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<mods:title id="0BEB7BB304F5A9B6E566C0600402B96C">Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India)</mods:title>
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<mods:namePart id="E92278EECA5483E86622F8888064925B">Balasubramanian, C.</mods:namePart>
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<paragraph id="8BE836C76356085FFF38FC03DE90FC62" blockId="1.[151,1001,976,1035]" box="[151,1001,976,1002]" pageId="1" pageNumber="189">
<heading id="D0A081AB6356085FFF38FC03DE90FC62" bold="true" box="[151,1001,976,1002]" fontSize="11" level="1" pageId="1" pageNumber="189" reason="1">
<emphasis id="B923EAD56356085FFF38FC03DE90FC62" bold="true" box="[151,1001,976,1002]" pageId="1" pageNumber="189">
<taxonomicName id="4C574D446356085FFF38FC03DE0EFC62" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[151,887,976,1002]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="189" phylum="Arthropoda" rank="species" species="frequentus" status="comb. nov.">
<emphasis id="B923EAD56356085FFF38FC03DCD2FC62" bold="true" box="[151,427,976,1002]" italics="true" pageId="1" pageNumber="189">Tenuibaetis frequentus</emphasis>
(
<bibRefCitation id="EFC64B366356085FFE15FC03DE16FC62" author="Muller-Liebenau" box="[442,879,976,1002]" pageId="1" pageNumber="189" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental region (Insecta, Ephemeroptera). The Florida Entomologist, 68, 537 - 561. http: // dx. doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
)
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<taxonomicNameLabel id="A21057AE6356085FFCD1FC03DE90FC62" box="[894,1001,976,1002]" pageId="1" pageNumber="189">n. comb.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BE836C76356085FFF38FC21DC74FB83" blockId="1.[151,1001,976,1035]" box="[151,269,1010,1035]" pageId="1" pageNumber="189">
<figureCitation id="136C2A426356085FFF38FC21DC74FB83" box="[151,269,1010,1035]" captionStart-0="FIGURES 1 9" captionStart-1="FIGURES 10 16" captionStart-2="FIGURES 17 21" captionStartId-0="2.[151,264,1660,1683]" captionStartId-1="3.[151,264,1927,1950]" captionStartId-2="4.[151,264,1050,1073]" captionTargetBox-0="[386,1191,456,1628]" captionTargetBox-1="[435,1157,752,1880]" captionTargetBox-2="[369,1213,203,1015]" captionTargetId-0="figure@2.[376,1211,445,1639]" captionTargetId-1="figure@3.[406,1180,733,1907]" captionTargetId-2="figure@4.[333,1253,193,1029]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." captionText-1="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." captionText-2="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri-0="https://zenodo.org/record/243147/files/figure.png" httpUri-1="https://zenodo.org/record/243148/files/figure.png" httpUri-2="https://zenodo.org/record/243149/files/figure.png" pageId="1" pageNumber="189">Figs. 121</figureCitation>
</paragraph>
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<taxonomicName id="4C574D446356085FFF38FBE8DFC9FBD8" ID-CoL="KCY9" authority="Muller-Liebenau &amp; Hubbard, 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[151,688,1082,1104]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="189" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56356085FFF38FBE8DC3FFBD8" box="[151,326,1082,1104]" italics="true" pageId="1" pageNumber="189">Baetis frequentus</emphasis>
<bibRefCitation id="EFC64B366356085FFEE2FBE9DFC9FBD8" author="Muller-Liebenau" box="[333,688,1082,1104]" pageId="1" pageNumber="189" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental region (Insecta, Ephemeroptera). The Florida Entomologist, 68, 537 - 561. http: // dx. doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard, 1985</bibRefCitation>
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<emphasis id="B923EAD56356085FFF38FBAEDC00FB1E" bold="true" box="[151,377,1149,1174]" pageId="1" pageNumber="189">Material examined</emphasis>
.
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larvae and
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larvae and
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imagos,
<collectingCountry id="F34076576356085FFC48FBADD940FB1E" box="[999,1081,1150,1174]" name="India" pageId="1" pageNumber="189">INDIA</collectingCountry>
, Tamil Nadu, Theni, Kurangani stream, tributary of Vaigai River,
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,
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,
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,
<date id="FFE910076356085FFC5AFB77D90EFB35" box="[1013,1143,1188,1213]" pageId="1" pageNumber="189" value="2012-07-30">30.vii.2012</date>
, Colls. Balasubramanian, Kubendran and Selvakumar.
