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<document id="D738411DBC55EC01FC614A8F1DDCAEC1" ID-DOI="10.1111/zoj.12362" ID-ISSN="0024-4082" ID-Zenodo-Dep="10109299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235092894" checkinUser="marcus" docAuthor="Dellapé, Pablo M., Melo, María C. &amp; Henry, Thomas J." docDate="2016" docId="69250127E930505DFC5234CAFDC6E882" docLanguage="en" docName="zoj12362.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (1)" docSource="http://dx.doi.org/10.1111/zoj.12362" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus setosus Dellapé &amp; Melo &amp; Henry 2016, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="50" masterDocId="951C795FE9235048FF9E3541FFCCEE5A" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="48" updateTime="1700019240479" updateUser="ExternalLinkService">
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<mods:title id="EA23DDCB6D407D03CA7998EB5BE92215">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="A606FD1ED8E958C4D2DC9D7ED3A7436E">Dellapé, Pablo M.</mods:namePart>
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<mods:namePart id="A27A463F57AE079C570C040C098D547B">Melo, María C.</mods:namePart>
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<mods:namePart id="249E9C3804919431D01A8CF826392BB8">Henry, Thomas J.</mods:namePart>
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<mods:title id="45CB3E69DE0EDA1038B139ECD6469A59">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="87AC6EEB5FF8312DE71E3E12846E5ADF">2016</mods:date>
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<mods:number id="D35B5A2420AA41D8E6F01F117BFC8114">2016-04-17</mods:number>
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<treatment id="69250127E930505DFC5234CAFDC6E882" LSID="urn:lsid:plazi:treatment:69250127E930505DFC5234CAFDC6E882" httpUri="http://treatment.plazi.org/id/69250127E930505DFC5234CAFDC6E882" lastPageId="21" lastPageNumber="50" pageId="19" pageNumber="48">
<subSubSection id="A996E3BAE930505BFC5234CAFAC3EFFA" box="[972,1295,395,416]" pageId="19" pageNumber="48" type="nomenclature">
<paragraph id="E133B031E930505BFC5234CAFAC3EFFA" blockId="19.[972,1295,395,416]" box="[972,1295,395,416]" pageId="19" pageNumber="48">
<heading id="BA7B075DE930505BFC5234CAFAC3EFFA" box="[972,1295,395,416]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="48" reason="4">
<taxonomicName id="268CCBB2E930505BFC5234CAFB60EFC5" ID-CoL="3KXMV" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[972,1196,395,416]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="49" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">HERAEUS SETOSUS</taxonomicName>
<emphasis id="D3F86C23E930505BFB2A34CFFAC3EFFA" bold="true" box="[1204,1295,395,416]" pageId="19" pageNumber="48">
<taxonomicNameLabel id="C8CBD158E930505BFB2A34CFFAC3EFFA" box="[1204,1295,395,416]" pageId="19" pageNumber="48" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A996E3BAE930505DFC4E34F0FDC6E882" lastPageId="21" lastPageNumber="50" pageId="19" pageNumber="48" type="description">
<paragraph id="E133B031E930505BFC4E34F0FAC7EF9D" blockId="19.[976,1291,433,455]" box="[976,1291,433,455]" pageId="19" pageNumber="48">
(
<figureCitation id="79B7ACB4E930505BFC4634F0FBFBEF9C" box="[984,1079,433,454]" captionStart="Figure 3" captionStartId="12.[144,224,1458,1477]" captionTargetBox="[145,1424,195,1428]" captionTargetId="figure-137@12.[144,1424,195,1429]" captionTargetPageId="12" captionText="Figure 3. Habitus dorsal: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus guttatus (Dallas, 1852). G, Heraeus hollyae Baranowski, 2005. H, Heraeus triguttatus (Guérin-Méneville, 1857); I, Heraeus steineri sp. nov.; J, Heraeus caliginosus Slater &amp; Baranowski, 1994; K, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128581" httpUri="https://zenodo.org/record/10128581/files/figure.png" pageId="19" pageNumber="48">FIGS 3E</figureCitation>
,
<figureCitation id="79B7ACB4E930505BFBDA34F0FBAAEF9D" box="[1092,1126,433,455]" captionStart="Figure 5" captionStartId="14.[144,222,1347,1366]" captionTargetBox="[305,1261,197,1315]" captionTargetId="figure-232@14.[303,1263,195,1317]" captionTargetPageId="14" captionText="Figure 5. Legs: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus caliginosus Slater &amp; Baranowski, 1994; G, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128585" httpUri="https://zenodo.org/record/10128585/files/figure.png" pageId="19" pageNumber="48">5E</figureCitation>
