1441,1493,1759]" pageId="17" pageNumber="46">
<taxonomicName id="268CCBB2E9325059FCCD3094FBC9EBB0" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[851,1029,1493,1514]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="costalis" status="sp. nov.">
<emphasis id="D3F86C23E9325059FCCD3094FBC9EBB0" box="[851,1029,1493,1514]" italics="true" pageId="17" pageNumber="46">Heraeus costalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E9325059FB953094FB90EBB0" box="[1035,1116,1493,1514]" pageId="17" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="268CCBB2E9325059FB0D3094FA87EBB0" authorityName="Barber" authorityYear="1948" box="[1171,1355,1493,1514]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="cinnamomeus">
<emphasis id="D3F86C23E9325059FB0D3094FA87EBB0" box="[1171,1355,1493,1514]" italics="true" pageId="17" pageNumber="46">H. cinnamomeus</emphasis>
</taxonomicName>
are the only species of the
<taxonomicName id="268CCBB2E9325059FB8A30B2FB49E852" authorityName="Barber" authorityYear="1914" box="[1044,1157,1523,1544]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="coquilletti">
<emphasis id="D3F86C23E9325059FB8A30B2FB49E852" box="[1044,1157,1523,1544]" italics="true" pageId="17" pageNumber="46">coquilletti</emphasis>
</taxonomicName>
group that have a shiny head, contrasting with a dull pronotum and hemelytra, and the scutellum lacking erect setae. The general colouration of
<taxonomicName id="268CCBB2E9325059FC203311FBFFE83E" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[958,1075,1615,1637]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="costalis" status="sp. nov.">
<emphasis id="D3F86C23E9325059FC203311FBFFE83E" box="[958,1075,1615,1637]" italics="true" pageId="17" pageNumber="46">H. costalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E9325059FBA4330EFB41E83E" box="[1082,1165,1615,1636]" pageId="17" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
is dark brown, with the distiflagellomere mostly white with the apex darker, whereas the general colouration of
<taxonomicName id="268CCBB2E9325059FB5433CCFA4BE8F8" authorityName="Barber" authorityYear="1948" box="[1226,1415,1677,1698]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="cinnamomeus">
<emphasis id="D3F86C23E9325059FB5433CCFA4BE8F8" box="[1226,1415,1677,1698]" italics="true" pageId="17" pageNumber="46">H. cinnamomeus</emphasis>
</taxonomicName>
is cinnamon brown, with distiflagellomere and hemelytra uniformly coloured.
</paragraph>
<paragraph id="E133B031E9325059FCA433BCFC77E948" blockId="17.[826,1441,1789,1902]" box="[826,955,1789,1810]" pageId="17" pageNumber="46">
<emphasis id="D3F86C23E9325059FCA433BCFC77E948" box="[826,955,1789,1810]" italics="true" pageId="17" pageNumber="46">Description</emphasis>
</paragraph>
<paragraph id="E133B031E9325059FCA4325AFBB8E96B" blockId="17.[826,1441,1789,1902]" box="[826,1140,1818,1841]" pageId="17" pageNumber="46">
Total length 6.27 (
<figureCitation id="79B7ACB4E9325059FB8C325AFBABE96B" box="[1042,1127,1819,1841]" captionStart="Figure 3" captionStartId="12.[144,224,1458,1477]" captionTargetBox="[145,1424,195,1428]" captionTargetId="figure-137@12.[144,1424,195,1429]" captionTargetPageId="12" captionText="Figure 3. Habitus dorsal: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus guttatus (Dallas, 1852). G, Heraeus hollyae Baranowski, 2005. H, Heraeus triguttatus (Guérin-Méneville, 1857); I, Heraeus steineri sp. nov.; J, Heraeus caliginosus Slater &amp; Baranowski, 1994; K, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128581" httpUri="https://zenodo.org/record/10128581/files/figure.png" pageId="17" pageNumber="46">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="E133B031E932505AFCCD3278FD26EC13" blockId="17.[826,1441,1789,1902]" lastBlockId="18.[144,759,197,1657]" lastPageId="18" lastPageNumber="47" pageId="17" pageNumber="46">
<emphasis id="D3F86C23E9325059FCCD3278FC5AE914" box="[851,918,1849,1870]" italics="true" pageId="17" pageNumber="46">Head:</emphasis>
Convex dorsally, dark reddish brown, shiny, smooth, with short recumbent and long, erect setae. Head length 1.34, width 0.91. Postocular length 0.48. Eyes with setae between ommatidea; not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.46, interocellar width 0.27. Labium pale brown with erect setae, extending to mesocoxae. Labial segment lengths: I 0.93, II 0.96, III 0.88, and IV 0.37. Antennae brown, except scapus darker and distiflagellomere paler with a broad diffuse subbasal yellowish band, and abundant short recumbent and sparse erect setae. Antennal lengths: scape 0.62, pedicel 1.26, basiflagellomere 0.94, and distiflagellomere 1.30. Length of pale band on distiflagellomere 0.72.