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larva (on slide) and
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, Tamil Nadu, Bas. Vaigai, Riv. Valipparai,
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,
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,
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,
<date id="FFE910076356085FFCB1FB3DDED9FA8F" box="[798,928,1262,1287]" pageId="1" pageNumber="189" value="2012-07-29">29.vii.2012</date>
, Colls. Balasubramanian, Kubendran and Selvakumar.
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larva and
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larvae,
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, Tamil Nadu, Kodaikanal, Bas. Manjalaru, Riv. Mulaiyaru,
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,
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,
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,
<date id="FFE910076356085FFDCBFAEADF9FFADA" box="[612,742,1337,1362]" pageId="1" pageNumber="189" value="2012-07-29">29.vii.2012</date>
, Colls. Balasubramanian, Kubendran and Selvakumar.
</paragraph>
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<emphasis id="B923EAD56356085FFF68FA8EDC09FAFE" bold="true" box="[199,368,1373,1398]" pageId="1" pageNumber="189">Mature larva.</emphasis>
Maximal length: Body 9.0
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; cerci
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; terminal filament
<quantity id="4CAF9B226356085FFB31FA8ED983FAFD" box="[1182,1274,1373,1398]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="189" unit="mm" value="3.5">3.5 mm</quantity>
. Size variable between different populations. Colouration: Head uniformly brown (
<figureCitation id="136C2A426356085FFC1CFA52DE81FA12" box="[947,1016,1409,1434]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 1</figureCitation>
). Thorax brown with a pale yellow transverse band on the posterior half of mesonotum, legs light yellow with dark pattern (
<figureCitation id="136C2A426356085FFBC2FA76D9C8FA36" box="[1133,1201,1445,1470]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 2</figureCitation>
). Abdominal tergites dark brown with paired pale yellow maculae on abdominal tergum IV and abdominal terga IXX pale yellow (
<figureCitation id="136C2A426356085FFAC0FA1ADDDDF98E" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 1</figureCitation>
); sternites pale brown except sternites VIII and IX pale yellow (
<figureCitation id="136C2A426356085FFCD9FA3EDEC2F98E" box="[886,955,1517,1542]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE836C76356085CFF68F9C2DF94FF5C" blockId="1.[151,1437,1149,2010]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="190" pageId="1" pageNumber="189">
<emphasis id="B923EAD56356085FFF68F9C2DC7EF9A2" bold="true" box="[199,263,1553,1578]" pageId="1" pageNumber="189">Head</emphasis>
: Antennae: scape and pedicel bare (
<figureCitation id="136C2A426356085FFD33F9C2DF99F9A2" box="[668,736,1553,1578]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 3</figureCitation>
), light yellow. Labrum (
<figureCitation id="136C2A426356085FFC5EF9C2D94EF9A2" box="[1009,1079,1553,1578]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 4</figureCitation>
): rounded with an anteromedial emargination, dorsal surface with one central seta and a distolateral row of 5 short to long simple setae, abundant short fine setae scattered on the surface, subapical row of feathered setae on anterior margin. Hypopharynx (
<figureCitation id="136C2A426356085FFAE6F98AD8F4F9FA" box="[1353,1421,1625,1650]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 5</figureCitation>
): lingua covered with short thin setae, much longer apically; superlingua with thin setae apically and laterally. Left mandible (
<figureCitation id="136C2A426356085FFEBAF972DC18F932" box="[277,353,1697,1722]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 6</figureCitation>
): with incisors composed of 7 denticles; outer denticle longer than others; prostheca with elongated, thin and comb-shaped structure; margin between prostheca and mola not crenate, almost straight without hump and without setae. Right mandible (
<figureCitation id="136C2A426356085FFD7AF93ADE65F88A" box="[725,796,1769,1794]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 7</figureCitation>
): with incisors composed of 7 denticles; stout prostheca with small denticles; margin between prostheca and mola not crenate, without setae. Maxillae (
<figureCitation id="136C2A426356085FFB5FF8DED841F8AE" box="[1264,1336,1805,1830]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 8</figureCitation>
): with 4 caninae, lacinia with 2 rows of setae, one row with abundant thin setae ending with stout and long setae, second row with 2 long stout dentisetae; palp 2- segmented longer than galea-lacinia, segment 1 equal in length to segment 2, apex of segment 2 rounded with few fine setae. Labium (
<figureCitation id="136C2A426356085FFCE0F8AADEEEF81A" box="[847,919,1913,1938]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="1" pageNumber="189">Fig. 9</figureCitation>
): glossae shorter than paraglossae; inner and apical margins of glossae with long setae; paraglossae apically rounded with 3 rows of long setae; labial palp threesegmented, segment 1 longer than segment 2 and 3 combined; segment 2 slightly produced inward to form a moderately expanded lobe at distal corner, dorsally with a row of 6 medium setae; segment 3 conical, slightly asymmetrical and covered with small and stout setae.
