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<mods:title id="2E345738ECD6F64ADDC6D4F848FCC2D9">Contributions to the knowledge of North American tenebrionids of the subtribe Cylindrinotina (Coleoptera: Tenebrionidae: Helopini)</mods:title>
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<taxonomicName id="4C574D76FFE51E382CE9BCDFDA65FF44" box="[151,260,151,177]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382CE9BCDFDA65FF44" bold="true" box="[151,260,151,177]" italics="true" pageId="2" pageNumber="157">Nalassus</emphasis>
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(s. str.)
<taxonomicName id="4C574D76FFE51E382D19BCDFD861FF47" authority="Mannerheim, 1843" authorityName="Mannerheim" authorityYear="1843" box="[359,768,151,178]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="californicus" status="comb. nov.">
<emphasis id="B923EAE7FFE51E382D19BCDFDA99FF44" bold="true" box="[359,504,151,177]" italics="true" pageId="2" pageNumber="157">californicus</emphasis>
(
<bibRefCitation id="EFC64B04FFE51E382E79BCD0D996FF47" author="Mannerheim" box="[519,759,152,178]" pageId="2" pageNumber="157" refString="Mannerheim, M. le Comte. (1843) Memoire sur la recolte d'Insectes Coleopteres faite em 1842. Bulletin de la Societe Imperiale des naturalistes de Moscou, 16 (1), 70 - 99." type="journal article" year="1843">Mannerheim, 1843</bibRefCitation>
)
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,
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(
<figureCitation id="136C2A70FFE51E382CE1BCF2DA68FF26" box="[159,265,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1875,1897]" captionStartId-1="3.[151,250,1529,1551]" captionStartId-2="4.[151,250,1103,1125]" captionTargetBox-0="[189,1397,754,1850]" captionTargetBox-1="[164,1436,199,1481]" captionTargetBox-2="[257,1317,201,1075]" captionTargetId-0="figure@2.[189,1398,754,1850]" captionTargetId-1="figure@3.[151,1436,171,1508]" captionTargetId-2="figure@4.[252,1323,195,1079]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Nalassus californicus, head, SEM. A—ventral side with well expressed ventral posterior grooves; B—the same, right ventral posterior groove; C—ventral side with poorly expressed ventral posterior grooves; D—the same, right ventral groove. Arrow shows ventral posterior grooves." captionText-1="FIGURE 2. Nalassus californicus. A—lectotype, male; B—upper label; C—lower labels." captionText-2="FIGURE 3. Nalassus californicus, genital structures. A—aedeagus ventrally; B—aedeagus laterally; C—male inner sternite VIII; D—gastral spicula; E—female genital tubes. g—the spemathecal accessory gland; s—spermatheca; bds—basal duct of spermatheca; o—oviduct; v—vagina. Scale bars 1 mm." httpUri-0="https://zenodo.org/record/265053/files/figure.png" httpUri-1="https://zenodo.org/record/265054/files/figure.png" httpUri-2="https://zenodo.org/record/265055/files/figure.png" pageId="2" pageNumber="157">Figs. 13</figureCitation>
)
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<emphasis id="B923EAE7FFE51E382CE9BD4ADA71FEED" box="[151,272,258,280]" italics="true" pageId="2" pageNumber="157">californicus</emphasis>
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: 287
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(
<emphasis id="B923EAE7FFE51E382E5FBD4AD908FEED" box="[545,617,258,280]" italics="true" pageId="2" pageNumber="157">Helops</emphasis>
)
</taxonomicName>
;
<treatmentCitation id="0AF610E4FFE51E382EFCBD4BD848FEED" author="Horn" box="[642,809,258,280]" page="394" pageId="2" pageNumber="157" year="1870">
