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<document id="7C470F9CAC7BBFAB8B222039D915035A" ID-CLB-Dataset="26691" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.30778" ID-GBIF-Dataset="00106674-20d9-4d70-bb0f-8f000fb311f2" ID-Pensoft-Pub="2535-0730-2-169" ID-ZooBank="C0E89FEC8BE54DE9803D784FF6727BA0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-2-169" ModsDocOrigin="Evolutionary Systematics 2 (2)" ModsDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" checkinTime="1558621757476" checkinUser="pensoft" docAuthor="Framenau, Volker W. &amp; Baehr, Barbara C." docDate="2018" docId="1F6F73940C10792F082C5B283F62F8B2" docLanguage="en" docName="EvolutSyst 2(2): 169-241" docOrigin="Evolutionary Systematics 2 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.2.30778" docTitle="Artoria equipalus Framenau &amp; Baehr, 2018, sp. n." docType="treatment" docUuid="1B77FE1F-B9F6-4517-A47C-4D74CFEFDA13" docUuidSource="ZooBank" docVersion="6" lastPageNumber="169" masterDocId="FFE8FFB72743554D943BFFB83D324B46" masterDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="241" masterPageNumber="169" pageNumber="169" updateTime="1701381204780" updateUser="ExternalLinkService">
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<mods:title id="8666F5D96C4FF94A3F9EE90920FDEA16">The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
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<mods:namePart id="4ACD518FA79AEF254314CAF98292556E">Framenau, Volker W.</mods:namePart>
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<mods:namePart id="219791E2D89E313751E395FBF7E99231">Baehr, Barbara C.</mods:namePart>
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<mods:title id="58F4C7E1A72CE6412527FFA1E3AA56C4">Evolutionary Systematics</mods:title>
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<mods:date id="6AD47615317D628B9E87F5E72ABD2295">2018</mods:date>
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<mods:url id="2BF07468A53CD239F29AF36769C8799A">http://dx.doi.org/10.3897/evolsyst.2.30778</mods:url>
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<treatment id="1F6F73940C10792F082C5B283F62F8B2" ID-GBIF-Taxon="157251913" LSID="urn:lsid:zoobank.org:act:1B77FE1F-B9F6-4517-A47C-4D74CFEFDA13" httpUri="http://treatment.plazi.org/id/1F6F73940C10792F082C5B283F62F8B2" lastPageNumber="169" pageId="0" pageNumber="169">
<subSubSection id="4C4E6690282C713A9AC13C1C78864F16" pageId="0" pageNumber="169" type="nomenclature">
<paragraph id="349D72952CC437C17EBC072CBB76F235" pageId="0" pageNumber="169">
<taxonomicName id="F42742ED237631A7087D1ED00D84E962" ID-CoL="H2BL" LSID="http://zoobank.org/1B77FE1F-B9F6-4517-A47C-4D74CFEFDA13" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria equipalus" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="equipalus">Artoria equipalus</taxonomicName>
<taxonomicNameLabel id="71ADC85484D3B5919784A70F65E6F797" pageId="0" pageNumber="169">sp. n.</taxonomicNameLabel>
Figs 15, 17
<normalizedToken id="0F3B0012229AB094CFF8647F788D2BF8" originalValue="AH">A-H</normalizedToken>
, 47H Horse Swamp Forest Runner
</paragraph>
</subSubSection>
<subSubSection id="97A54900595594E1A532081AA3D85E64" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="F33B12FEAE76F79F84598CC2773192BE" pageId="0" pageNumber="169">Material examined.</paragraph>
<paragraph id="6E1F93FC2A21CA6E866D7F1DEFAF6EF8" pageId="0" pageNumber="169">
Holotype male, Barrington Tops National Park, 100 m from Horse Swamp Track (
<geoCoordinate id="CE4D759488F53749648C0D7BC5BD09C9" direction="south" orientation="latitude" precision="15" value="-31.933332">
31°56
<normalizedToken id="3B838D4CBA601421D009EEE87CEB0F06" originalValue="00”">'00&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="1BDFE770AE27768350B920F96D2960B1" direction="east" orientation="longitude" precision="15" value="151.38445">
151°23
<normalizedToken id="16AE43CA33F0B78803EFF1CAAA4DBF18" originalValue="04”">'04&quot;</normalizedToken>
E
</geoCoordinate>
, New South Wales, AUSTRALIA), 11-21 January 2012, J.R. Gollan, M.A. Ashcroft, pitfall trap, upland swamp (AM KS128077). Paratype: 1 female, same data as holotype (AM KS122934).