</paragraph>
<paragraph id="8BE836C76355085CFF68FF0CDE6DFE00" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="190">
<emphasis id="B923EAD56355085CFF68FF0CDC59FF70" bold="true" box="[199,288,223,248]" pageId="2" pageNumber="190">Thorax</emphasis>
: Legs (
<figureCitation id="136C2A426355085CFED8FF0CDCB4FF70" box="[375,461,223,248]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 10</figureCitation>
): dorsal margin of femora with one row of robust setae; lateral surface with spatulate setae; villopore on anteromedial corner. Dorsal margin of tibiae with short spatulate setae; ventral margin with short pointed setae. Dorsal margin of tarsi with few short and pointed setae, ventral margin with stout setae increasing in length towards apex. Tarsal claws with a single row of about 12 acute teeth increasing in length towards apex (
<figureCitation id="136C2A426355085CFE94FEBCDCF5FE00" box="[315,396,367,392]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 11</figureCitation>
). Hindwing pads present (
<figureCitation id="136C2A426355085CFD1CFEBCDE7FFE00" box="[691,774,367,392]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 12</figureCitation>
).
</paragraph>
<caption id="DF28664F6355085CFF38F9AFDC02F959" httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="2" pageNumber="190" targetBox="[386,1191,456,1628]" targetPageId="2">
<paragraph id="8BE836C76355085CFF38F9AFDC02F959" blockId="2.[151,1436,1660,1745]" pageId="2" pageNumber="190">
<emphasis id="B923EAD56355085CFF38F9AFDC4EF91A" bold="true" box="[151,311,1660,1683]" pageId="2" pageNumber="190">FIGURES 19</emphasis>
. Larva of
<taxonomicName id="4C574D446355085CFE05F9AFD96BF91A" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[426,1042,1660,1682]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="190" phylum="Arthropoda" rank="species" species="frequentus" status="comb. nov.">
<emphasis id="B923EAD56355085CFE05F9AFDFE8F91A" box="[426,657,1660,1682]" italics="true" pageId="2" pageNumber="190">Tenuibaetis frequentus</emphasis>
(Müller-Liebenau &amp; Hubbard 1985)
</taxonomicName>
<taxonomicNameLabel id="A21057AE6355085CFBB3F9AED902F91A" box="[1052,1147,1661,1682]" pageId="2" pageNumber="190">
<emphasis id="B923EAD56355085CFBB3F9AED902F91A" bold="true" box="[1052,1147,1661,1682]" pageId="2" pageNumber="190">n. comb.</emphasis>
</taxonomicNameLabel>
1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium.
</paragraph>
</caption>
<paragraph id="8BE836C76355085CFF68F92BDFC2F84E" blockId="2.[151,1436,1784,2025]" pageId="2" pageNumber="190">
<emphasis id="B923EAD56355085CFF68F92BDC45F899" bold="true" box="[199,316,1784,1809]" pageId="2" pageNumber="190">Abdomen</emphasis>
: Single lamellate gills present on segments 17 with poorly developed tracheation and serrated margin (
<figureCitation id="136C2A426355085CFF56F8CFDC29F8BD" box="[249,336,1820,1845]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 14</figureCitation>
); gill 1 (
<figureCitation id="136C2A426355085CFE14F8CFDF68F8BD" box="[443,529,1820,1845]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 13</figureCitation>
) and 7 smaller than others. Paraproct (
<figureCitation id="136C2A426355085CFC71F8CFD94FF8BD" box="[990,1078,1820,1845]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 15</figureCitation>
): with patch of notched scales medially; margin with numerous short spines; margin of lateral extension with about 9 short to medium triangular spines. Tergites with scale bases; distal margin with triangular spines (
<figureCitation id="136C2A426355085CFC1DF8B7D97FF8F5" box="[946,1030,1892,1917]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="2" pageNumber="190">Fig. 16</figureCitation>
); sternites with abundant thin setae; without scales and scale bases; posterior margin smooth. Cerci light brown without dark stripe; median terminal filament shorter ¾ of cerci and with lateral hairs.
