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<document id="67E18313DD6D8E1F0D08D90D3E486CC1" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D64267FFD282AA3F2A4C37F949" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paramblopusa sumikawai Tasaku, Ono &amp; Maruyama 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="283" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A657F1EE16190BEA424F870C1A5251E4">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="795B18D42B8EEE89EAF09BB4D34B7AE7" type="personal">
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<mods:namePart id="BE67C7FED883C7183E98E9751D3735EF">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="5B29AFB0662137C1941C337BB16D5635">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="993945FC1CC5B70A224923FD44EAC33C">Ono, Hiroki</mods:namePart>
<mods:affiliation id="99E305EFFDF2A7E09629AD4AE294FBE2">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
</mods:name>
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<mods:namePart id="94DF6889C2A87044F8D14B45F41B0C97">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="4235D1245BE49CCC59E48AE5693C3F7F">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:title id="077739D924792FF963AA3E7A84AFD9BD">Zootaxa</mods:title>
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<mods:part id="9B503C93884D0011940E372F710FAE5F">
<mods:date id="4B9605ACDA82EE6AF79A6E692F1E88BA">2023</mods:date>
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<mods:number id="B57E5C460AA747800084D332A0A5BC80">2023-12-12</mods:number>
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<mods:number id="9FCFC08F84C86F076BA6C477FDE20B83">5383</mods:number>
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<mods:number id="297886F3BD341BFB76F78B3685B049DF">3</mods:number>
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<treatment id="03FE87D64267FFD282AA3F2A4C37F949" ID-DOI="http://doi.org/10.5281/zenodo.10361683" ID-Zenodo-Dep="10361683" LSID="urn:lsid:plazi:treatment:03FE87D64267FFD282AA3F2A4C37F949" httpUri="http://treatment.plazi.org/id/03FE87D64267FFD282AA3F2A4C37F949" lastPageId="35" lastPageNumber="286" pageId="32" pageNumber="283">
<subSubSection id="C34D654B4267FFD182AA3F2A4D06FA1B" pageId="32" pageNumber="283" type="nomenclature">
<paragraph id="8BE836C04267FFD182AA3F2A4D60FA37" blockId="32.[151,891,1402,1501]" box="[151,891,1402,1429]" pageId="32" pageNumber="283">
<heading id="D0A081AC4267FFD182AA3F2A4D60FA37" bold="true" box="[151,891,1402,1429]" fontSize="11" level="1" pageId="32" pageNumber="283" reason="1">
<emphasis id="B923EAD24267FFD182AA3F2A4D60FA37" bold="true" box="[151,891,1402,1429]" pageId="32" pageNumber="283">
<taxonomicName id="4C574D434267FFD182AA3F2A4D37FA37" authority="Tasaku, Ono &amp; Maruyama" authorityName="Tasaku, Ono &amp; Maruyama" authorityYear="2023" box="[151,812,1402,1429]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="sumikawai" status="sp. nov.">
<emphasis id="B923EAD24267FFD182AA3F2A4FCAFA37" bold="true" box="[151,465,1402,1429]" italics="true" pageId="32" pageNumber="283">Paramblopusa sumikawai</emphasis>
Tasaku, Ono &amp; Maruyama
</taxonomicName>
,
<taxonomicNameLabel id="A21057A94267FFD181053F2B4D60FA37" box="[824,891,1403,1429]" pageId="32" pageNumber="283" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE836C04267FFD182AA3FCF4D06FA1B" blockId="32.[151,891,1402,1501]" box="[151,797,1439,1465]" pageId="32" pageNumber="283">[Japanese name: Menashi-nise-nokoba-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B4267FFD182AA3F934F23FA7F" box="[151,312,1475,1501]" pageId="32" pageNumber="283" type="description">
<paragraph id="8BE836C04267FFD182AA3F934F23FA7F" blockId="32.[151,891,1402,1501]" box="[151,312,1475,1501]" pageId="32" pageNumber="283">
(
