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<document id="F52FCBF8DE2B2A23C350CC09D444CCF7" ID-CLB-Dataset="32399" ID-DOI="10.11646/zootaxa.4296.1.1" ID-GBIF-Dataset="ad746972-3837-4f1d-bf97-c09e5d7f1bd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="843325" ID-ZooBank="EFE86EDD-D170-4D97-87D9-3125400713EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502790552672" checkinUser="plazi" docAuthor="Filander, Zoleka &amp; Griffiths, Charles" docDate="2017" docId="03FE87F0FF86FFFDFF57FEBFFCD0FC4D" docLanguage="en" docName="zootaxa.4296.1.1.pdf.imf" docOrigin="Zootaxa 4296 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Coelopleurus maillardi" docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFC7FF88FFA7FFDCFFC0FF97FFC9FFF3" masterDocTitle="Illustrated guide to the echinoid (Echinodermata: Echinoidea) fauna of South Africa" masterLastPageNumber="72" masterPageNumber="1" pageNumber="34" updateTime="1698466659567" updateUser="ExternalLinkService">
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<mods:title id="8435C3958E7BCE7E4CE5105D511BD6E0">Illustrated guide to the echinoid (Echinodermata: Echinoidea) fauna of South Africa</mods:title>
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<mods:namePart id="FC05D70A39C064AC154943AFC41E3CC6">Filander, Zoleka</mods:namePart>
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<mods:namePart id="2D9502B3323D02E2588371BAA1D6D059">Griffiths, Charles</mods:namePart>
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<mods:title id="344C7BFFE1E107CBFC39BCCF7317E777">Zootaxa</mods:title>
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<mods:date id="312C4E014C0C3C30120E4AB7623F1999">2017</mods:date>
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<treatment id="03FE87F0FF86FFFDFF57FEBFFCD0FC4D" ID-DOI="http://doi.org/10.5281/zenodo.6022689" ID-GBIF-Taxon="132650422" ID-Zenodo-Dep="6022689" LSID="urn:lsid:plazi:treatment:03FE87F0FF86FFFDFF57FEBFFCD0FC4D" httpUri="http://treatment.plazi.org/id/03FE87F0FF86FFFDFF57FEBFFCD0FC4D" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="C34D656DFF86FFFDFF57FEBFFEE3FE90" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BE836E6FF86FFFDFF57FEBFFD48FEB1" blockId="33.[151,641,296,355]" box="[151,641,296,322]" pageId="33" pageNumber="34">
<heading id="D0A0818AFF86FFFDFF57FEBFFD48FEB1" bold="true" box="[151,641,296,322]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<taxonomicName id="4C574D65FF86FFFDFF57FEBFFD48FEB1" ID-CoL="WSJF" authority="(Michelin, 1862)" baseAuthorityName="Michelin" baseAuthorityYear="1862" box="[151,641,296,322]" class="Echinoidea" family="Arbaciidae" genus="Coelopleurus" kingdom="Animalia" order="Arbacioida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="maillardi">
<emphasis id="B923EAF4FF86FFFDFF57FEBFFD48FEB1" bold="true" box="[151,641,296,322]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF57FEBFFE64FEB1" bold="true" box="[151,429,296,322]" italics="true" pageId="33" pageNumber="34">Coelopleurus maillardi</emphasis>
(
<bibRefCitation id="EFC64B17FF86FFFDFE7DFEBFFDB0FEB1" author="Michelin" box="[445,633,296,322]" pageId="33" pageNumber="34" refString="Michelin, H. (1862) Annex A. Echinides et Stellerides. In: Maillard, L. (Eds.), Notes sur l'Ile de la Reunion (Bourbon). Palais Royal, Galerie D'Orleans, Paris, pp. 1 - 7." type="book chapter" year="1862">Michelin, 1862</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836E6FF86FFFDFF57FEDDFEE3FE90" blockId="33.[151,641,296,355]" box="[151,298,330,355]" pageId="33" pageNumber="34">
<figureCitation id="136C2A63FF86FFFDFF57FEDDFF2FFE90" box="[151,230,330,355]" captionStart="FIGURE 34" captionStartId="33.[151,250,1657,1679]" captionTargetBox="[382,1245,1014,1633]" captionTargetId="figure@33.[340,1247,1013,1635]" captionTargetPageId="33" captionText="FIGURE 34. Coelopleurus maillardi (MBC- 077798). A. Map showing distribution. B. Side view of preserved specimen. C. Close-up view of interambulacra." httpUri="https://zenodo.org/record/843395/files/figure.png" pageId="33" pageNumber="34">Fig. 34</figureCitation>
AC.