</paragraph>
</subSubSection>
<subSubSection id="C34D654C6355085DFF68F803D9E1FE47" lastPageId="3" lastPageNumber="191" pageId="2" pageNumber="190" type="description">
<paragraph id="8BE836C76355085DFF68F803D9E1FE47" blockId="2.[151,1436,1784,2025]" lastBlockId="3.[151,1437,151,680]" lastPageId="3" lastPageNumber="191" pageId="2" pageNumber="190">
<emphasis id="B923EAD56355085CFF68F803DC2FF861" bold="true" box="[199,342,2000,2025]" pageId="2" pageNumber="190">Male imago</emphasis>
: Length: male body
<quantity id="4CAF9B226355085CFDE5F802DFDFF860" box="[586,678,2001,2025]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="190" unit="mm" value="8.7">8.7 mm</quantity>
; fore wing
<quantity id="4CAF9B226355085CFC83F803DEF1F860" box="[812,904,2000,2025]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="190" unit="mm" value="7.6">7.6 mm</quantity>
; cerci
<quantity id="4CAF9B226355085CFC79F802D928F860" box="[982,1105,2000,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="190" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">1416 mm</quantity>
(
<figureCitation id="136C2A426355085CFBCCF803D9C0F861" box="[1123,1209,2000,2025]" captionStart="FIGURES 17 21" captionStartId="4.[151,264,1050,1073]" captionTargetBox="[369,1213,203,1015]" captionTargetId="figure@4.[333,1253,193,1029]" captionTargetPageId="4" captionText="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="2" pageNumber="190">Fig. 17</figureCitation>
). Colouration: head (
<figureCitation id="136C2A426354085DFF30FF44DD8DFF38" box="[159,244,151,176]" captionStart="FIGURES 17 21" captionStartId="4.[151,264,1050,1073]" captionTargetBox="[369,1213,203,1015]" captionTargetId="figure@4.[333,1253,193,1029]" captionTargetPageId="4" captionText="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="3" pageNumber="191">Fig. 18</figureCitation>
): antennal scape, pedicel and flagellum medium reddish brown; upper surface of compound eye medium reddish brown; lateral face lighter. Thorax: medium to reddish brown. Legs pale yellow without marking or stripe. Forewings (
<figureCitation id="136C2A426354085DFEB0FF0CDC0BFF70" box="[287,370,223,248]" captionStart="FIGURES 17 21" captionStartId="4.[151,264,1050,1073]" captionTargetBox="[369,1213,203,1015]" captionTargetId="figure@4.[333,1253,193,1029]" captionTargetPageId="4" captionText="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="3" pageNumber="191">Fig. 19</figureCitation>
) hyaline with light brown venation; pterostigma with about 4 cross veins generally reaching subcostal vein; double intercalary veins shorter or slightly longer than half distance between corresponding main veins. Hindwings (
<figureCitation id="136C2A426354085DFEC3FEF4DCC6FEC8" box="[364,447,295,320]" captionStart="FIGURES 17 21" captionStartId="4.[151,264,1050,1073]" captionTargetBox="[369,1213,203,1015]" captionTargetId="figure@4.[333,1253,193,1029]" captionTargetPageId="4" captionText="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="3" pageNumber="191">Fig. 20</figureCitation>
) with an erect costal spur at ¼ length of wing; two longitudinal veins reaching margin, none of them bifurcated. Abdomen: tergites IVI whitish without marking or pattern; tergites VIIX medium brown without marking or pattern; sternite colouration similar to corresponding tergites. Genitalia (
<figureCitation id="136C2A426354085DFA9AFEBCD8F6FE00" box="[1333,1423,367,392]" captionStart="FIGURES 17 21" captionStartId="4.[151,264,1050,1073]" captionTargetBox="[369,1213,203,1015]" captionTargetId="figure@4.[333,1253,193,1029]" captionTargetPageId="4" captionText="FIGURES 17 21. Imago of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia." httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="3" pageNumber="191">Fig. 21</figureCitation>
): segment 1 and 2 completely fused; segment 3 elongated, apically slightly expanded; well-developed sclerotized plate between forceps, as broad as distance between forceps, apically convex without setae.