,
<figureCitation id="79B7ACB4E930505BFBEA34F0FB48EF9C" box="[1140,1156,433,454]" captionStart="Figure 7" captionStartId="16.[144,225,1240,1259]" captionTargetBox="[147,1421,199,1208]" captionTargetId="figure-380@16.[144,1424,196,1211]" captionTargetPageId="16" captionText="Figure 7. Distributional map of Heraeus cinnamomeus Barber, 1948, Heraeus coquilletti Barber, 1914, Heraeus costalis sp. nov., Heraeus itzelae sp. nov., and Heraeus setosus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128589" httpUri="https://zenodo.org/record/10128589/files/figure.png" pageId="19" pageNumber="48">7</figureCitation>
,
<figureCitation id="79B7ACB4E930505BFB0F34F0FB78EF9C" box="[1169,1204,433,454]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="19" pageNumber="48">8C</figureCitation>
,
<figureCitation id="79B7ACB4E930505BFB5F34F0FAC8EF9D" box="[1217,1284,433,455]" captionStart="Figure 9" captionStartId="20.[144,220,1496,1515]" captionTargetBox="[152,1416,199,1459]" captionTargetId="figure-54@20.[144,1424,194,1466]" captionTargetPageId="20" captionText="Figure 9. Male genitalia. Heraeus setosus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus caliginosus Slater &amp; Baranowski, 1994: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus dominicanus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128593" httpUri="https://zenodo.org/record/10128593/files/figure.png" pageId="19" pageNumber="48">9A–D</figureCitation>
)
</paragraph>
<paragraph id="E133B031E930505BFCA43499FC66EFB7" blockId="19.[826,1442,472,830]" box="[826,938,472,493]" pageId="19" pageNumber="48">
<emphasis id="D3F86C23E930505BFCA43499FC66EFB7" box="[826,938,472,493]" italics="true" pageId="19" pageNumber="48">Diagnosis</emphasis>
</paragraph>
<paragraph id="E133B031E930505BFCA434B7FBB8EC13" blockId="19.[826,1442,472,830]" pageId="19" pageNumber="48">Antennae uniformly coloured; pronotum, scutellum, and hemelytra with erect setae; labium extending to or slightly beyond metacoxae.</paragraph>
<paragraph id="E133B031E930505BFCCD3712FAE0ED64" blockId="19.[826,1442,472,830]" pageId="19" pageNumber="48">
<taxonomicName id="268CCBB2E930505BFCCD3712FBC4EC32" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[851,1032,595,616]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="D3F86C23E930505BFCCD3712FBC4EC32" box="[851,1032,595,616]" italics="true" pageId="19" pageNumber="48">Heraeus setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E930505BFB913713FBAFEC3D" box="[1039,1123,594,615]" pageId="19" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="268CCBB2E930505BFB003712FAC4EC3D" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[1182,1288,594,616]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="itzelae" status="sp. nov.">
<emphasis id="D3F86C23E930505BFB003712FAC4EC3D" box="[1182,1288,594,616]" italics="true" pageId="19" pageNumber="48">H. itzelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E930505BFA913713FAAFEC3D" box="[1295,1379,594,615]" pageId="19" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
have long erect setae on pronotum and scutellum. In
<taxonomicName id="268CCBB2E930505BFCA437D1FC64ECFF" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[826,936,656,677]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="D3F86C23E930505BFCA437D1FC64ECFF" box="[826,936,656,677]" italics="true" pageId="19" pageNumber="48">H. setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E930505BFC3137D1FBCFECFF" box="[943,1027,656,677]" pageId="19" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, the hemelytra also have erect setae, the distiflagellomere lacks a pale band, and the labium extends to the metacoxae, whereas in
<taxonomicName id="268CCBB2E930505BFB41378CFA89ECBB" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[1247,1349,716,738]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="itzelae" status="sp. nov.">
<emphasis id="D3F86C23E930505BFB41378CFA89ECBB" box="[1247,1349,716,738]" italics="true" pageId="19" pageNumber="48">H. itzelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E930505BFAD5378CFA52ECB8" box="[1355,1438,717,738]" pageId="19" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, the distiflagellomere has a pale band, the hemelytra have only scattered, short, recumbent setae, and the labium is shorter, extending to mesocoxae.