</paragraph>
</subSubSection>
<subSubSection id="F1BB12F0B90C855B2991F4FC59B8979D" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="52EA0E35CE0F13617248FE385059E709" pageId="0" pageNumber="169">Other material examined.</paragraph>
<paragraph id="8C166F5F8C4F3003F06C9F71F42DB6E5" pageId="0" pageNumber="169">Known only from type material.</paragraph>
</subSubSection>
<subSubSection id="CD479CD2172066B62DCD6ACABE29418D" pageId="0" pageNumber="169" type="etymology">
<paragraph id="145838BC4FF7A2098C5FB976DF564E64" pageId="0" pageNumber="169">Etymology.</paragraph>
<paragraph id="688570D75312FF03E04A7F8849987FDE" pageId="0" pageNumber="169">The specific epithet is a compound noun in apposition derived from the Latin equus (= horse) and palus (= swamp) and refers to the type locality, near Horse Swamp Track.</paragraph>
</subSubSection>
<subSubSection id="683E0D84B597811CEEAEF59AB46A80A9" pageId="0" pageNumber="169" type="diagnosis">
<paragraph id="1E20BD3ECA27F3CB661D35428283FCDA" pageId="0" pageNumber="169">Diagnosis.</paragraph>
<paragraph id="5227595670E986A625C72561AD80943D" pageId="0" pageNumber="169">
Males of
<taxonomicName id="CC19211237FFF111E85653441C7532B0" lsidName="A. equipalus" pageId="0" pageNumber="169" rank="species" species="equipalus">A. equipalus</taxonomicName>
sp. n. share with
<taxonomicName id="2FD68DE0796432EF889A9FB132E1E21B" lsidName="A. booderee" pageId="0" pageNumber="169" rank="species" species="booderee">A. booderee</taxonomicName>
sp. n.,
<taxonomicName id="5B6D9516A777E0076DCAE4422E2FB424" lsidName="A. corowa" pageId="0" pageNumber="169" rank="species" species="corowa">A. corowa</taxonomicName>
sp. n. and
<taxonomicName id="6F61672D6F0ECF47F80E4DE5440134C3" lsidName="A. munmorah" pageId="0" pageNumber="169" rank="species" species="munmorah">A. munmorah</taxonomicName>
sp. n. a distinctly bi-lobed tegular apophysis, but differ from these by the basal lobe of the tegular apophysis being more sclerotized and oriented more
<normalizedToken id="AFEBACF9E4E055992584757C2FFC0F14" originalValue="horizontally">'horizontally'</normalizedToken>
in ventral view. The cymbium tip in
<taxonomicName id="57647BC3F0F0A849545608D9AD0EEF9E" lsidName="A. equipalus" pageId="0" pageNumber="169" rank="species" species="equipalus">A. equipalus</taxonomicName>
has 4-5 macrosetae, which are absent in
<taxonomicName id="6F053B96760237DA67DC01337EFC04F0" lsidName="A. corowa" pageId="0" pageNumber="169" rank="species" species="corowa">A. corowa</taxonomicName>
sp. n. and
<taxonomicName id="1F849BB719B514F5433F070A7305295D" lsidName="A. munmorah" pageId="0" pageNumber="169" rank="species" species="munmorah">A. munmorah</taxonomicName>
sp. n.