<figureCitation id="136C2A454267FFD182A23F934F2BFA7F" box="[159,304,1475,1501]" captionStart="FIGURE 21" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[162,1424,191,1909]" captionTargetId="figure-19@33.[152,1435,181,1919]" captionTargetPageId="33" captionText="FIGURE 21 Habitus &amp; genitalia of Paramblopusa spp. A–D, P. borealis; E–H, P. sumikawai sp. n. A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H." figureDoi="http://doi.org/10.5281/zenodo.10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="32" pageNumber="283">Figs. 21E–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4267FFD182AA3C5B4D67F9CF" pageId="32" pageNumber="283" type="materials_examined">
<paragraph id="8BE836C04267FFD182AA3C5B4D67F9CF" blockId="32.[151,1436,1546,2041]" pageId="32" pageNumber="283">
<emphasis id="B923EAD24267FFD182AA3C5B4F5FF986" bold="true" box="[151,324,1546,1573]" pageId="32" pageNumber="283">Type material.</emphasis>
<materialsCitation id="3B3F3C9D4267FFD183773C5B4D7CF9EB" collectingDate="2023-05-02" collectionCode="VIII-X, KUM" collectorName="D. Sumikawa" country="Japan" location="Male" pageId="32" pageNumber="283" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="B923EAD24267FFD183773C5B4FA5F987" bold="true" box="[330,446,1547,1573]" pageId="32" pageNumber="283">
<typeStatus id="54EC88624267FFD183773C5B4FA2F987" box="[330,441,1547,1573]" pageId="32" pageNumber="283" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8E88601B4267FFD183F93C5B4C1BF987" LSID="urn:lsid:plazi:treatment:03FE87D64267FFD282AA3F2A4C37F949:8E88601B4267FFD183F93C5B4C1BF987" box="[452,512,1547,1573]" country="Japan" name="Male" pageId="32" pageNumber="283" stateProvince="Hokkaido">Male</location>
, “
<collectingCountry id="F34076504267FFD1802B3C5B4C47F987" box="[534,604,1547,1573]" name="Japan" pageId="32" pageNumber="283">Japan</collectingCountry>
:
<collectingRegion id="4993F8224267FFD180583C5B4CC8F987" box="[613,723,1547,1573]" country="Japan" name="Hokkaido" pageId="32" pageNumber="283">Hokkaidô</collectingRegion>
/ Samani-tyô, Horoman /
<date id="FFE910004267FFD181C83C5B4A73F987" box="[1013,1128,1547,1573]" pageId="32" pageNumber="283" value="2023-05-02">
<collectingDate id="EFADE9E84267FFD181C83C5B4A73F987" box="[1013,1128,1547,1573]" pageId="32" pageNumber="283" value="2023-05-02">2. V. 2023</collectingDate>
</date>
,
<collectorName id="26A253164267FFD1864F3C5B4B16F987" box="[1138,1293,1547,1573]" pageId="32" pageNumber="283">D. Sumikawa</collectorName>
” (abdominal segments
<collectionCode id="ED46AE054267FFD1833B3C7F4F4FF9EB" box="[262,340,1583,1609]" pageId="32" pageNumber="283">VIII-X</collectionCode>
and aedeagus are dissected) (
<figureCitation id="136C2A454267FFD180A33C7F4D1AF9EB" box="[670,769,1583,1609]" captionStart="FIGURE 21" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[162,1424,191,1909]" captionTargetId="figure-19@33.[152,1435,181,1919]" captionTargetPageId="33" captionText="FIGURE 21 Habitus &amp; genitalia of Paramblopusa spp. A–D, P. borealis; E–H, P. sumikawai sp. n. A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H." figureDoi="http://doi.org/10.5281/zenodo.10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="32" pageNumber="283">Fig. 21E</figureCitation>
) (
<collectionCode id="ED46AE054267FFD181253C7F4D44F9EB" box="[792,863,1583,1609]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="32" pageNumber="283" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3F3C9D4267FFD1814C3C7F4D63F9CF" collectingDate="2023-05-02" collectionCode="KUM" collectorName="D. Sumikawa" country="Japan" location="Male" pageId="32" pageNumber="283" specimenCount="1" specimenCount-female="1" stateProvince="Hokkaido" typeStatus="paratype">
<emphasis id="B923EAD24267FFD1814C3C7F4DFEF9EB" bold="true" box="[881,997,1583,1609]" pageId="32" pageNumber="283">