</paragraph>
<paragraph id="E133B031E931505AFF363712FD6BEB1D" blockId="18.[144,759,197,1657]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFF363712FECDEC32" box="[168,257,595,616]" italics="true" pageId="18" pageNumber="47">Thorax:</emphasis>
Pronotum pruinose, with sparse, short, recumbent, and erect setae. Pronotum dark brown; anterior lobe darker; humeral angles with a small pale spot. Pronotum more conspicuously punctate on posterior lobe. Collar length 0.07, anterior lobe length 0.86, posteri- or lobe length 0.48; anterior lobe width 0.98, posteri- or lobe width 1.50. Pleurae dark brown, acetabular areas reddish brown; punctate, with short recumbent setae. Evaporative area short. Scutellum dark brown, pruinose, punctate, with short recumbent setae. Hemelytra pruinose, with short recumbent setae. Hemelytron brown, claval commisure and about apical half of inner margin of clavus, base of corium, anterior threequarters of lateral margins, and a small subapical spot yellowish; apex of corium and adjacent area of membrane pale, membrane pale brown with a diffuse pale apical spot, veins concolorous. Legs: Coxae and protrochanters reddish brown; profemur reddish brown, except base and apex paler; meso- and metafemur bicoloured, base yellowish and apex brown; tibiae and tarsi pale brown, apex of tibiae darker (
<figureCitation id="79B7ACB4E931505AFDC931F7FD67EA96" box="[599,683,1206,1228]" captionStart="Figure 5" captionStartId="14.[144,222,1347,1366]" captionTargetBox="[305,1261,197,1315]" captionTargetId="figure-232@14.[303,1263,195,1317]" captionTargetPageId="14" captionText="Figure 5. Legs: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus caliginosus Slater &amp; Baranowski, 1994; G, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128585" httpUri="https://zenodo.org/record/10128585/files/figure.png" pageId="18" pageNumber="47">Fig. 5C</figureCitation>
). With sparse, short, semi-erect setae. Profemur conspicuously incrassate, spines on profemur small except for one spine contrastingly larger. Protibia with small tubercles bearing spiniform setae on inner surface.