</paragraph>
</subSubSection>
<subSubSection id="C34D654C6354085DFF68FE0FD817FD20" pageId="3" pageNumber="191" type="diagnosis">
<paragraph id="8BE836C76354085DFF68FE0FD817FD20" blockId="3.[151,1437,151,680]" pageId="3" pageNumber="191">
<emphasis id="B923EAD56354085DFF68FE0FDC43FE7D" bold="true" box="[199,314,476,501]" pageId="3" pageNumber="191">Diagnosis</emphasis>
.
<taxonomicName id="4C574D446354085DFEE8FE0FDEA4FE7D" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[327,989,476,501]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56354085DFEE8FE0FDF3AFE7C" box="[327,579,476,501]" italics="true" pageId="3" pageNumber="191">Tenuibaetis frequentus</emphasis>
(
<bibRefCitation id="EFC64B366354085DFDFEFE0FDEADFE7D" author="Muller-Liebenau" box="[593,980,476,501]" pageId="3" pageNumber="191" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental region (Insecta, Ephemeroptera). The Florida Entomologist, 68, 537 - 561. http: // dx. doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
)
</taxonomicName>
can be differentiated from other species of the genus described so far by the following combination of characters in the larval stage: (i) dark brown tergal color pattern typical of the genus but with a pale yellow transverse band on the posterior half of mesonotum, paired pale yellow maculae on abdominal tergum IV and abdominal terga IXX pale yellow (
<figureCitation id="136C2A426354085DFBD2FD94D9B8FDE8" box="[1149,1217,583,608]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="3" pageNumber="191">Fig. 1</figureCitation>
); (ii) segment 2 of labial palp with a row of 6 long setae (
<figureCitation id="136C2A426354085DFDFCFDBFDFE1FD0C" box="[595,664,620,645]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="3" pageNumber="191">Fig. 9</figureCitation>
); (iii) abdominal gills 17 with poorly developed tracheae and with serrated margin (
<figureCitation id="136C2A426354085DFEFBFD5CDCADFD20" box="[340,468,655,680]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="3" pageNumber="191">Figs 13, 14</figureCitation>
) and (iv) number and size of the spines of the distal margin of paraproct (
<figureCitation id="136C2A426354085DFAA2FD5CD818FD20" box="[1293,1377,655,680]" captionStart="FIGURES 10 16" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[435,1157,752,1880]" captionTargetId="figure@3.[406,1180,733,1907]" captionTargetPageId="3" captionText="FIGURES 10 16. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="3" pageNumber="191">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF28664F6354085DFF38F854D95FF835" httpUri="https://zenodo.org/record/243148/files/figure.png" pageId="3" pageNumber="191" targetBox="[435,1157,752,1880]" targetPageId="3">
<paragraph id="8BE836C76354085DFF38F854D95FF835" blockId="3.[151,1435,1927,1981]" pageId="3" pageNumber="191">
<emphasis id="B923EAD56354085DFF38F854DC2CF815" bold="true" box="[151,341,1927,1950]" pageId="3" pageNumber="191">FIGURES 1016.</emphasis>
Larva of
<taxonomicName id="4C574D446354085DFE12F85BD967F816" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[445,1054,1928,1950]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="frequentus" status="comb. nov.">
<emphasis id="B923EAD56354085DFE12F85BDFDAF816" box="[445,675,1928,1950]" italics="true" pageId="3" pageNumber="191">Tenuibaetis frequentus</emphasis>
(Müller-Liebenau &amp; Hubbard 1985)
</taxonomicName>
<taxonomicNameLabel id="A21057AE6354085DFB88F85BD9FCF815" box="[1063,1157,1928,1949]" pageId="3" pageNumber="191">
<emphasis id="B923EAD56354085DFB88F85BD9FCF815" bold="true" box="[1063,1157,1928,1949]" pageId="3" pageNumber="191">n. comb.</emphasis>
</taxonomicNameLabel>
10, Foreleg; 11, Claw; 12, Hindwing pad; 13, Gill 1; 14, Gill 4; 15, Paraproct; 16, Posterior marginal spines of terga.