<typeStatus id="54EC88624267FFD1814C3C7F4DFBF9EB" box="[881,992,1583,1609]" pageId="32" pageNumber="283" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCount id="9D51FD494267FFD181D13C7F4A4BF9EB" box="[1004,1104,1583,1609]" count="1" pageId="32" pageNumber="283" type="female">1 female</specimenCount>
, same data as holotype (head, abdominal segments VIII–X, and aedeagus are dissected) (
<collectionCode id="ED46AE054267FFD181143C034D6BF9CF" box="[809,880,1619,1645]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="32" pageNumber="283" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4267FFD182FA3C264D90F917" pageId="32" pageNumber="283" type="diagnosis">
<paragraph id="8BE836C04267FFD182FA3C264D90F917" blockId="32.[151,1436,1546,2041]" pageId="32" pageNumber="283">
<emphasis id="B923EAD24267FFD182FA3C264F59F932" bold="true" box="[199,322,1654,1680]" pageId="32" pageNumber="283">Diagnosis.</emphasis>
This new species is similar to other species of
<taxonomicName id="4C574D434267FFD181463C274A3BF933" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[891,1056,1655,1681]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD24267FFD181463C274A3BF933" box="[891,1056,1655,1681]" italics="true" pageId="32" pageNumber="283">Paramblopusa</emphasis>
</taxonomicName>
but can be distinguished by the following characteristics: eyes absent, body color yellowish brown.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4267FFD282FA3CEE4B86F9B6" lastPageId="35" lastPageNumber="286" pageId="32" pageNumber="283" type="description">
<paragraph id="8BE836C04267FFD182FA3CEE4F65F85B" blockId="32.[151,1436,1546,2041]" pageId="32" pageNumber="283">
<emphasis id="B923EAD24267FFD182FA3CEE4F42F97A" bold="true" box="[199,345,1726,1752]" pageId="32" pageNumber="283">Description.</emphasis>
Body slender, and flattened. Body color yellowish brown. Head almost circular; eyes absent; antennae longer than head and pronotum combined; segment I longest, as long as segment II and III combined; segment II shorter than I; segments III–VIII as long as wide; segments IX and X slightly wider than long; segment XI about 1.5 times longer than wide. Labrum rectangular and transverse, with about 20 setae. Mandibles symmetric, with 3 or 4 small teeth between apex and median tooth. Maxillary palpi 4 segmented; segment I very short; segment II and III ovular; segment IV thin, as long as II. Labium: prementum without medial seta; ligula unilobed, short; labial palpi 3 articled, segment II thinner than I. Mentum almost triangular, incised at apex. Pronotum flattened and subquadrate. Elytra longer than wide, and almost adherent. Abdomen almost parallel-sided. Legs: tarsi very short; tarsal formula 4-4-5.
</paragraph>
<caption id="DF2866484266FFD082AA3DC74ED0F855" ID-DOI="http://doi.org/10.5281/zenodo.10361653" ID-Zenodo-Dep="10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="33" pageNumber="284" startId="33.[151,250,1943,1967]" targetBox="[162,1424,191,1909]" targetPageId="33" targetType="figure">
<paragraph id="8BE836C04266FFD082AA3DC74ED0F855" blockId="33.[151,1436,1943,2039]" pageId="33" pageNumber="284">
<emphasis id="B923EAD24266FFD082AA3DC74F01F80D" bold="true" box="[151,282,1943,1967]" pageId="33" pageNumber="284">FIGURE 21</emphasis>
Habitus &amp; genitalia of
<taxonomicName id="4C574D434266FFD080323DC74CF5F80D" box="[527,750,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="284" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B923EAD24266FFD080323DC74CBFF80D" box="[527,676,1943,1967]" italics="true" pageId="33" pageNumber="284">Paramblopusa</emphasis>
spp. A
</taxonomicName>
–D,
<taxonomicName id="4C574D434266FFD081263DC84D91F80D" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[795,906,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="284" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B923EAD24266FFD081263DC84D31F80D" box="[795,810,1944,1967]" italics="true" pageId="33" pageNumber="284">P</emphasis>
.