</paragraph>
<paragraph id="E133B031E931505AFF36300EFED2E823" blockId="18.[144,759,197,1657]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFF36300EFED4EB3E" box="[168,280,1359,1380]" italics="true" pageId="18" pageNumber="47">Abdomen:</emphasis>
Reddish brown; with abundant, short, recumbent setae. Male genitalia: Pygophore (
<figureCitation id="79B7ACB4E931505AFD18302FFD21EBD9" box="[646,749,1390,1412]" captionStart="Figure 6" captionStartId="15.[164,243,1703,1722]" captionTargetBox="[243,1361,199,1673]" captionTargetId="figure-0@15.[234,1372,192,1684]" captionTargetPageId="15" captionText="Figure 6. Male genitalia. Heraeus cinnamomeus Barber, 1948: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus coquilletti Barber, 1914: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus costalis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus itzelae sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128587" httpUri="https://zenodo.org/record/10128587/files/figure.png" pageId="18" pageNumber="47">Fig. 6I, J</figureCitation>
) rounded, anterior margin of dorsal aperture slightly rounded, inner projections subquadrangular; declivent posteriorly in lateral view. Parameres:
<figureCitation id="79B7ACB4E931505AFDD0308BFD62EBBA" box="[590,686,1482,1504]" captionStart="Figure 6" captionStartId="15.[164,243,1703,1722]" captionTargetBox="[243,1361,199,1673]" captionTargetId="figure-0@15.[234,1372,192,1684]" captionTargetPageId="15" captionText="Figure 6. Male genitalia. Heraeus cinnamomeus Barber, 1948: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus coquilletti Barber, 1914: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus costalis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus itzelae sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128587" httpUri="https://zenodo.org/record/10128587/files/figure.png" pageId="18" pageNumber="47">Figure 6</figureCitation>
(K, L). Aedeagus (
<figureCitation id="79B7ACB4E931505AFE9130A9FEA8EBA5" box="[271,356,1512,1535]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="18" pageNumber="47">Fig. 8B</figureCitation>
): conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with a few minute spines laterally and two sclerotized lobes with a few minute spines distally;
<emphasis id="D3F86C23E931505AFE793304FD71E800" box="[487,701,1605,1626]" italics="true" pageId="18" pageNumber="47">processus gonopori</emphasis>
long and slender.
</paragraph>
<paragraph id="E133B031E931505AFF0E33E1FED7E8EF" blockId="18.[144,759,1696,1779]" box="[144,283,1696,1717]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFF0E33E1FED7E8EF" box="[144,283,1696,1717]" italics="true" pageId="18" pageNumber="47">Distribution</emphasis>
</paragraph>
<paragraph id="E133B031E931505AFF0E33FEFF24E8A9" blockId="18.[144,759,1696,1779]" pageId="18" pageNumber="47">
<collectingCountry id="999BF0A1E931505AFF0E33FEFEDAE88F" box="[144,278,1727,1749]" name="Costa Rica" pageId="18" pageNumber="47">Costa Rica</collectingCountry>
,
<collectingCountry id="999BF0A1E931505AFEB833FEFE67E88F" box="[294,427,1727,1749]" name="Guatemala" pageId="18" pageNumber="47">Guatemala</collectingCountry>
,
<collectingCountry id="999BF0A1E931505AFE2233FEFDFFE88F" box="[444,563,1727,1749]" name="Honduras" pageId="18" pageNumber="47">Honduras</collectingCountry>
, and
<collectingCountry id="999BF0A1E931505AFDE233FEFD3BE88F" box="[636,759,1727,1749]" name="Nicaragua" pageId="18" pageNumber="47">Nicaragua</collectingCountry>
(
<figureCitation id="79B7ACB4E931505AFF06339CFF16E8A9" box="[152,218,1757,1779]" captionStart="Figure 7" captionStartId="16.[144,225,1240,1259]" captionTargetBox="[147,1421,199,1208]" captionTargetId="figure-380@16.[144,1424,196,1211]" captionTargetPageId="16" captionText="Figure 7. Distributional map of Heraeus cinnamomeus Barber, 1948, Heraeus coquilletti Barber, 1914, Heraeus costalis sp. nov., Heraeus itzelae sp. nov., and Heraeus setosus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128589" httpUri="https://zenodo.org/record/10128589/files/figure.png" pageId="18" pageNumber="47">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="E133B031E931505AFF0E325BFECBE975" blockId="18.[144,758,1818,1902]" box="[144,263,1818,1839]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFF0E325BFECBE975" box="[144,263,1818,1839]" italics="true" pageId="18" pageNumber="47">Etymology</emphasis>
</paragraph>
<paragraph id="E133B031E931505AFF0E3278FE79E934" blockId="18.[144,758,1818,1902]" pageId="18" pageNumber="47">
This species is named ‘
<taxonomicName id="268CCBB2E931505AFE103278FE13E914" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[398,479,1849,1870]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="D3F86C23E931505AFE103278FE13E914" box="[398,479,1849,1870]" italics="true" pageId="18" pageNumber="47">costalis</emphasis>
</taxonomicName>
’ to denote the pale costal margin of the hemelytra.