</paragraph>
</caption>
<caption id="DF28664F6353085AFF38FBC9DF8EFBD8" httpUri="https://zenodo.org/record/243149/files/figure.png" pageId="4" pageNumber="192" targetBox="[369,1213,203,1015]" targetPageId="4">
<paragraph id="8BE836C76353085AFF38FBC9DF8EFBD8" blockId="4.[151,1436,1050,1104]" pageId="4" pageNumber="192">
<emphasis id="B923EAD56353085AFF38FBC9DC2AFBB8" bold="true" box="[151,339,1050,1073]" pageId="4" pageNumber="192">FIGURES 1721.</emphasis>
Imago of
<taxonomicName id="4C574D446353085AFE15FBC8D976FBB9" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[442,1039,1051,1073]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus" status="comb. nov.">
<emphasis id="B923EAD56353085AFE15FBC8DFE7FBB9" box="[442,670,1051,1073]" italics="true" pageId="4" pageNumber="192">Tenuibaetis frequentus</emphasis>
(Müller-Liebenau &amp; Hubbard 1985)
</taxonomicName>
<taxonomicNameLabel id="A21057AE6353085AFBBAFBC8D908FBB8" box="[1045,1137,1051,1072]" pageId="4" pageNumber="192">
<emphasis id="B923EAD56353085AFBBAFBC8D908FBB8" bold="true" box="[1045,1137,1051,1072]" pageId="4" pageNumber="192">n. comb.</emphasis>
</taxonomicNameLabel>
17, Male; 18, Lateral view of male head; 19, Forewing; 20, Hindwing; 21, Male genitalia.
</paragraph>
</caption>
<subSubSection id="C34D654C6353085AFF68FBAFD9C1F859" pageId="4" pageNumber="192" type="discussion">
<paragraph id="8BE836C76353085AFF68FBAFDDACFA75" blockId="4.[151,1436,1148,2001]" pageId="4" pageNumber="192">
<emphasis id="B923EAD56353085AFF68FBAFDC3CFB1D" bold="true" box="[199,325,1148,1173]" pageId="4" pageNumber="192">Discussion</emphasis>
.
<taxonomicName id="4C574D446353085AFEFBFBAFDCADFB1D" box="[340,468,1148,1173]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFEFBFBAFDCADFB1D" box="[340,468,1148,1173]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
was originally established as a subgenus by
<bibRefCitation id="EFC64B366353085AFC7AFBAED9AEFB1D" author="Kang" box="[981,1239,1148,1173]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">Kang and Yang (1994)</bibRefCitation>
within the genus
<taxonomicName id="4C574D446353085AFF38FB72DDA4FB30" box="[151,221,1185,1208]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFF38FB72DDA4FB30" box="[151,221,1185,1208]" italics="true" pageId="4" pageNumber="192">Baetis</emphasis>
</taxonomicName>
designating
<taxonomicName id="4C574D446353085AFEDDFB72DE1AFB31" authority="Muller-Liebenau 1985" authorityName="Muller-Liebenau" authorityYear="1985" box="[370,867,1184,1209]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="pseudofrequentus">
<emphasis id="B923EAD56353085AFEDDFB72DF20FB31" box="[370,601,1184,1209]" italics="true" pageId="4" pageNumber="192">B. pseudofrequentus</emphasis>
Müller-Liebenau 1985
</taxonomicName>
from
<collectingCountry id="F34076576353085AFC02FB73DE86FB31" box="[941,1023,1184,1209]" name="Taiwan" pageId="4" pageNumber="192">Taiwan</collectingCountry>
as
<typeStatus id="54EC88656353085AFB84FB72D922FB31" box="[1067,1115,1185,1209]" pageId="4" pageNumber="192">type</typeStatus>
species and describing two more species viz.,
<taxonomicName id="4C574D446353085AFEC6FB16DE0BFB55" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[361,882,1220,1245]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="inornatus" subGenus="Tenuibaetis">