<emphasis id="B923EAD24266FFD081053DC74D91F80D" box="[824,906,1943,1967]" italics="true" pageId="33" pageNumber="284">borealis</emphasis>
</taxonomicName>
; E–H,
<taxonomicName id="4C574D434266FFD081E83DC84A45F80D" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[981,1118,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="284" phylum="Arthropoda" rank="species" species="sumikawai" status="sp. n.">
<emphasis id="B923EAD24266FFD081E83DC84DFFF80D" box="[981,996,1944,1967]" italics="true" pageId="33" pageNumber="284">P</emphasis>
.
<emphasis id="B923EAD24266FFD081CF3DC74A45F80D" box="[1010,1118,1943,1967]" italics="true" pageId="33" pageNumber="284">sumikawai</emphasis>
</taxonomicName>
<emphasis id="B923EAD24266FFD0865B3DC74ABBF80D" bold="true" box="[1126,1184,1943,1967]" pageId="33" pageNumber="284">
<taxonomicNameLabel id="A21057A94266FFD0865B3DC74ABBF80D" box="[1126,1184,1943,1967]" pageId="33" pageNumber="284" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H.
</paragraph>
</caption>
<caption id="DF2866484265FFD382AA3DC74D67F871" ID-DOI="http://doi.org/10.5281/zenodo.10361659" ID-Zenodo-Dep="10361659" httpUri="https://zenodo.org/record/10361659/files/figure.png" pageId="34" pageNumber="285" startId="34.[151,250,1943,1967]" targetBox="[193,1404,190,1911]" targetPageId="34" targetType="figure">
<paragraph id="8BE836C04265FFD382AA3DC74D67F871" blockId="34.[151,1436,1943,2004]" pageId="34" pageNumber="285">
<emphasis id="B923EAD24265FFD382AA3DC74F01F80D" bold="true" box="[151,282,1943,1967]" pageId="34" pageNumber="285">FIGURE 22</emphasis>
Mouth parts of
<taxonomicName id="4C574D434265FFD383FC3DC74CB4F80D" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[449,687,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B923EAD24265FFD383FC3DC74CB4F80D" box="[449,687,1943,1967]" italics="true" pageId="34" pageNumber="285">Paramblopusa borealis</emphasis>
</taxonomicName>
. A, labrum (right side, dorsal view; left side, ventral view (epipharynx)); B, mentum; C, labium; D, maxilla; E, right mandible. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="DF2866484264FFD282AA3EE74C1BFA99" ID-DOI="http://doi.org/10.5281/zenodo.10361663" ID-Zenodo-Dep="10361663" httpUri="https://zenodo.org/record/10361663/files/figure.png" pageId="35" pageNumber="286" startId="35.[151,250,1207,1231]" targetBox="[151,1436,181,1183]" targetPageId="35" targetType="figure">
<paragraph id="8BE836C04264FFD282AA3EE74C1BFA99" blockId="35.[151,1437,1207,1339]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282AA3EE74F01FB6D" bold="true" box="[151,282,1207,1231]" pageId="35" pageNumber="286">FIGURE 23</emphasis>
Distribution and habitat of
<taxonomicName id="4C574D434264FFD280043EE74CD6FB6D" authorityName="Sawada" authorityYear="1971" box="[569,717,1207,1231]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="aokii">
<emphasis id="B923EAD24264FFD280043EE74CD6FB6D" box="[569,717,1207,1231]" italics="true" pageId="35" pageNumber="286">Diaulota aokii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C574D434264FFD280E63EE84D40FB6D" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[731,859,1207,1231]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="decipiens" status="sp. n.">
<emphasis id="B923EAD24264FFD280E63EE84CF6FB6D" box="[731,749,1208,1231]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD280C63EE74D40FB6D" box="[763,859,1207,1231]" italics="true" pageId="35" pageNumber="286">decipiens</emphasis>
</taxonomicName>