</paragraph>
<paragraph id="E133B031E931505AFCB83584FC0FEE80" blockId="18.[806,1421,197,280]" box="[806,963,197,218]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFCB83584FC0FEE80" box="[806,963,197,218]" italics="true" pageId="18" pageNumber="47">Type material</emphasis>
</paragraph>
<paragraph id="E133B031E931505AFCB835A2FBB8EF42" blockId="18.[806,1421,197,280]" pageId="18" pageNumber="47">
<materialsCitation id="51E4BA6CE931505AFCB835A2FBB8EF42" collectingDate="1965-07-13" collectionCode="USNM" collectorName="P. J. Spangler" country="Costa Rica" location="Las Canas" pageId="18" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="D3F86C23E931505AFCB835A2FC54EEA2" box="[806,920,227,248]" italics="true" pageId="18" pageNumber="47">
<typeStatus id="3E370E93E931505AFCB835A2FC5EEEA2" box="[806,914,227,248]" pageId="18" pageNumber="47" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E931505AFC3835A2FC73EEA3" box="[934,959,227,249]" pageId="18" pageNumber="47" type="male">♂</specimenCount>
,
<emphasis id="D3F86C23E931505AFC4A35A2FB4BEEA3" bold="true" box="[980,1159,227,249]" pageId="18" pageNumber="47">
<collectingCountry id="999BF0A1E931505AFC4A35A2FB4BEEA3" box="[980,1159,227,249]" name="Costa Rica" pageId="18" pageNumber="47">COSTA RICA</collectingCountry>
</emphasis>
,
<location id="E453E6EAE931505AFB0235A5FAE8EEA2" LSID="urn:lsid:plazi:treatment:69250127E932505AFEAD33D0FB31EDD1:E453E6EAE931505AFB0235A5FAE8EEA2" box="[1180,1316,227,249]" country="Costa Rica" name="Las Canas" pageId="18" pageNumber="47">Las Canas</location>
,
<date id="953296F1E931505AFAAB35A2FC93EF42" pageId="18" pageNumber="47" value="1965-07-13">
<collectingDate id="85766F19E931505AFAAB35A2FC93EF42" pageId="18" pageNumber="47" value="1965-07-13">13-VII- 1965</collectingDate>
</date>
,
<collectorName id="4C79D5E7E931505AFCF03443FBC8EF42" box="[878,1028,258,280]" pageId="18" pageNumber="47">P.J. Spangler</collectorName>
(
<collectionCode id="879D28F4E931505AFB8B3443FBA5EF42" box="[1045,1129,258,280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="47" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E133B031E931505AFCB83471FB31EDD1" blockId="18.[806,1422,303,907]" pageId="18" pageNumber="47">
<emphasis id="D3F86C23E931505AFCB83471FC69EF1F" box="[806,933,304,325]" italics="true" pageId="18" pageNumber="47">
<typeStatus id="3E370E93E931505AFCB83471FC53EF1F" box="[806,927,304,325]" pageId="18" pageNumber="47" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E931505AFC2C3471FC24EF1F" box="[946,1000,303,325]" pageId="18" pageNumber="47" type="male">10♂</specimenCount>
,
<specimenCount id="F78A7BB8E931505AFC633471FBE3EF1F" box="[1021,1071,303,325]" pageId="18" pageNumber="47" type="female">20♀</specimenCount>
, same data as for
<typeStatus id="3E370E93E931505AFAB83471FA41EF1C" box="[1318,1421,304,326]" pageId="18" pageNumber="47" type="holotype">holotype</typeStatus>
(USNM);
<specimenCount id="F78A7BB8E931505AFC32340FFC1EEF3E" box="[940,978,334,356]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
,
<specimenCount id="F78A7BB8E931505AFC7F340FFBCEEF3E" box="[993,1026,334,356]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
, same data as for
<typeStatus id="3E370E93E931505AFB49340FFAFBEF3E" box="[1239,1335,334,356]" pageId="18" pageNumber="47" type="holotype">holotype</typeStatus>
(MLP);
<specimenCount id="F78A7BB8E931505AFCA1342CFCADEFD8" box="[831,865,364,386]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
, same data as for
<typeStatus id="3E370E93E931505AFBA5342CFB50EFD9" box="[1083,1180,365,387]" pageId="18" pageNumber="47" type="holotype">holotype</typeStatus>
(UKANS);
<specimenCount id="F78A7BB8E931505AFABC342CFA89EFD8" box="[1314,1349,364,386]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
,
<quantity id="26741DD4E931505AFACA342CFA41EFD9" box="[1364,1421,365,387]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="18" pageNumber="47" unit="mi" value="9.0">9 mi.</quantity>
NW Esparta,
<date id="953296F1E931505AFC7834CAFBBEEFFB" box="[998,1138,395,417]" pageId="18" pageNumber="47" value="1965-07-22">22-VII-1965</date>
, P.J. Spangler (USNM).