<bibRefCitation id="EFC64B04FFE51E382EFCBD4BD997FEED" author="Horn" box="[642,758,258,280]" pageId="2" pageNumber="157" refString="Horn, G. H. (1870) Revision of the Tenebrionidae of America, north of Mexico. Transactions of the American Philosophical Society, 14, 64 - 404. http: // dx. doi. org / 10.2307 / 1005214" type="journal article" year="1870">Horn 1870</bibRefCitation>
: 394
</treatmentCitation>
(
<taxonomicName id="4C574D76FFE51E382F42BD4AD8E5FEED" box="[828,900,258,280]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382F42BD4AD8E5FEED" box="[828,900,258,280]" italics="true" pageId="2" pageNumber="157">Helops</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AF610E4FFE51E382FE2BD4ADF25FEED" author="Allard" box="[924,1092,258,280]" page="19" pageId="2" pageNumber="157" year="1876">
<bibRefCitation id="EFC64B04FFE51E382FE2BD4ADF7CFEED" author="Allard" box="[924,1053,258,280]" pageId="2" pageNumber="157" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard 1876</bibRefCitation>
: 19
</treatmentCitation>
(
<taxonomicName id="4C574D76FFE51E382828BD4ADFC4FEED" box="[1110,1189,258,280]" class="Insecta" family="Tenebrionidae" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382828BD4ADFC4FEED" box="[1110,1189,258,280]" italics="true" pageId="2" pageNumber="157">Omalus</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AF610E4FFE51E3828C0BD4ADE06FEED" author="Allard" box="[1214,1383,258,280]" page="37" pageId="2" pageNumber="157" year="1877">
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: 37
</treatmentCitation>
, 158 (
<taxonomicName id="4C574D76FFE51E382CB1BD69DA42FEC2" box="[207,291,289,311]" class="Insecta" family="Tenebrionidae" genus="Omaleis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382CB1BD69DA42FEC2" box="[207,291,289,311]" italics="true" pageId="2" pageNumber="157">Omaleis</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AF610E4FFE51E382D47BD69DA92FEC2" author="Seidlitz" box="[313,499,289,311]" page="695" pageId="2" pageNumber="157" year="1896">
<bibRefCitation id="EFC64B04FFE51E382D47BD69DAA2FEC2" author="Seidlitz" box="[313,451,289,311]" pageId="2" pageNumber="157" refString="Seidlitz, G. (1896) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. von (Eds.), Naturgeschichte der Insecten Deutschlands. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800." type="book chapter" year="1896">Seidlitz 1896</bibRefCitation>
: 695
</treatmentCitation>
(
<taxonomicName id="4C574D76FFE51E382E7CBD69D92BFEC2" box="[514,586,289,311]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382E7CBD69D92BFEC2" box="[514,586,289,311]" italics="true" pageId="2" pageNumber="157">Helops</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AF610E4FFE51E382E21BD69D8C1FEC2" author="Cifuentes-Ruiz" box="[607,928,289,311]" page="194" pageId="2" pageNumber="157" year="2014">
<bibRefCitation id="EFC64B04FFE51E382E21BD69D80EFEC2" author="Cifuentes-Ruiz" box="[607,879,289,311]" pageId="2" pageNumber="157" refString="Cifuentes-Ruiz, P., Zaragoza-Caballero, S., Ochoterena-Booth, H. &amp; Moron, M. A. (2014) A preliminary phylogenetic analysis of the New World Helopini (Coleoptera, Tenebrionidae, Tenebrioninae) indicates the need for profound rearrangements of the classification. ZooKeys, 415, 191 - 216. http: // dx. doi. org / 10.3897 / zookeys. 415.6882" type="journal article" year="2014">
Cifuentes-Ruiz
<emphasis id="B923EAE7FFE51E382E81BD6AD84AFEC2" box="[767,811,289,311]" italics="true" pageId="2" pageNumber="157">et al</emphasis>