<emphasis id="B923EAD24264FFD2815F3EE74D87FB6D" bold="true" box="[866,924,1207,1231]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="A21057A94264FFD2815F3EE74D87FB6D" box="[866,924,1207,1231]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C574D434264FFD281EB3EE84A77FB6D" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[982,1132,1207,1231]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="hokkaidona">
<emphasis id="B923EAD24264FFD281EB3EE84DF3FB6D" box="[982,1000,1208,1231]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD281CB3EE74A77FB6D" box="[1014,1132,1207,1231]" italics="true" pageId="35" pageNumber="286">hokkaidona</emphasis>
</taxonomicName>
. A, known localities in Japan (red ring, type locarity of
<taxonomicName id="4C574D434264FFD283983E8C4FE2FB51" authorityName="Sawada" authorityYear="1971" box="[421,505,1243,1267]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="aokii">
<emphasis id="B923EAD24264FFD283983E8C4FACFB51" box="[421,439,1244,1267]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD283FB3E8B4FE2FB51" box="[454,505,1243,1267]" italics="true" pageId="35" pageNumber="286">aokii</emphasis>
</taxonomicName>
; red circle,
<taxonomicName id="4C574D434264FFD2804B3E8C4CD1FB51" authorityName="Sawada" authorityYear="1971" box="[630,714,1243,1267]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="aokii">
<emphasis id="B923EAD24264FFD2804B3E8C4C93FB51" box="[630,648,1244,1267]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD280AA3E8B4CD1FB51" box="[663,714,1243,1267]" italics="true" pageId="35" pageNumber="286">aokii</emphasis>
</taxonomicName>
; blue square,
<taxonomicName id="4C574D434264FFD281613E8C4DE8FB51" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[860,1011,1243,1267]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="hokkaidona">
<emphasis id="B923EAD24264FFD281613E8C4D75FB51" box="[860,878,1244,1267]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD281403E8B4DE8FB51" box="[893,1011,1243,1267]" italics="true" pageId="35" pageNumber="286">hokkaidona</emphasis>
</taxonomicName>
; yellow triangle,
<taxonomicName id="4C574D434264FFD286963E8C4B30FB51" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[1195,1323,1243,1267]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="decipiens" status="sp. n.">
<emphasis id="B923EAD24264FFD286963E8C4AA6FB51" box="[1195,1213,1244,1267]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD286F63E8B4B30FB51" box="[1227,1323,1243,1267]" italics="true" pageId="35" pageNumber="286">decipiens</emphasis>
</taxonomicName>
<emphasis id="B923EAD24264FFD2870E3E8B4B75FB51" bold="true" box="[1331,1390,1243,1267]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="A21057A94264FFD2870E3E8B4B75FB51" box="[1331,1390,1243,1267]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); B, habitat of
<taxonomicName id="4C574D434264FFD2833D3F504F48FAB5" authorityName="Sawada" authorityYear="1971" box="[256,339,1279,1303]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="aokii">
<emphasis id="B923EAD24264FFD2833D3F504F09FAB5" box="[256,274,1280,1303]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD2831D3EAF4F48FAB5" box="[288,339,1279,1303]" italics="true" pageId="35" pageNumber="286">aokii</emphasis>
</taxonomicName>
(Izu-ôshima); C, habitat of
<taxonomicName id="4C574D434264FFD280493F504D11FAB5" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[628,778,1279,1303]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="hokkaidona">
<emphasis id="B923EAD24264FFD280493F504C9DFAB5" box="[628,646,1280,1303]" italics="true" pageId="35" pageNumber="286">D</emphasis>
.
<emphasis id="B923EAD24264FFD280A93EAF4D11FAB5" box="[660,778,1279,1303]" italics="true" pageId="35" pageNumber="286">hokkaidona</emphasis>
</taxonomicName>
; white arrow, adults of eating barnacle (Nemuro-shi, Hokkaidô, Photographed by Tomoki Kikuchi).