</paragraph>
<paragraph id="E133B031E930505BFCA436C4FC77EDC0" blockId="19.[826,1442,901,1902]" box="[826,955,901,922]" pageId="19" pageNumber="48">
<emphasis id="D3F86C23E930505BFCA436C4FC77EDC0" box="[826,955,901,922]" italics="true" pageId="19" pageNumber="48">Description</emphasis>
</paragraph>
<paragraph id="E133B031E930505BFCA436E2FA5CEDE3" blockId="19.[826,1442,901,1902]" box="[826,1424,931,953]" pageId="19" pageNumber="48">
Total length 6.46 (
<figureCitation id="79B7ACB4E930505BFB8C36E2FBABEDE3" box="[1042,1127,931,953]" captionStart="Figure 3" captionStartId="12.[144,224,1458,1477]" captionTargetBox="[145,1424,195,1428]" captionTargetId="figure-137@12.[144,1424,195,1429]" captionTargetPageId="12" captionText="Figure 3. Habitus dorsal: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus guttatus (Dallas, 1852). G, Heraeus hollyae Baranowski, 2005. H, Heraeus triguttatus (Guérin-Méneville, 1857); I, Heraeus steineri sp. nov.; J, Heraeus caliginosus Slater &amp; Baranowski, 1994; K, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128581" httpUri="https://zenodo.org/record/10128581/files/figure.png" pageId="19" pageNumber="48">Fig. 3E</figureCitation>
). Strongly setose species.
</paragraph>
<paragraph id="E133B031E930505BFCCD3683FAADEB1D" blockId="19.[826,1442,901,1902]" pageId="19" pageNumber="48">
<emphasis id="D3F86C23E930505BFCCD3683FC5FED8D" box="[851,915,962,983]" italics="true" pageId="19" pageNumber="48">Head:</emphasis>
Strongly convex dorsally, brown, coriaceous with abundant recumbent and erect setae. Head length 1.27, width 0.88. Postocular length 0.41. Eyes slightly protruding, not surpassing dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.45, interocellar width 0.30. Labium light brown with erect setae, extending to metacoxae. Labial segment lengths: I 0.90, II 0.99, III 0.90, and IV 0.37. Antennal colouration yellowish brown. Pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.62, pedicel 1.60, basiflagellomere, and distiflagellomere missing.
</paragraph>
<paragraph id="E133B031E930505BFCCD300EFBCCE934" blockId="19.[826,1442,901,1902]" pageId="19" pageNumber="48">
<emphasis id="D3F86C23E930505BFCCD300EFC66EB3E" box="[851,938,1359,1380]" italics="true" pageId="19" pageNumber="48">Thorax:</emphasis>
Pronotum pruinose; punctate, those of anterior lobe shallow; dark brown, posterior lobe and humeral angles paler; pruinose, with abundant, long, erect setae. Collar length 0.08, anterior lobe length 0.82, posterior lobe length 0.54; anterior lobe width 1.14, posterior lobe width 1.68. Pleurae brown. Evaporative area short. Scutellum brown, pruinose, punctate, with abundant long erect setae. Hemelytra pruinose, brown, with abundant erect setae shorter than those on scutellum; clavus unicolorous; costal margin pale on proximal three-quarters. Inner corial spot well defined. Membrane brown, veins paler. Legs: Coxae and protrochanter brown; profemur brown with apex paler, rest of legs pale brown; meso- and metafemur slightly darkened distally, tibiae darker at apices (
<figureCitation id="79B7ACB4E930505BFAA133BDFA58E948" box="[1343,1428,1788,1810]" captionStart="Figure 5" captionStartId="14.[144,222,1347,1366]" captionTargetBox="[305,1261,197,1315]" captionTargetId="figure-232@14.[303,1263,195,1317]" captionTargetPageId="14" captionText="Figure 5. Legs: A, Heraeus cinnamomeus Barber, 1948; B, Heraeus coquilletti Barber, 1914; C, Heraeus costalis sp. nov.; D, Heraeus itzelae sp. nov.; E, Heraeus setosus sp. nov.; F, Heraeus caliginosus Slater &amp; Baranowski, 1994; G, Heraeus dominicanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128585" httpUri="https://zenodo.org/record/10128585/files/figure.png" pageId="19" pageNumber="48">Fig. 5E</figureCitation>
). Strongly setose, with long erect and semi-erect setae. Protibia with small tubercles bearing spiniform setae on inner surface.