. 2014
</bibRefCitation>
: 194
</treatmentCitation>
, figs. 5E, 6C, F (
<taxonomicName id="4C574D76FFE51E382833BD69DFDEFEC2" box="[1101,1215,289,311]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382833BD69DFDEFEC2" box="[1101,1215,289,311]" italics="true" pageId="2" pageNumber="157">Odocnemis</emphasis>
</taxonomicName>
), fig. 2B (
<taxonomicName id="4C574D76FFE51E382957BD69DE10FEC2" box="[1321,1393,289,311]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382957BD69DE10FEC2" box="[1321,1393,289,311]" italics="true" pageId="2" pageNumber="157">Helops</emphasis>
</taxonomicName>
). =
<emphasis id="B923EAE7FFE51E382CD5BD08DA61FEA3" box="[171,256,320,342]" italics="true" pageId="2" pageNumber="157">carolina</emphasis>
<bibRefCitation id="EFC64B04FFE51E382D78BD09DAE8FEA3" author="Manee" box="[262,393,320,342]" pageId="2" pageNumber="157" refString="Manee, A. H. (1924) Ecological observations on Rhynchophora in Southern Pines, N. C. Bulletin of the Brooklyn Entomological Society, 19, 40 - 43." type="journal article" year="1924">Manee 1924</bibRefCitation>
: (
<taxonomicName id="4C574D76FFE51E382DE4BD08DA83FEA3" box="[410,482,320,342]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E382DE4BD08DA83FEA3" box="[410,482,320,342]" italics="true" pageId="2" pageNumber="157">Helops</emphasis>
</taxonomicName>
). The name was synonymized by
<bibRefCitation id="EFC64B04FFE51E382F42BD08D8B1FEA3" author="Steiner" box="[828,976,320,342]" pageId="2" pageNumber="157" refString="Steiner, W. E. (2009) The Helopini (Coleoptera: Tenebrionidae) of Virginia. In: Roble, S. M. &amp; Mitchell, J. C. (Eds), A Lifetime of Contributions to Myriapodology and the Natural History of Virginia: A Festschrift in Honor of Richard L. Hoffman's 80 th Birthday. Virginia Museum of Natural History Special Publication No. 16, Martinsville, VA, pp. 331 - 339." type="book chapter" year="2009">Steiner (2009)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D657EFFE51E382CE9BDCADEFDFE16" pageId="2" pageNumber="157" type="materials_examined">
<paragraph id="8BE836F5FFE51E382CE9BDCADE27FE4A" blockId="2.[151,1437,386,699]" pageId="2" pageNumber="157">
<emphasis id="B923EAE7FFE51E382CE9BDCADA2BFE6E" bold="true" box="[151,330,386,411]" pageId="2" pageNumber="157">
<typeStatus id="54EC8857FFE51E382CE9BDCADBB3FE6E" box="[151,210,386,411]" pageId="2" pageNumber="157">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54EC8857FFE51E382D29BDCBDAA6FE6E" box="[343,455,387,411]" pageId="2" pageNumber="157" type="lectotype">Lectotype</typeStatus>
(♂), designated here, with labels: “
<taxonomicName id="4C574D76FFE51E382F07BDCADE51FE6E" authority="Californ. Esch." authorityName="Californ. Esch." box="[889,1328,386,411]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE51E382F07BDCADF10FE6E" box="[889,1137,386,411]" italics="true" pageId="2" pageNumber="157">Nalassus californicus</emphasis>
Californ. Esch.
</taxonomicName>
”, “
<typeStatus id="54EC8857FFE51E382929BDCBDEECFE6E" box="[1367,1421,387,411]" pageId="2" pageNumber="157">type</typeStatus>
”, square with symbol “♂” and green circle.
<typeStatus id="54EC8857FFE51E382E11BDEFD9BEFE4A" box="[623,735,423,447]" pageId="2" pageNumber="157" type="lectotype">Lectotype</typeStatus>
is deposited in
<collectionCode id="ED46AE30FFE51E382FF1BDEFD887FE4B" box="[911,998,423,446]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="157">ZMMU</collectionCode>
in collection of V. Motshulsky.