<emphasis id="B923EAD56353085AFEC6FB16DCF8FB54" box="[361,385,1221,1244]" italics="true" pageId="4" pageNumber="192">B.</emphasis>
(
<emphasis id="B923EAD56353085AFE3DFB17DF6BFB55" box="[402,530,1220,1245]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
)
<emphasis id="B923EAD56353085AFD8CFB16DFF6FB54" box="[547,655,1221,1244]" italics="true" pageId="4" pageNumber="192">inornatus</emphasis>
<bibRefCitation id="EFC64B366353085AFD39FB16DE0BFB55" author="Kang" box="[662,882,1221,1245]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C574D446353085AFC05FB16D8E0FB55" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[938,1433,1220,1245]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="arduus" subGenus="Tenuibaetis">
<emphasis id="B923EAD56353085AFC05FB16DEBBFB54" box="[938,962,1221,1244]" italics="true" pageId="4" pageNumber="192">B.</emphasis>
(
<emphasis id="B923EAD56353085AFC7BFB17D92DFB55" box="[980,1108,1220,1245]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
)
<emphasis id="B923EAD56353085AFBCBFB17D9C8FB55" box="[1124,1201,1220,1245]" italics="true" pageId="4" pageNumber="192">arduus</emphasis>
<bibRefCitation id="EFC64B366353085AFB16FB16D8E0FB55" author="Kang" box="[1209,1433,1221,1245]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
</taxonomicName>
. The subgenus
<taxonomicName id="4C574D446353085AFE90FB3BDCC6FA89" box="[319,447,1256,1281]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFE90FB3BDCC6FA89" box="[319,447,1256,1281]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
was established based on the following combination of characters: (i) mandible with smooth medial margin; (ii) Conical segment 3 of labial palpus; (iii) presence of villopore on femur and (iv) paraproct with a patch of notched scales (
<bibRefCitation id="EFC64B366353085AFD36FAE2DE16FAC1" author="Kang" box="[665,879,1328,1353]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">
Kang
<emphasis id="B923EAD56353085AFD4AFAE2DE65FAC1" box="[741,796,1328,1353]" italics="true" pageId="4" pageNumber="192">et al</emphasis>
. 1994
</bibRefCitation>
). Subsequently,
<bibRefCitation id="EFC64B366353085AFB9CFAE3D849FAC1" box="[1075,1328,1328,1353]" pageId="4" pageNumber="192" refString="Fujitani, T., Hirowatari, T. &amp; Tanida, K. (2003) Nymphs of Nigrobaetis, Alainites, Labiobaetis, Tenuibaetis and Baetis from Japan (Ephemerotpera: Baetidae): diagnosis and keys for genera and species. In: Gaino, E. (Ed.), Research update on Ephemeroptera &amp; Plecoptera. University of Perugia, Perugia, pp. 127 - 133." type="book chapter">
Fujitani
<emphasis id="B923EAD56353085AFB37FAE2D9AEFAC1" box="[1176,1239,1328,1353]" italics="true" pageId="4" pageNumber="192">et al.</emphasis>
(2003)
</bibRefCitation>
elevated
<taxonomicName id="4C574D446353085AFF38FA87DC6EFAE5" box="[151,279,1364,1389]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFF38FA87DC6EFAE5" box="[151,279,1364,1389]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
to the generic level based on the presence of robust setae with median ridge on the dorsomedian surface of the nymphal femur as the exclusive diagnostic features of this genus.