</paragraph>
</subSubSection>
<subSubSection id="C34D656DFF86FFFDFF57FE05FDEDFE15" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8BE836E6FF86FFFDFF57FE05FDEDFE15" blockId="33.[151,1436,402,486]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB4711C8FF86FFFDFF57FE05FDEDFE15" pageId="33" pageNumber="34">
<taxonomicName id="4C574D65FF86FFFDFF57FE05FDB6FE5B" ID-CoL="WSJF" authority=": A. Agassiz, 1872: 104" authorityName=": A. Agassiz" authorityPageNumber="104" authorityYear="1872" box="[151,639,402,424]" class="Echinoidea" family="Arbaciidae" genus="Coelopleurus" kingdom="Animalia" order="Arbacioida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="maillardi">
<emphasis id="B923EAF4FF86FFFDFF57FE05FE4CFE5B" box="[151,389,402,424]" italics="true" pageId="33" pageNumber="34">Coelopleurus maillardi</emphasis>
:
<treatmentCitation id="0AF610F7FF86FFFDFE55FE04FDB6FE5B" author="Agassiz" box="[405,639,402,424]" page="104" pageId="33" pageNumber="34" year="1872">
<bibRefCitation id="EFC64B17FF86FFFDFE55FE04FD82FE5B" author="Agassiz" box="[405,587,402,424]" pageId="33" pageNumber="34" refString="Agassiz, A. (1872) Revision of the Echini. Memoirs of the Museum of Comparative Zoology at Harvard College, 3, 1 - 378." type="journal article" year="1872">A. Agassiz, 1872</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
, 406;
<treatmentCitation id="0AF610F7FF86FFFDFD02FE05FC68FE5B" author="Doderlein" box="[706,929,402,424]" page="182" pageId="33" pageNumber="34" year="1906">
<bibRefCitation id="EFC64B17FF86FFFDFD02FE05FCA4FE5B" author="Doderlein" box="[706,877,402,424]" pageId="33" pageNumber="34" refString="Doderlein, L. (1906) Die Echinoiden der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee- Expedition, 5, 61 - 290." type="journal article" year="1906">Döderlein, 1906</bibRefCitation>
: 182
</treatmentCitation>
. Pl. XLV,
<figureCitation id="136C2A63FF86FFFDFBCEFE05FB85FE5B" box="[1038,1100,402,424]" captionStart="FIGURE 1" captionStartId="6.[151,250,1227,1249]" captionTargetBox="[151,1430,448,1187]" captionTargetId="figure@6.[151,1436,445,1206]" captionTargetPageId="6" captionText="FIGURE 1. Detail of regular echinoid: upper (Stomopneustes variolaris) and lower (Stereocidaris excavata) side, showing terminology of various body parts." httpUri="https://zenodo.org/record/843327/files/figure.png" pageId="33" pageNumber="34">Fig. 1</figureCitation>
;
<treatmentCitation id="0AF610F7FF86FFFDFB9BFE04FA88FE5B" author="Mortensen" box="[1115,1345,402,424]" page="627" pageId="33" pageNumber="34" year="1935">
<bibRefCitation id="EFC64B17FF86FFFDFB9BFE04FAC4FE5B" author="Mortensen" box="[1115,1293,402,424]" pageId="33" pageNumber="34" refString="Mortensen, T., (1935) A Monograph of the Echiniodea. II. Bothriocidarioda, Melonechinoida, Lepidocebtroida, and Stirodonta. Reitzel, Copenhagen, 647 pp." type="book" year="1935">Mortensen, 1935</bibRefCitation>
: 627
</treatmentCitation>
631. Pl. LXVII,
<figureCitation id="136C2A63FF86FFFDFED9FE26FE91FE34" box="[281,344,433,455]" captionStart="FIGURE 3" captionStartId="7.[151,250,1253,1275]" captionTargetBox="[194,1367,185,1223]" captionTargetId="figure@7.[154,1439,158,1263]" captionTargetPageId="7" captionText="FIGURE 3. The body plan of an irregular echinoids or heart urchin (Echinocardium cordatum), showing terminology of different body parts." httpUri="https://zenodo.org/record/843331/files/figure.png" pageId="33" pageNumber="34">Fig. 3</figureCitation>
. Pl. LXVIII,