</paragraph>
<caption id="B5F3E0B9E937505CFF0E3099FE55E83A" ID-DOI="http://doi.org/10.5281/zenodo.10128593" ID-Zenodo-Dep="10128593" httpUri="https://zenodo.org/record/10128593/files/figure.png" pageId="20" pageNumber="49" startId="20.[144,220,1496,1515]" targetBox="[152,1416,199,1459]" targetPageId="20">
<paragraph id="E133B031E937505CFF0E3099FE55E83A" blockId="20.[144,1422,1496,1632]" pageId="20" pageNumber="49">
<emphasis id="D3F86C23E937505CFF0E3099FF3FEBB1" bold="true" box="[144,243,1496,1515]" pageId="20" pageNumber="49">Figure 9.</emphasis>
Male genitalia.
<emphasis id="D3F86C23E937505CFE053099FD68EBB1" bold="true" box="[411,676,1496,1515]" pageId="20" pageNumber="49">
<taxonomicName id="268CCBB2E937505CFE053099FD82EBB1" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[411,590,1496,1515]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="D3F86C23E937505CFE053099FD82EBB1" bold="true" box="[411,590,1496,1515]" italics="true" pageId="20" pageNumber="49">Heraeus setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E937505CFDCD3099FD68EBB1" box="[595,676,1496,1515]" pageId="20" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view.
<taxonomicName id="268CCBB2E937505CFD1130B7FB52E852" authority="Slater &amp; Baranowski, 1994" authorityName="Slater &amp; Baranowski" authorityYear="1994" box="[655,1182,1525,1544]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="caliginosus">
<emphasis id="D3F86C23E937505CFD1130B7FCAAE852" box="[655,870,1525,1544]" italics="true" pageId="20" pageNumber="49">Heraeus caliginosus</emphasis>
<bibRefCitation id="851DCDC0E937505CFCEF30B4FB52E852" author="Slater JA &amp; Baranowski RM" box="[881,1182,1525,1544]" pageId="20" pageNumber="49" pagination="488 - 494" refId="ref78146" refString="Slater JA, Baranowski RM. 1994. Three new species of rhyparochromine Lygaeidae (Hemiptera: Heteroptera) from Hispaniola. Florida Entomologist 77: 488 - 494." type="journal article" year="1994">Slater &amp; Baranowski, 1994</bibRefCitation>
</taxonomicName>
: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view.
<emphasis id="D3F86C23E937505CFAB93352FE48E819" bold="true" pageId="20" pageNumber="49">
<taxonomicName id="268CCBB2E937505CFAB93352FEE4E818" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="dominicanus" status="sp. nov.">
<emphasis id="D3F86C23E937505CFAB93352FEE4E818" bold="true" italics="true" pageId="20" pageNumber="49">Heraeus dominicanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E937505CFEB03371FE48E819" box="[302,388,1584,1603]" pageId="20" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view.
</paragraph>
</caption>
<paragraph id="E133B031E937505CFF3633FEFE40E934" blockId="20.[144,759,1727,1902]" pageId="20" pageNumber="49">
<emphasis id="D3F86C23E937505CFF3633FEFED6E88E" box="[168,282,1727,1748]" italics="true" pageId="20" pageNumber="49">Abdomen:</emphasis>
Brown with abundant short recumbent setae. Male genitalia: Pygophore rounded, anterior margin of dorsal aperture slightly rounded (
<figureCitation id="79B7ACB4E937505CFD0333BDFF68E96A" captionStart="Figure 9" captionStartId="20.[144,220,1496,1515]" captionTargetBox="[152,1416,199,1459]" captionTargetId="figure-54@20.[144,1424,194,1466]" captionTargetPageId="20" captionText="Figure 9. Male genitalia. Heraeus setosus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus caliginosus Slater &amp; Baranowski, 1994: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus dominicanus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128593" httpUri="https://zenodo.org/record/10128593/files/figure.png" pageId="20" pageNumber="49">Fig. 9A, B</figureCitation>
). Parameres: Figure (9C, D). Aedeagus with minute spines; vesical lobes sclerotized;