</paragraph>
<paragraph id="8BE836F5FFE51E382CB9BD82DEFDFE16" blockId="2.[151,1437,386,699]" box="[199,1436,458,483]" pageId="2" pageNumber="157">
<emphasis id="B923EAE7FFE51E382CB9BD82DAB6FE16" bold="true" box="[199,471,458,483]" pageId="2" pageNumber="157">Other studied material</emphasis>
(prepared for genitalia): 2♂♂ and 2♀♀ with press labels: “California Leconte” (
<collectionCode id="ED46AE30FFE51E382922BD83DEF1FE16" box="[1372,1424,459,483]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="157">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D657EFFE51E392CB9BDA6DAE4F94E" lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157" type="discussion">
<paragraph id="8BE836F5FFE51E382CB9BDA6D84AFD4E" blockId="2.[151,1437,386,699]" pageId="2" pageNumber="157">
<emphasis id="B923EAE7FFE51E382CB9BDA6DA2EFDF2" bold="true" box="[199,335,494,519]" pageId="2" pageNumber="157">Comments.</emphasis>
Victor Motshulsky changed many original labels of other entomologists to his own labels for uniformity in the collection. For example, green label was mark North
<collectingCountry id="F3407665FFE51E382F95BE5ADF2EFDDE" box="[1003,1103,530,555]" name="United States of America" pageId="2" pageNumber="157">America</collectingCountry>
. Additionally, sometimes he changed the generic names according to his understanding of beetle classification. Motshulsky always carefully labeled specimens from the
<typeStatus id="54EC8857FFE51E382DA6BE13D969FD86" box="[472,520,603,627]" pageId="2" pageNumber="157">type</typeStatus>
series as “
<typeStatus id="54EC8857FFE51E382EFABE13D9DBFD86" box="[644,698,603,627]" pageId="2" pageNumber="157">type</typeStatus>
”. So, the name “
<taxonomicName id="4C574D76FFE51E382F02BE12DF0EFD86" box="[892,1135,602,627]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE51E382F02BE12DF0EFD86" box="[892,1135,602,627]" italics="true" pageId="2" pageNumber="157">Nalassus californicus</emphasis>
</taxonomicName>
” (the genus
<taxonomicName id="4C574D76FFE51E38297DBE12DE09FD86" box="[1283,1384,602,627]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAE7FFE51E38297DBE12DE09FD86" box="[1283,1384,602,627]" italics="true" pageId="2" pageNumber="157">Nalassus</emphasis>
</taxonomicName>
was described only in 1854) was pinned later by Motschulsky to the
<typeStatus id="54EC8857FFE51E382F12BE37D8FDFD62" box="[876,924,639,663]" pageId="2" pageNumber="157">type</typeStatus>
specimen of M. Mannerheim, who described the species under name “
<taxonomicName id="4C574D76FFE51E382DD1BEEAD87CFD4E" authority="Eschscholtz" authorityName="Eschscholtz" box="[431,797,674,699]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE51E382DD1BEEAD9EAFD4E" box="[431,651,674,699]" italics="true" pageId="2" pageNumber="157">Helops californicus</emphasis>
Eschscholtz
</taxonomicName>
”.
</paragraph>
<caption id="DF28667DFFE51E382CE9BB1BD91DF853" httpUri="https://zenodo.org/record/265053/files/figure.png" pageId="2" pageNumber="157" targetBox="[189,1397,754,1850]" targetPageId="2">
<paragraph id="8BE836F5FFE51E382CE9BB1BD91DF853" blockId="2.[151,1436,1875,1958]" pageId="2" pageNumber="157">
<emphasis id="B923EAE7FFE51E382CE9BB1BDA75F89D" bold="true" box="[151,276,1875,1897]" pageId="2" pageNumber="157">FIGURE 1.</emphasis>
<taxonomicName id="4C574D76FFE51E382D62BB1BDA96F89C" box="[284,503,1875,1897]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE51E382D62BB1BDA96F89C" box="[284,503,1875,1897]" italics="true" pageId="2" pageNumber="157">Nalassus californicus</emphasis>
</taxonomicName>
, head, SEM. A—ventral side with well expressed ventral posterior grooves; B—the same, right ventral posterior groove; C—ventral side with poorly expressed ventral posterior grooves; D—the same, right ventral groove. Arrow shows ventral posterior grooves.