<figureCitation id="136C2A63FF86FFFDFE21FE26FDE9FE34" box="[481,544,433,455]" captionStart="FIGURE 4" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[170,1416,1376,1667]" captionTargetId="figure@7.[151,1436,1356,1675]" captionTargetPageId="7" captionText="FIGURE 4. Tubercle types: A. perforated and non-crenulated (Stereocidaris alcocki). B. non-perforated and non-crenulated (Stomopneustes variolaris). C. perforated and crenulated (Chaetodiadema africanum)." httpUri="https://zenodo.org/record/843333/files/figure.png" pageId="33" pageNumber="34">Fig. 4</figureCitation>
. Pl. LXIX, Figs
<date id="FFE91026FF86FFFDFD0BFE26FD3DFE34" box="[715,756,433,455]" bridgedPair="-" pageId="33" pageNumber="34">I3.</date>
Pl. LXXXVIII,
<figureCitation id="136C2A63FF86FFFDFC5DFE26FBDBFE34" box="[925,1042,433,455]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStartId-0="24.[151,250,1517,1539]" captionStartId-1="25.[151,250,875,897]" captionTargetBox-0="[372,1246,833,1492]" captionTargetBox-1="[370,1246,194,845]" captionTargetId-0="figure@24.[340,1247,832,1495]" captionTargetId-1="figure@25.[342,1249,193,848]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 22. Phormosoma busarium (SAMC-A 22621). A. Map showing distribution. B. Aboral view of preserved specimen. C. Oral view of preserved specimen." captionText-1="FIGURE 23. Phormosoma placenta africana (SAMC- A 22250). A. Map showing distribution. B. Aboral view of preserved specimen. C. Oral view of preserved specimen." httpUri-0="https://zenodo.org/record/843371/files/figure.png" httpUri-1="https://zenodo.org/record/843373/files/figure.png" pageId="33" pageNumber="34">Figs 2223</figureCitation>
;
<treatmentCitation id="0AF610F7FF86FFFDFBDEFE26FA5EFE34" author="Clark" box="[1054,1431,433,455]" page="229" pageId="33" pageNumber="34" year="1976">
<bibRefCitation id="EFC64B17FF86FFFDFBDEFE26FAAAFE34" author="Clark" box="[1054,1379,433,455]" pageId="33" pageNumber="34" refString="Clark, A. &amp; Courtman-Stock, J. (1976) The Echinoderms of Southern Africa. Publication no. 766. British Museum of Natural History, London, 277 pp." type="book" year="1976">Clark &amp; Courtman-Stock, 1976</bibRefCitation>
: 229
</treatmentCitation>
;
<treatmentCitation id="0AF610F7FF86FFFDFF07FE47FE4EFE15" author="Schultz" box="[199,391,464,486]" page="126" pageId="33" pageNumber="34" year="2010">
<bibRefCitation id="EFC64B17FF86FFFDFF07FE47FE9FFE15" author="Schultz" box="[199,342,464,486]" pageId="33" pageNumber="34" refString="Schultz, H. (2010) Sea-Urchins, a Guide to Worldwide Shallow Water Species. 3 rd Edition. Scientific Publication, Hemdingen, 484 pp." type="book" year="2010">Schultz, 2010</bibRefCitation>
: 126
</treatmentCitation>
, Figs 229232.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D656DFF86FFFDFF57FD85FB4FFD4C" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BE836E6FF86FFFDFF57FD85FE87FD80" blockId="33.[151,1436,530,958]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF57FD85FE4BFDD8" bold="true" box="[151,386,530,555]" pageId="33" pageNumber="34">Material examined.</emphasis>
MBC-A077794; MBC-A077798; MBC-A077802; MBC-A077803;
<collectionCode id="ED46AE23FF86FFFDFB50FD84FB36FDD8" box="[1168,1279,531,555]" pageId="33" pageNumber="34">SAMC-A</collectionCode>
23724; MBC- A077796; MBC-A077804; MBC-A077797; MBC-A077795; MBC-A077799; MBC-A077800; MBC-A077801; MBC-A077805.
</paragraph>
<paragraph id="8BE836E6FF86FFFDFF07FD15FB4FFD4C" blockId="33.[151,1436,530,958]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF07FD15FEB9FD68" bold="true" box="[199,368,642,667]" pageId="33" pageNumber="34">Identification.</emphasis>
Test creamish; conspicuous naked interambulacra, naked zone pink to light lilac, with yellow patches, bordered by red line. Spines long, fusiform, green and spotted or banded reddish.