<emphasis id="D3F86C23E937505CFE6A327BFD0DE915" box="[500,705,1850,1871]" italics="true" pageId="20" pageNumber="49">processus gonopori</emphasis>
long and slender (
<figureCitation id="79B7ACB4E937505CFEB43219FE4CE934" box="[298,384,1880,1902]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="20" pageNumber="49">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="E133B031E937505CFCB833FEFB52E88E" blockId="20.[806,1273,1727,1779]" box="[806,1182,1727,1748]" pageId="20" pageNumber="49">
<emphasis id="D3F86C23E937505CFCB833FEFB52E88E" box="[806,1182,1727,1748]" italics="true" pageId="20" pageNumber="49">
Variability observed in
<typeStatus id="3E370E93E937505CFBAE33FEFB52E88E" box="[1072,1182,1727,1748]" pageId="20" pageNumber="49" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E133B031E937505CFCB8339CFB35E8A9" blockId="20.[806,1273,1727,1779]" box="[806,1273,1757,1779]" pageId="20" pageNumber="49">All antennal segments yellowish brown.</paragraph>
<paragraph id="E133B031E937505CFCB83278FC7DE914" blockId="20.[806,1159,1849,1902]" box="[806,945,1849,1870]" pageId="20" pageNumber="49">
<emphasis id="D3F86C23E937505CFCB83278FC7DE914" box="[806,945,1849,1870]" italics="true" pageId="20" pageNumber="49">Distribution</emphasis>
</paragraph>
<paragraph id="E133B031E937505CFCB83219FB4BE937" blockId="20.[806,1159,1849,1902]" box="[806,1159,1880,1902]" pageId="20" pageNumber="49">
<collectingCountry id="999BF0A1E937505CFCB83219FC5BE934" box="[806,919,1880,1902]" name="Honduras" pageId="20" pageNumber="49">Honduras</collectingCountry>
and
<collectingCountry id="999BF0A1E937505CFC483219FBEAE934" box="[982,1062,1880,1902]" name="Mexico" pageId="20" pageNumber="49">Mexico</collectingCountry>
(
<figureCitation id="79B7ACB4E937505CFBA93219FBB6E937" box="[1079,1146,1880,1902]" captionStart="Figure 7" captionStartId="16.[144,225,1240,1259]" captionTargetBox="[147,1421,199,1208]" captionTargetId="figure-380@16.[144,1424,196,1211]" captionTargetPageId="16" captionText="Figure 7. Distributional map of Heraeus cinnamomeus Barber, 1948, Heraeus coquilletti Barber, 1914, Heraeus costalis sp. nov., Heraeus itzelae sp. nov., and Heraeus setosus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128589" httpUri="https://zenodo.org/record/10128589/files/figure.png" pageId="20" pageNumber="49">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="E133B031E936505DFF3835C2FCA5EEC1" blockId="21.[166,873,131,155]" box="[166,873,131,155]" pageId="21" pageNumber="50">
50 A REVISION OF
<taxonomicName id="268CCBB2E936505DFE2E35C4FDFCEEC0" authorityName="Stal" authorityYear="1862" box="[432,560,133,154]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="D3F86C23E936505DFE2E35C4FDFCEEC0" box="[432,560,133,154]" italics="true" pageId="21" pageNumber="50">HERAEUS</emphasis>
</taxonomicName>
(
<taxonomicName id="268CCBB2E936505DFDA135C4FCAFEEC1" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[575,867,133,155]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="family">RHYPAROCHROMIDAE</taxonomicName>
)
</paragraph>
<caption id="B5F3E0B9E936505DFF3A3197FA6DEAB3" ID-DOI="http://doi.org/10.5281/zenodo.10128595" ID-Zenodo-Dep="10128595" box="[164,1441,1237,1257]" httpUri="https://zenodo.org/record/10128595/files/figure.png" pageId="21" pageNumber="50" startId="21.[164,243,1238,1257]" targetBox="[164,1444,194,1208]" targetPageId="21">
<paragraph id="E133B031E936505DFF3A3197FA6DEAB3" blockId="21.[164,1441,1237,1257]" box="[164,1441,1237,1257]" pageId="21" pageNumber="50">
<emphasis id="D3F86C23E936505DFF3A3197FED5EAB3" bold="true" box="[164,281,1238,1257]" pageId="21" pageNumber="50">Figure 10.</emphasis>
Distributional map of
<taxonomicName id="268CCBB2E936505DFD8C3197FBC1EAB3" authority="Slater &amp; Baranowski, 1994" authorityName="Slater &amp; Baranowski" authorityYear="1994" box="[530,1037,1238,1257]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="caliginosus">