<emphasis id="D3F86C23E931505AFCA134EBFC3EEF9A" bold="true" box="[831,1010,426,448]" pageId="18" pageNumber="47">
<collectingCountry id="999BF0A1E931505AFCA134EBFC21EF9A" box="[831,1005,426,448]" name="Guatemala" pageId="18" pageNumber="47">GUATEMALA</collectingCountry>
:
</emphasis>
<specimenCount id="F78A7BB8E931505AFC6734EBFBD3EF9A" box="[1017,1055,426,448]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
, Los Amates, J.S. Hine (OSUC).
<emphasis id="D3F86C23E931505AFCA13489FC28EF84" bold="true" box="[831,996,456,478]" pageId="18" pageNumber="47">
<collectingCountry id="999BF0A1E931505AFCA13489FC13EF84" box="[831,991,456,478]" name="Honduras" pageId="18" pageNumber="47">HONDURAS</collectingCountry>
:
</emphasis>
<specimenCount id="F78A7BB8E931505AFC723488FBC3EF84" box="[1004,1039,456,478]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
, intercepted at Atlanta (Georgia) from Honduras,
<date id="953296F1E931505AFC6034A6FBB8EFA7" box="[1022,1140,487,509]" pageId="18" pageNumber="47" value="1999-03-05">5-III-1999</date>
, on
<taxonomicName id="268CCBB2E931505AFB3B34A6FC6DEC46" authority="(USNM)" baseAuthorityName="USNM" class="Tubothalamea" family="Milioliporidae" genus="Cucurbita" kingdom="Chromista" order="Miliolida" pageId="18" pageNumber="47" phylum="Foraminifera" rank="species" species="moschata">
<emphasis id="D3F86C23E931505AFB3B34A6FA41EFA6" box="[1189,1421,487,508]" italics="true" pageId="18" pageNumber="47">Cucurbita moschata</emphasis>
(USNM)
</taxonomicName>
;
<specimenCount id="F78A7BB8E931505AFC313747FC1AEC41" box="[943,982,517,539]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
, intercepted at Miami (Florida) from Honduras,
<date id="953296F1E931505AFC263765FBD9EC60" box="[952,1045,548,570]" pageId="18" pageNumber="47" value="2005-01-03">3-I-2005</date>
, on
<taxonomicName id="268CCBB2E931505AFBDC3764FA96EC61" authority="(USNM)" baseAuthorityName="USNM" box="[1090,1370,549,571]" class="Magnoliopsida" family="Cucurbitaceae" genus="Cucumis" kingdom="Plantae" order="Cucurbitales" pageId="18" pageNumber="47" phylum="Tracheophyta" rank="species" species="sativus">
<emphasis id="D3F86C23E931505AFBDC3764FB39EC60" box="[1090,1269,549,570]" italics="true" pageId="18" pageNumber="47">Cucumis sativus</emphasis>
(USNM)
</taxonomicName>
;
<specimenCount id="F78A7BB8E931505AFAFB3764FA4AEC60" box="[1381,1414,548,570]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
, intercepted at West Palm Beach (Florida) from Honduras,
<date id="953296F1E931505AFC5A3723FB8AEC2D" box="[964,1094,610,631]" pageId="18" pageNumber="47" value="1993-04-19">19-IV-1993</date>
, on
<taxonomicName id="268CCBB2E931505AFBE33723FAD5EC2D" box="[1149,1305,610,631]" class="Tubothalamea" family="Milioliporidae" genus="Cucurbita" kingdom="Chromista" order="Miliolida" pageId="18" pageNumber="47" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="D3F86C23E931505AFBE33723FB3DEC2D" box="[1149,1265,610,631]" italics="true" pageId="18" pageNumber="47">Cucurbita</emphasis>
sp.