</paragraph>
</subSubSection>
<subSubSection id="C34D656DFF86FFFDFF07FD5AFF29FCDD" pageId="33" pageNumber="34" type="description">
<paragraph id="8BE836E6FF86FFFDFF07FD5AFC88FD15" blockId="33.[151,1436,530,958]" box="[199,833,717,742]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF07FD5AFE1BFD15" bold="true" box="[199,466,717,742]" pageId="33" pageNumber="34">Global maximum size.</emphasis>
Maximum test diameter
<quantity id="4CAF9B03FF86FFFDFD2BFD59FCF4FD15" box="[747,829,718,742]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="34" unit="mm" value="40.0">40 mm</quantity>
.
</paragraph>
<paragraph id="8BE836E6FF86FFFDFF07FD65FF29FCDD" blockId="33.[151,1436,530,958]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF07FD65FE7CFCF8" bold="true" box="[199,437,754,779]" pageId="33" pageNumber="34">Global distribution.</emphasis>
East coast of South Africa to Philippines at
<quantity id="4CAF9B03FF86FFFDFC74FD65FBEEFCF9" box="[948,1063,754,778]" metricMagnitude="2" metricUnit="m" metricValue="1.85" metricValueMax="3.0" metricValueMin="0.7" pageId="33" pageNumber="34" unit="m" value="185.0" valueMax="300.0" valueMin="70.0">70300 m</quantity>
depth (
<bibRefCitation id="EFC64B17FF86FFFDFBBDFD65FAF2FCF8" author="Mortensen" box="[1149,1339,754,779]" pageId="33" pageNumber="34" refString="Mortensen, T., (1935) A Monograph of the Echiniodea. II. Bothriocidarioda, Melonechinoida, Lepidocebtroida, and Stirodonta. Reitzel, Copenhagen, 647 pp." type="book" year="1935">Mortensen 1935</bibRefCitation>
;
<bibRefCitation id="EFC64B17FF86FFFDFA88FD65FF1DFCDD" author="Schultz" pageId="33" pageNumber="34" refString="Schultz, H. (2010) Sea-Urchins, a Guide to Worldwide Shallow Water Species. 3 rd Edition. Scientific Publication, Hemdingen, 484 pp." type="book" year="2010">Schultz 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D656DFF86FFFDFF07FCADFCD0FC4D" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BE836E6FF86FFFDFF07FCADFCD0FC4D" blockId="33.[151,1436,530,958]" pageId="33" pageNumber="34">
<emphasis id="B923EAF4FF86FFFDFF07FCADFEF2FCA0" bold="true" box="[199,315,826,851]" pageId="33" pageNumber="34">Remarks.</emphasis>
Samples identified through this study extend previously known distribution northwards towards southern
<collectingCountry id="F3407676FF86FFFDFEC3FCCAFE5EFC85" box="[259,407,861,886]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
border. Color may distinguish the two
<taxonomicName id="4C574D65FF86FFFDFCB0FCCAFBCEFC85" box="[880,1031,861,886]" class="Echinoidea" family="Arbaciidae" genus="Coelopleurus" kingdom="Animalia" order="Arbacioida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="B923EAF4FF86FFFDFCB0FCCAFBCEFC85" box="[880,1031,861,886]" italics="true" pageId="33" pageNumber="34">Coelopleurus</emphasis>
</taxonomicName>
species; the naked interambulacra zones of
<taxonomicName id="4C574D65FF86FFFDFF3CFC14FE4AFC68" authorityName=": A. Agassiz" authorityYear="1872" box="[252,387,898,923]" class="Echinoidea" family="Arbaciidae" genus="Coelopleurus" kingdom="Animalia" order="Arbacioida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="maillardi">
<emphasis id="B923EAF4FF86FFFDFF3CFC14FE4AFC68" box="[252,387,898,923]" italics="true" pageId="33" pageNumber="34">C. maillardi</emphasis>
</taxonomicName>
are violet with orange tints bordered by a red line; whereas
<taxonomicName id="4C574D65FF86FFFDFBE6FC14FB0DFC69" authorityName=": H. L. Clark" authorityYear="1923" box="[1062,1220,899,922]" class="Echinoidea" family="Arbaciidae" genus="Coelopleurus" kingdom="Animalia" order="Arbacioida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="interruptus">
<emphasis id="B923EAF4FF86FFFDFBE6FC14FB0DFC69" box="[1062,1220,899,922]" italics="true" pageId="33" pageNumber="34">C. interruptus</emphasis>
</taxonomicName>
has a brown naked patch with transverse violet bands and lateral white lines.
</paragraph>
</subSubSection>
</treatment>
</document>