<taxonomicName id="7DA788F5E1DF6530042B155FF070BFC2" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria booderee" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="booderee">Artoria booderee</taxonomicName>
sp. n. has 2-3 macrosetae on the cymbium tip which is more elongate than that of
<taxonomicName id="2EB34275FAD1CF9F4C9CBAB5F1E3FBF5" lsidName="A. equipalus" pageId="0" pageNumber="169" rank="species" species="equipalus">A. equipalus</taxonomicName>
sp. n. The epigyne of female
<taxonomicName id="1B6D02830ED4ED2CB6B2E25F9C853D76" lsidName="A. equipalus" pageId="0" pageNumber="169" rank="species" species="equipalus">A. equipalus</taxonomicName>
sp. n. is most similar to that of
<taxonomicName id="B8379495EFEDC560EC177D829FF2111D" lsidName="A. mungo" pageId="0" pageNumber="169" rank="species" species="mungo">A. mungo</taxonomicName>
sp. n., but the shape of the spermathecal heads is very different in both species with in particular with respect to the insertion of the spermathecal stalks (postero-medial in
<taxonomicName id="E93CB7820AFD26A047FCCACBB60B88B2" lsidName="A. equipalus" pageId="0" pageNumber="169" rank="species" species="equipalus">A. equipalus</taxonomicName>
sp. n., lateral in
<taxonomicName id="C9B46AEF5B30E8E42AA83BD6D990D780" lsidName="A. mungo" pageId="0" pageNumber="169" rank="species" species="mungo">A. mungo</taxonomicName>
sp. n.).
</paragraph>
</subSubSection>
<subSubSection id="DD3586EED4C269D9659F374C0A9762D7" pageId="0" pageNumber="169" type="description">
<paragraph id="D2FB1D21F249037F9A59E253B07E771B" pageId="0" pageNumber="169">Description.</paragraph>
<paragraph id="93CDDA20F10A2FCED65B78ACE10AA17B" pageId="0" pageNumber="169">Male (based on holotype, AM KS1128077).</paragraph>
<paragraph id="72253CD07E19D17C675636B8B7D0B588" pageId="0" pageNumber="169">Total length 3.0.</paragraph>
<paragraph id="BF9F4F459BE4A92BDF4F552CA8A0F68E" pageId="0" pageNumber="169">Prosoma. Length 1.7, width 1.2; carapace dark grey; with distinct broad lighter marginal band and broad lighter central band, widening towards eye region (Fig. 17A); sternum light brown, margin slightly grey (Fig. 17B).</paragraph>
<paragraph id="FA4A9E476CA28C91186EBCA4CDFE8164" pageId="0" pageNumber="169">Eyes. Diameter of AME: 0.07; ALE: 0.08; PME: 0.23; PLE: 0.16.</paragraph>
<paragraph id="3C32602CAE33EE2B0D81FF0823A3054E" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, evenly spaced.</paragraph>
<paragraph id="E224480A2C70028B1C3AB3C891EAB8F0" pageId="0" pageNumber="169">Chelicerae. Dark brown.</paragraph>
<paragraph id="9E99C237AEF81039861293E2EF2CB85C" pageId="0" pageNumber="169">Labium. Dark brown, with lighter anterior rim (Fig. 17B)</paragraph>
<paragraph id="6DE4971F2762B72544F4F2C95C1B392E" pageId="0" pageNumber="169">Pedipalp coxae. Dark brown, with lighter anterior rim (Fig. 17B).</paragraph>
<paragraph id="2085317224F83CAD88877115A00941BE" pageId="0" pageNumber="169">Legs. Tibiae, metatarsi and tarsi of leg I very dark to almost black; other legs light brown, annulated (Fig. 17A).</paragraph>
<paragraph id="70BB04A3D2D0293F713E06106A24D063" pageId="0" pageNumber="169">Opisthosoma. Length 1.3, width 1.0; cinnamon-brown with light yellow-brown anterior cardiac mark and dark grey irregular pattern (Fig. 17A). Venter cinnamon with darker pattern laterally (Fig. 17B); spinnerets dark grey.</paragraph>
<paragraph id="D846D0B50568C481DE12BD129CAB9FD2" pageId="0" pageNumber="169">Pedipalps. Tibia as long as broad; cymbium tip with 4-5 macrosetae (Fig. 17E, F); dorsal scopula patch absent; tegular apophysis distally scooped, deeply indented, bent retrolaterally, basally narrowed to 1/3 width, tip reaching margin of cymbium (Fig. 17E); palea about as long as wide, basoembolic apophysis large, triangular; embolus broad, straight; terminal apophysis broad, retrolaterally bifurcate (Fig. 47H).</paragraph>