</paragraph>
</caption>
<paragraph id="8BE836C04264FFD282FA3F394B7FFA05" blockId="35.[151,1437,1385,1772]" pageId="35" pageNumber="286">
Male: tergite VIII, apical margin truncate; sternite VIII, apical margin slightly prolonged posteriorly; median lobe short and thick, apical lobe short, slightly pointed (
<figureCitation id="136C2A454264FFD281353FDD4D77FA05" box="[776,876,1421,1447]" captionStart="FIGURE 21" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[162,1424,191,1909]" captionTargetId="figure-19@33.[152,1435,181,1919]" captionTargetPageId="33" captionText="FIGURE 21 Habitus &amp; genitalia of Paramblopusa spp. A–D, P. borealis; E–H, P. sumikawai sp. n. A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H." figureDoi="http://doi.org/10.5281/zenodo.10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="35" pageNumber="286">Fig. 21F</figureCitation>
); apical lobe of paramere slender (
<figureCitation id="136C2A454264FFD286D23FDD4B43FA05" box="[1263,1368,1421,1447]" captionStart="FIGURE 21" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[162,1424,191,1909]" captionTargetId="figure-19@33.[152,1435,181,1919]" captionTargetPageId="33" captionText="FIGURE 21 Habitus &amp; genitalia of Paramblopusa spp. A–D, P. borealis; E–H, P. sumikawai sp. n. A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H." figureDoi="http://doi.org/10.5281/zenodo.10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="35" pageNumber="286">Fig. 21H</figureCitation>
).
</paragraph>
<paragraph id="8BE836C04264FFD282FA3FE14C9EFA4D" blockId="35.[151,1437,1385,1772]" pageId="35" pageNumber="286">
Female: tergite VIII, apical margin truncate; sternite VIII, apical margin rounded; spermatheca small, apical part circular, coiled twice at base (
<figureCitation id="136C2A454264FFD2802A3F854C66FA4D" box="[535,637,1493,1519]" captionStart="FIGURE 21" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[162,1424,191,1909]" captionTargetId="figure-19@33.[152,1435,181,1919]" captionTargetPageId="33" captionText="FIGURE 21 Habitus &amp; genitalia of Paramblopusa spp. A–D, P. borealis; E–H, P. sumikawai sp. n. A &amp; E, Habitus; B &amp; F, median lobe of aedeagus; C &amp; G, spermatheca; D &amp; H, apical lobe of paramere. Scale bars: 1.0 mm for A &amp; E; 0.1 mm for B–D, F–H." figureDoi="http://doi.org/10.5281/zenodo.10361653" httpUri="https://zenodo.org/record/10361653/files/figure.png" pageId="35" pageNumber="286">Fig. 21G</figureCitation>
)
</paragraph>
<paragraph id="8BE836C04264FFD282FA3FA94B86F9B6" blockId="35.[151,1437,1385,1772]" box="[199,1437,1529,1556]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282FA3FA94F65F9B1" bold="true" box="[199,382,1529,1555]" pageId="35" pageNumber="286">Measurements.</emphasis>
(
<typeStatus id="54EC88624264FFD283B13FA94FE2F9B1" box="[396,505,1529,1555]" pageId="35" pageNumber="286" type="holotype">Holotype</typeStatus>
, male): FBL,
<quantity id="4CAF9B254264FFD280A93FA94CE0F9B6" box="[660,763,1529,1556]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="35" pageNumber="286" unit="mm" value="0.84">0.84 mm</quantity>
; HW,
<quantity id="4CAF9B254264FFD2817D3FA94DBCF9B6" box="[832,935,1529,1556]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="35" pageNumber="286" unit="mm" value="0.36">0.36 mm</quantity>
; PL,
<quantity id="4CAF9B254264FFD281DC3FA94A53F9B6" box="[993,1096,1529,1556]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="35" pageNumber="286" unit="mm" value="0.34">0.34 mm</quantity>
; PW,
<quantity id="4CAF9B254264FFD286B43FA94AEBF9B6" box="[1161,1264,1529,1556]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="35" pageNumber="286" unit="mm" value="0.36">0.36 mm</quantity>