<emphasis id="D3F86C23E936505DFD8C3197FD29EAB3" box="[530,741,1238,1257]" italics="true" pageId="21" pageNumber="50">Heraeus caliginosus</emphasis>
<bibRefCitation id="851DCDC0E936505DFD723197FBC1EAB3" author="Slater JA &amp; Baranowski RM" box="[748,1037,1238,1257]" pageId="21" pageNumber="50" pagination="488 - 494" refId="ref78146" refString="Slater JA, Baranowski RM. 1994. Three new species of rhyparochromine Lygaeidae (Hemiptera: Heteroptera) from Hispaniola. Florida Entomologist 77: 488 - 494." type="journal article" year="1994">Slater &amp; Baranowski, 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="268CCBB2E936505DFBDB3197FA89EAB2" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[1093,1349,1237,1257]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="dominicanus" status="sp. nov.">
<emphasis id="D3F86C23E936505DFBDB3197FA89EAB2" bold="true" box="[1093,1349,1237,1257]" italics="true" pageId="21" pageNumber="50">Heraeus dominicanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E936505DFAD53197FA6DEAB3" box="[1355,1441,1238,1257]" pageId="21" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E133B031E936505DFF3A3061FED7EB6F" blockId="21.[164,779,1312,1395]" box="[164,283,1312,1333]" pageId="21" pageNumber="50">
<emphasis id="D3F86C23E936505DFF3A3061FED7EB6F" box="[164,283,1312,1333]" italics="true" pageId="21" pageNumber="50">Etymology</emphasis>
</paragraph>
<paragraph id="E133B031E936505DFF3A307FFDF1EB28" blockId="21.[164,779,1312,1395]" pageId="21" pageNumber="50">
This species is named ‘
<taxonomicName id="268CCBB2E936505DFE22307EFDC0EB0E" authority="Latin" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[444,524,1343,1364]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="D3F86C23E936505DFE22307EFDC0EB0E" box="[444,524,1343,1364]" italics="true" pageId="21" pageNumber="50">setosus</emphasis>
</taxonomicName>
’, Latin for bristly, to denote the long, erect dorsal setae.
</paragraph>
<paragraph id="E133B031E936505DFF3A30E1FE8DEBEF" blockId="21.[164,779,1440,1554]" box="[164,321,1440,1461]" pageId="21" pageNumber="50">
<emphasis id="D3F86C23E936505DFF3A30E1FE8DEBEF" box="[164,321,1440,1461]" italics="true" pageId="21" pageNumber="50">Type material</emphasis>
</paragraph>
<paragraph id="E133B031E936505DFF3A30FFFDBCE848" blockId="21.[164,779,1440,1554]" pageId="21" pageNumber="50">
<materialsCitation id="51E4BA6CE936505DFF3A30FFFDBCE848" collectionCode="USNM" collectorName="Lk. Yooa &amp; Agua Azul, U. F. Co. &amp; F. S. Blanton &amp; A. B. Brose &amp; R. E. Woodruff" country="Honduras" location="Cortes" pageId="21" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Cortes" typeStatus="holotype">
<emphasis id="D3F86C23E936505DFF3A30FFFEC4EB89" box="[164,264,1470,1491]" italics="true" pageId="21" pageNumber="50">
<typeStatus id="3E370E93E936505DFF3A30FFFECFEB89" box="[164,259,1470,1491]" pageId="21" pageNumber="50" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E936505DFE9130FFFEEBEB8E" box="[271,295,1470,1492]" pageId="21" pageNumber="50" type="male">♂</specimenCount>
,
<emphasis id="D3F86C23E936505DFEAA30FFFE01EB8E" bold="true" box="[308,461,1470,1492]" pageId="21" pageNumber="50">
<collectingCountry id="999BF0A1E936505DFEAA30FFFE01EB8E" box="[308,461,1470,1492]" name="Honduras" pageId="21" pageNumber="50">HONDURAS</collectingCountry>
</emphasis>
, Dept
<collectingRegion id="23487ED3E936505DFD8830FFFD93EB89" box="[534,607,1470,1491]" country="Honduras" name="Cortes" pageId="21" pageNumber="50">Cortes</collectingRegion>
,
<collectorName id="4C79D5E7E936505DFDF730FFFD04EB8E" box="[617,712,1470,1492]" pageId="21" pageNumber="50">Lk. Yooa</collectorName>
,