<paragraph id="9B5FC0691CB43D1C61F0C3AD0710F587" pageId="0" pageNumber="169">Female (based on AM KS122934).</paragraph>
<paragraph id="BDDF9EB4BF97890EB5E0A5B93411274B" pageId="0" pageNumber="169">Total length 5.3.</paragraph>
<paragraph id="D79EA091B5DB4B9A3CDCD42830C7C4C7" pageId="0" pageNumber="169">Prosoma. Length 2.5, width 1.8; carapace and sternum colouration as male (Fig. 17C, D).</paragraph>
<paragraph id="FD08802165954DC66E5ED3B571AA922D" pageId="0" pageNumber="169">Eyes. Diameter of AME 0.10, ALE 0.14, PME 0.26, PLE 0.17.</paragraph>
<paragraph id="30DFE625B905566D3C9FB0633A80C0FA" pageId="0" pageNumber="169">Anterior eye row. Straight, evenly spaced.</paragraph>
<paragraph id="143B5EC471E16EEBCB8992D42E4B13C3" pageId="0" pageNumber="169">Opisthosoma. Length 2.8, width 2.1; otherwise as male, but legs I not darker, not annulated and opisthosoma pattern more obscure (Fig. 17C, D).</paragraph>
<paragraph id="C9B256AF2F19E1570D64946FEC25DA1E" pageId="0" pageNumber="169">Epigyne slightly wider than long, poorly sclerotised at posterior tips, atrium lighter inverted u-shaped (Fig. 17G); spermathecal heads globular about diameter apart, spermathecal stalks short, attached posteriorly (Fig. 17H).</paragraph>
</subSubSection>
<subSubSection id="40C0DD91AA107D501A302F61DE0FF19A" pageId="0" pageNumber="169" type="life history and habitat preferences">
<paragraph id="5E572AD6F046F06456B4E7689EBD38D0" pageId="0" pageNumber="169">Life history and habitat preferences.</paragraph>
<paragraph id="B7B221C79E783F08C045087F9B46217F" pageId="0" pageNumber="169">
The habitat description with the type material is 'upland
<normalizedToken id="181FF183C1155920307CD0E91F2FA8A4" originalValue="swamp">swamp'</normalizedToken>
. Spiders were found in January suggesting
<taxonomicName id="D44A51F76234AB23F23C984FD5DA4EC8" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria equipalus" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="equipalus">Artoria equipalus</taxonomicName>
sp. n. to be summer mature.
</paragraph>
</subSubSection>
<subSubSection id="21BB64723C15C68D84DFAC4CD2C896A7" pageId="0" pageNumber="169" type="distribution">
<paragraph id="8066D0C5323AC9E38524577AB80F1AC1" pageId="0" pageNumber="169">Distribution.</paragraph>
<paragraph id="8D9470BC0EB6AF70F5841EA178EF0C7B" pageId="0" pageNumber="169">
<taxonomicName id="11889F24E46AF19D8DB494E179AF7BFC" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria equipalus" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="equipalus">Artoria equipalus</taxonomicName>
sp. n. is known only from its type locality in Barrington Tops National Park in the NSW North Coast (NNC) IBRA region (Fig. 15).
</paragraph>
<caption id="AB2FF456384665715B7A7D9D8B408628" pageId="0" pageNumber="169">
<paragraph id="4B73A18C6DFF2F60279A109C579CB8D5" pageId="0" pageNumber="169">
Figure 17.
<taxonomicName id="51C40DEF291709EA3A7DE3758829065C" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria equipalus" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="equipalus">Artoria equipalus</taxonomicName>
sp. n., male holotype (AM KS128077), female paratype (AM KS122934): A, male habitus, dorsal view; B, male habitus, ventral view; C, female habitus, dorsal view; D, female habitus, ventral view; E, male pedipalp, ventral view; F, male pedipalp, retrolateral view; G, epigyne, ventral view; H, epigyne, dorsal view. Scale bars: habitus 1.0 mm; pedipalp, epigyne 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>