; EW,
<quantity id="4CAF9B254264FFD2870F3FA94B82F9B6" box="[1330,1433,1529,1556]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="286" unit="mm" value="0.33">0.33 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4264FFD282FA3C4D4F3EF9F9" pageId="35" pageNumber="286" type="etymology">
<paragraph id="8BE836C04264FFD282FA3C4D4F3EF9F9" blockId="35.[151,1437,1385,1772]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282FA3C4D4F55F995" bold="true" box="[199,334,1565,1591]" pageId="35" pageNumber="286">Etymology.</emphasis>
The specific name is named after Mr. Daisuke Sumikawa, the collector of the
<typeStatus id="54EC88624264FFD286E53C4E4B13F99A" box="[1240,1288,1566,1592]" pageId="35" pageNumber="286">type</typeStatus>
series of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4264FFD282FA3C354A12F965" pageId="35" pageNumber="286" type="discussion">
<paragraph id="8BE836C04264FFD282FA3C354B77F9DD" blockId="35.[151,1437,1385,1772]" box="[199,1388,1637,1664]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282FA3C354F20F9DD" bold="true" box="[199,315,1637,1663]" pageId="35" pageNumber="286">Remarks.</emphasis>
Measurements were taken after the
<typeStatus id="54EC88624264FFD280EC3C354D28F9DD" box="[721,819,1637,1663]" pageId="35" pageNumber="286" type="holotype">holotype</typeStatus>
other than BL because the
<typeStatus id="54EC88624264FFD286593C364ADFF922" box="[1124,1220,1638,1664]" pageId="35" pageNumber="286" type="paratype">paratype</typeStatus>
was dissected.
</paragraph>
<paragraph id="8BE836C04264FFD282FA3CD94A12F965" blockId="35.[151,1437,1385,1772]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282FA3CD94F31F901" bold="true" box="[199,298,1673,1699]" pageId="35" pageNumber="286">Habitat.</emphasis>
This new species was collected from a beach gravel (
<quantity id="4CAF9B254264FFD281453CD94DEDF906" box="[888,1014,1673,1700]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="2.0" metricValueMin="0.15" pageId="35" pageNumber="286" unit="mm" value="10.75" valueMax="20.0" valueMin="1.5">1.5–20 mm</quantity>
in diameter) together with
<taxonomicName id="4C574D434264FFD287233CD94EEEF965" authorityName="Casey" authorityYear="1893" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B923EAD24264FFD287233CD94EEEF965" italics="true" pageId="35" pageNumber="286">Amblopusa brevipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434264FFD283163CFD4C29F965" authorityName="Ahn &amp; Ashe" authorityYear="1996" box="[299,562,1709,1735]" class="Insecta" family="Staphylinidae" genus="Paramblopusa" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B923EAD24264FFD283163CFD4C29F965" box="[299,562,1709,1735]" italics="true" pageId="35" pageNumber="286">Paramblopusa borealis</emphasis>
</taxonomicName>
(personal comments from D. Sumikawa).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4264FFD282FA3C814C37F949" box="[199,556,1745,1772]" pageId="35" pageNumber="286" type="distribution">
<paragraph id="8BE836C04264FFD282FA3C814C37F949" blockId="35.[151,1437,1385,1772]" box="[199,556,1745,1772]" pageId="35" pageNumber="286">
<emphasis id="B923EAD24264FFD282FA3C814F7AF949" bold="true" box="[199,353,1745,1771]" pageId="35" pageNumber="286">Distribution.</emphasis>
<collectingCountry id="F34076504264FFD2835A3C824FB0F94E" box="[359,427,1746,1772]" name="Japan" pageId="35" pageNumber="286">Japan</collectingCountry>
:
<collectingRegion id="4993F8224264FFD2838B3C814C33F949" box="[438,552,1745,1771]" country="Japan" name="Hokkaido" pageId="35" pageNumber="286">Hokkaidô</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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