</taxonomicName>
(USNM);
<emphasis id="D3F86C23E931505AFCA137C0FC0AECCC" box="[831,966,641,662]" italics="true" pageId="18" pageNumber="47">
<collectingRegion id="23487ED3E931505AFCA137C0FC0AECCC" box="[831,966,641,662]" country="Honduras" name="Comayagua" pageId="18" pageNumber="47">Comayagua</collectingRegion>
</emphasis>
:
<specimenCount id="F78A7BB8E931505AFC4937C1FC36ECCC" box="[983,1018,640,662]" pageId="18" pageNumber="47" type="female">2♀</specimenCount>
, 6-VI-[19]66, J.M. Matta (USNM);
<specimenCount id="F78A7BB8E931505AFCA137DEFCA9ECEE" box="[831,869,670,692]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
, 13-IV-[19]66, mosquito trap (USNM);
<specimenCount id="F78A7BB8E931505AFAAC37DEFA95ECEE" box="[1330,1369,670,692]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
, Tegucigalpa, M. Mojica, 62–6225 (USNM).
<emphasis id="D3F86C23E931505AFA8737FCFC4DECA8" bold="true" pageId="18" pageNumber="47">NICARA- GUA:</emphasis>
<specimenCount id="F78A7BB8E931505AFC13379DFC7FECA8" box="[909,947,732,754]" pageId="18" pageNumber="47" type="male">1♂</specimenCount>
, intercepted at Ft. Lauderdale (Florida) from Nicaragua,
<date id="953296F1E931505AFB9137BAFB45ED4B" box="[1039,1161,763,785]" pageId="18" pageNumber="47" value="1996-02-21">21-II-1996</date>
on
<taxonomicName id="268CCBB2E931505AFB2037BAFC6AED75" authority="(USNM)" baseAuthorityName="USNM" class="Magnoliopsida" family="Cucurbitaceae" genus="Citrullus" kingdom="Plantae" order="Cucurbitales" pageId="18" pageNumber="47" phylum="Tracheophyta" rank="species" species="lanatus">
<emphasis id="D3F86C23E931505AFB2037BAFA41ED4A" box="[1214,1421,763,784]" italics="true" pageId="18" pageNumber="47">Citrullus lanatus</emphasis>
(USNM)
</taxonomicName>
; one without abdomen, intercepted at Ft. Lauderdale (Florida) from Nicaragua,
<date id="953296F1E931505AFADE3679FCB6ED36" pageId="18" pageNumber="47" value="1994-03-28">28-III- 1994</date>
on
<taxonomicName id="268CCBB2E931505AFC2A3617FB0BED37" authority="(USNM)" baseAuthorityName="USNM" box="[948,1223,854,877]" class="Liliopsida" family="Amaryllidaceae" genus="Allium" kingdom="Plantae" order="Asparagales" pageId="18" pageNumber="47" phylum="Tracheophyta" rank="species" species="cepa">
<emphasis id="D3F86C23E931505AFC2A3617FB83ED36" box="[948,1103,854,876]" italics="true" pageId="18" pageNumber="47">Allium cepa</emphasis>
(USNM)
</taxonomicName>
;
<specimenCount id="F78A7BB8E931505AFB453616FB33ED36" box="[1243,1279,854,876]" pageId="18" pageNumber="47" type="female">1♀</specimenCount>
, Altamira, Managua,
<date id="953296F1E931505AFC253634FBD4EDD0" box="[955,1048,885,906]" pageId="18" pageNumber="47" value="1970-09">IX-1970</date>
, E. Moore (USNM).
</paragraph>
</subSubSection>
</treatment>
</document>
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