</paragraph>
</caption>
<caption id="DF28667DFFE41E392CE9B9B1DF48F9FA" box="[151,1065,1529,1551]" httpUri="https://zenodo.org/record/265054/files/figure.png" pageId="3" pageNumber="158" targetBox="[164,1436,199,1481]" targetPageId="3">
<paragraph id="8BE836F5FFE41E392CE9B9B1DF48F9FA" blockId="3.[151,1065,1529,1551]" box="[151,1065,1529,1551]" pageId="3" pageNumber="158">
<emphasis id="B923EAE7FFE41E392CE9B9B1DA73F9FB" bold="true" box="[151,274,1529,1551]" pageId="3" pageNumber="158">FIGURE 2.</emphasis>
<taxonomicName id="4C574D76FFE41E392D67B9B1DA95F9FA" box="[281,500,1529,1551]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE41E392D67B9B1DA95F9FA" box="[281,500,1529,1551]" italics="true" pageId="3" pageNumber="158">Nalassus californicus</emphasis>
</taxonomicName>
. A—lectotype, male; B—upper label; C—lower labels.
</paragraph>
</caption>
<paragraph id="8BE836F5FFE41E392CB9BA7EDAE4F94E" blockId="3.[151,1436,1590,1759]" pageId="3" pageNumber="158">
<taxonomicName id="4C574D76FFE41E392CB9BA7EDADBF9BA" box="[199,442,1590,1615]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B923EAE7FFE41E392CB9BA7EDADBF9BA" box="[199,442,1590,1615]" italics="true" pageId="3" pageNumber="158">Nalassus californicus</emphasis>
</taxonomicName>
is similar to the Palaearctic species
<taxonomicName id="4C574D76FFE41E392F22BA7FDF3AF9BA" authority="Gene, 1839" authorityName="Gene" authorityYear="1839" box="[860,1115,1590,1615]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="genei">
<emphasis id="B923EAE7FFE41E392F22BA7FD8DAF9BB" box="[860,955,1591,1614]" italics="true" pageId="3" pageNumber="158">N. genei</emphasis>
(Gené, 1839)
</taxonomicName>
,
<taxonomicName id="4C574D76FFE41E392817BA7FDEF6F9BA" authority="Kuster, 1850" authorityName="Kuster" authorityYear="1850" box="[1129,1431,1590,1615]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="plebejus">
<emphasis id="B923EAE7FFE41E392817BA7FDF89F9BA" box="[1129,1256,1590,1615]" italics="true" pageId="3" pageNumber="158">N. plebejus</emphasis>
(Küster, 1850)
</taxonomicName>
, and
<taxonomicName id="4C574D76FFE41E392CB9BA13D94AF986" authority="Fairmaire, 1888" authorityName="Fairmaire" authorityYear="1888" box="[199,555,1626,1651]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="B923EAE7FFE41E392CB9BA13DA3CF986" box="[199,349,1626,1651]" italics="true" pageId="3" pageNumber="158">N. pekinensis</emphasis>
(Fairmaire, 1888)
</taxonomicName>
of the nominative subgenus, which are also macropterous and lack a brush of setae on the first male abdominal ventrite. Sometimes posterior ventral grooves of head are poorly developed and unclear (figs. 1C, D).
</paragraph>
</subSubSection>
<subSubSection id="C34D657EFFE41E392CB9BA8EDFD2F92A" box="[199,1203,1734,1759]" pageId="3" pageNumber="158" type="distribution">
<paragraph id="8BE836F5FFE41E392CB9BA8EDFD2F92A" blockId="3.[151,1436,1590,1759]" box="[199,1203,1734,1759]" pageId="3" pageNumber="158">
<emphasis id="B923EAE7FFE41E392CB9BA8EDA01F92A" bold="true" box="[199,352,1734,1759]" pageId="3" pageNumber="158">Distribution.</emphasis>
<collectingCountry id="F3407665FFE41E392D19BA8FDAC2F92A" box="[359,419,1735,1759]" name="United States of America" pageId="3" pageNumber="158">USA</collectingCountry>
: California, Oregon, Washington, Nevada;
<collectingCountry id="F3407665FFE41E392FF9BA8ED8BEF92A" box="[903,991,1734,1759]" name="Canada" pageId="3" pageNumber="158">Canada</collectingCountry>
: British
<collectingCountry id="F3407665FFE41E392843BA8EDFCEF92A" box="[1085,1199,1734,1759]" name="United States of America" pageId="3" pageNumber="158">Columbia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>