<collectorName id="4C79D5E7E936505DFD4F30FFFE88EBA8" pageId="21" pageNumber="50">Agua Azul, U.F. Co.</collectorName>
, 1-VI-[19]64,
<collectorName id="4C79D5E7E936505DFE40309CFDA0EBA9" box="[478,620,1501,1523]" pageId="21" pageNumber="50">F.S. Blanton</collectorName>
,
<collectorName id="4C79D5E7E936505DFDE9309CFD23EBA8" box="[631,751,1501,1522]" pageId="21" pageNumber="50">A.B. Brose</collectorName>
&amp;
<collectorName id="4C79D5E7E936505DFF3A30BDFE8AE84B" box="[164,326,1531,1553]" pageId="21" pageNumber="50">R.E. Woodruff</collectorName>
,
<collectingMethod id="38CDC826E936505DFECD30BAFE33E84B" box="[339,511,1531,1553]" pageId="21" pageNumber="50">blacklight trap</collectingMethod>
(
<collectionCode id="879D28F4E936505DFD8F30BDFDA9E848" box="[529,613,1532,1554]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="50" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E133B031E936505DFF3A336BFDC6E882" blockId="21.[164,779,1577,1752]" pageId="21" pageNumber="50">
<materialsCitation id="51E4BA6CE936505DFF3A336BFEB0E804" collectingDate="2003-09-26" collectionCode="USNM" country="Mexico" location="Aircraft" pageId="21" pageNumber="50" specimenCount="2" specimenCount-female="2" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="D3F86C23E936505DFF3A336BFED7E865" box="[164,283,1578,1599]" italics="true" pageId="21" pageNumber="50">
<typeStatus id="3E370E93E936505DFF3A336BFED9E865" box="[164,277,1578,1599]" pageId="21" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<emphasis id="D3F86C23E936505DFEBC3368FE55E865" bold="true" box="[290,409,1577,1599]" pageId="21" pageNumber="50">
<collectingCountry id="999BF0A1E936505DFEBC3368FE5FE865" box="[290,403,1577,1599]" name="Mexico" pageId="21" pageNumber="50">MEXICO</collectingCountry>
:
</emphasis>
<specimenCount id="F78A7BB8E936505DFE013368FE0EE865" box="[415,450,1577,1599]" pageId="21" pageNumber="50" type="female">2♀</specimenCount>
,
<collectingRegion id="23487ED3E936505DFE4E3368FDECE865" box="[464,544,1577,1599]" country="Mexico" name="Mexico" pageId="21" pageNumber="50">Mexico</collectingRegion>
<location id="E453E6EAE936505DFDBB3368FD4EE865" LSID="urn:lsid:plazi:treatment:69250127E930505DFC5234CAFDC6E882:E453E6EAE936505DFDBB3368FD4EE865" box="[549,642,1577,1599]" country="Mexico" name="Aircraft" pageId="21" pageNumber="50" stateProvince="Mexico">Aircraft</location>
,
<date id="953296F1E936505DFD133368FCC6E865" box="[653,778,1577,1599]" pageId="21" pageNumber="50" value="2003-09-26">
<collectingDate id="85766F19E936505DFD133368FCC6E865" box="[653,778,1577,1599]" pageId="21" pageNumber="50" value="2003-09-26">26-IX-2003</collectingDate>
</date>
(
<collectionCode id="879D28F4E936505DFF5A3309FED8E804" box="[196,276,1608,1630]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="50" type="Museum">USNM</collectionCode>
);
<collectingRegion id="23487ED3E936505DFEBA3309FEB0E804" box="[292,380,1608,1630]" country="Mexico" name="Nayarit" pageId="21" pageNumber="50">Nayarit</collectingRegion>
</materialsCitation>
,
<materialsCitation id="51E4BA6CE936505DFE1A3309FCCAE827" collectingDate="1974-07-20" collectionCode="USNM" collectorName="M. E. &amp; P. D. Perkins" country="Mexico" location="18 km SW Compostela" pageId="21" pageNumber="50" specimenCount="1" specimenCount-female="1" stateProvince="Nayarit" typeStatus="paratype">
<specimenCount id="F78A7BB8E936505DFE1A3309FE69E807" box="[388,421,1607,1629]" pageId="21" pageNumber="50" type="female">1♀</specimenCount>
,
<emphasis id="D3F86C23E936505DFE2E3308FDC8E804" box="[432,516,1609,1630]" italics="true" pageId="21" pageNumber="50">
<collectingRegion id="23487ED3E936505DFE2E3308FDC8E804" box="[432,516,1609,1630]" country="Mexico" name="Nayarit" pageId="21" pageNumber="50">Nayarit</collectingRegion>
</emphasis>
:
<location id="E453E6EAE936505DFD8E3309FCCBE804" LSID="urn:lsid:plazi:treatment:69250127E930505DFC5234CAFDC6E882:E453E6EAE936505DFD8E3309FCCBE804" box="[528,775,1608,1630]" country="Mexico" name="18 km SW Compostela" pageId="21" pageNumber="50" stateProvince="Nayarit">
<quantity id="26741DD4E936505DFD8E3309FD98E804" box="[528,596,1608,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="50" unit="km" value="18.0">18 km</quantity>
SW Compostela
</location>
,
<date id="953296F1E936505DFF223326FE88E826" box="[188,324,1639,1660]" pageId="21" pageNumber="50" value="1974-07-20">
<collectingDate id="85766F19E936505DFF223326FE88E826" box="[188,324,1639,1660]" pageId="21" pageNumber="50" value="1974-07-20">20-VII-1974</collectingDate>
</date>
, blacklight,
<collectorName id="4C79D5E7E936505DFE583326FD6EE827" box="[454,674,1639,1661]" pageId="21" pageNumber="50">M.E. &amp; P.D. Perkins</collectorName>
(
<collectionCode id="879D28F4E936505DFD303326FD33E827" box="[686,767,1639,1661]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="50" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE936505DFF2233C4FED6E8E3" collectingDate="1974-07-20" collectionCode="MLP" collectorName="M. E. &amp; P. D. Perkins" country="Mexico" location="Chiapas" pageId="21" pageNumber="50" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="F78A7BB8E936505DFF2233C4FF11E8C1" box="[188,221,1669,1691]" pageId="21" pageNumber="50" type="female">1♀</specimenCount>
,
<date id="953296F1E936505DFF7633C4FEA6E8C0" box="[232,362,1669,1690]" pageId="21" pageNumber="50" value="1974-07-20">
<collectingDate id="85766F19E936505DFF7633C4FEA6E8C0" box="[232,362,1669,1690]" pageId="21" pageNumber="50" value="1974-07-20">20-VII-1974</collectingDate>
</date>
, blacklight,
<collectorName id="4C79D5E7E936505DFE7933C7FD77E8C1" box="[487,699,1669,1691]" pageId="21" pageNumber="50">M.E. &amp; P.D. Perkins</collectorName>
(
<collectionCode id="879D28F4E936505DFD5933C4FCCCE8C1" box="[711,768,1669,1691]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="21" pageNumber="50">MLP</collectionCode>
);
<emphasis id="D3F86C23E936505DFF2233E5FED6E8E3" box="[188,282,1700,1721]" italics="true" pageId="21" pageNumber="50">
<collectingRegion id="23487ED3E936505DFF2233E5FED6E8E3" box="[188,282,1700,1721]" country="Mexico" name="Chiapas" pageId="21" pageNumber="50">Chiapas</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="51E4BA6CE936505DFEB533E5FDCBE882" collectingDate="1964-07-22" collectionCode="USNM" collectorName="Spangler" country="Mexico" location="25 mi. E of Tuxtla Gutierrez" pageId="21" pageNumber="50" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E936505DFEB533E5FE82E8E3" box="[299,334,1699,1721]" pageId="21" pageNumber="50" type="female">1♀</specimenCount>
,
<location id="E453E6EAE936505DFEC033E5FD67E8E0" LSID="urn:lsid:plazi:treatment:69250127E930505DFC5234CAFDC6E882:E453E6EAE936505DFEC033E5FD67E8E0" box="[350,683,1700,1722]" country="Mexico" name="25 mi. E of Tuxtla Gutierrez" pageId="21" pageNumber="50">
<quantity id="26741DD4E936505DFEC033E5FE69E8E0" box="[350,421,1700,1722]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="21" pageNumber="50" unit="mi" value="25.0">25 mi.</quantity>
E of Tuxtla Gutiérrez
</location>
,
<date id="953296F1E936505DFD2733E5FF39E882" pageId="21" pageNumber="50" value="1964-07-22">
<collectingDate id="85766F19E936505DFD2733E5FF39E882" pageId="21" pageNumber="50" value="1964-07-22">22-VII- 1964</collectingDate>
</date>
, P.J.
<collectorName id="4C79D5E7E936505DFEAD3383FE56E882" box="[307,410,1730,1752]" pageId="21" pageNumber="50">Spangler</collectorName>
(
<collectionCode id="879D28F4E936505DFE353383FE33E882" box="[427,511,1730,1752]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="50" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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