<bibRefCitation id="EFC64B366353085AFC11FAABD9DCFA19" box="[958,1189,1400,1425]" pageId="4" pageNumber="192" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. http: // dx. doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article">
Fujitani
<emphasis id="B923EAD56353085AFBB3FAAAD92DFA19" box="[1052,1108,1400,1425]" italics="true" pageId="4" pageNumber="192">et al.</emphasis>
(2011)
</bibRefCitation>
associated nymphs of three
<taxonomicName id="4C574D446353085AFF7AFA4FDC2CFA3D" box="[213,341,1436,1461]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFF7AFA4FDC2CFA3D" box="[213,341,1436,1461]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
species with their respective imagos by rearing. They redescribed
<taxonomicName id="4C574D446353085AFB95FA4ED82BFA3D" authority="Gose 1980" authorityName="Gose" authorityYear="1980" box="[1082,1362,1436,1461]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="flexifemora">
<emphasis id="B923EAD56353085AFB95FA4ED9ADFA3D" box="[1082,1236,1436,1461]" italics="true" pageId="4" pageNumber="192">T. flexifemora</emphasis>
Gose 1980
</taxonomicName>
and
<taxonomicName id="4C574D446353085AFA28FA4EDF1DFA51" authority="Muller-Liebenau 1985" authorityName="Muller-Liebenau" authorityYear="1985" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="pseudofrequentus">
<emphasis id="B923EAD56353085AFA28FA4EDC25FA51" italics="true" pageId="4" pageNumber="192">T. pseudofrequentus</emphasis>
Müller-Liebenau 1985
</taxonomicName>
and also provided description of a new species,
<taxonomicName id="4C574D446353085AFB33FA12DDABFA75" authority="Fujitani 2011" authorityName="Fujitani" authorityYear="2011" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="parvipterus">
<emphasis id="B923EAD56353085AFB33FA12D842FA50" box="[1180,1339,1473,1496]" italics="true" pageId="4" pageNumber="192">T. parvipterus</emphasis>
<bibRefCitation id="EFC64B366353085AFAEAFA13DDABFA75" author="Fujitani" pageId="4" pageNumber="192" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. http: // dx. doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">Fujitani 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE836C76353085AFF68F9DBDFC2F8ED" blockId="4.[151,1436,1148,2001]" pageId="4" pageNumber="192">
According to
<bibRefCitation id="EFC64B366353085AFEC4F9DBDF1AF9A9" box="[363,611,1544,1569]" pageId="4" pageNumber="192" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. http: // dx. doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article">
Fujitani
<emphasis id="B923EAD56353085AFE61F9DADF72F9A9" box="[462,523,1544,1569]" italics="true" pageId="4" pageNumber="192">et al.</emphasis>
(2011)
</bibRefCitation>
,
<taxonomicName id="4C574D446353085AFDDBF9DBDF88F9A9" box="[628,753,1544,1569]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFDDBF9DBDF88F9A9" box="[628,753,1544,1569]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
presently encompasses five species:
<taxonomicName id="4C574D446353085AFB02F9DADDA3F9CD" authority="Gose 1980" authorityName="Gose" authorityYear="1980" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="flexifemora">
<emphasis id="B923EAD56353085AFB02F9DAD834F9A9" box="[1197,1357,1544,1569]" italics="true" pageId="4" pageNumber="192">T. flexifemora</emphasis>
(Gose 1980)
</taxonomicName>
,
<taxonomicName id="4C574D446353085AFF45F9FEDF95F9CD" authority="Muller-Liebenau 1985" authorityName="Muller-Liebenau" authorityYear="1985" box="[234,748,1580,1605]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="pseudofrequentus">
<emphasis id="B923EAD56353085AFF45F9FEDCB4F9CD" box="[234,461,1580,1605]" italics="true" pageId="4" pageNumber="192">T. pseudofrequentus</emphasis>
(Müller-Liebenau 1985)
</taxonomicName>
,
<taxonomicName id="4C574D446353085AFD54F9FED92EF9CD" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[763,1111,1580,1605]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B923EAD56353085AFD54F9FEDE1EF9CD" box="[763,871,1580,1605]" italics="true" pageId="4" pageNumber="192">T. arduus</emphasis>
<bibRefCitation id="EFC64B366353085AFCDEF9FED92EF9CD" author="Kang" box="[881,1111,1581,1605]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C574D446353085AFBC9F9FEDDABF9E1" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="B923EAD56353085AFBC9F9FED989F9CC" box="[1126,1264,1581,1604]" italics="true" pageId="4" pageNumber="192">T. inornatus</emphasis>
<bibRefCitation id="EFC64B366353085AFB54F9FEDDABF9E1" author="Kang" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C574D446353085AFE8DF982DFF9F9E1" authority="Fujitani 2011" authorityName="Fujitani" authorityYear="2011" box="[290,640,1616,1641]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="parvipterus">
<emphasis id="B923EAD56353085AFE8DF982DCB3F9E0" box="[290,458,1617,1640]" italics="true" pageId="4" pageNumber="192">T. parvipterus</emphasis>
<bibRefCitation id="EFC64B366353085AFE73F983DFF9F9E1" author="Fujitani" box="[476,640,1616,1641]" pageId="4" pageNumber="192" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. http: // dx. doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">Fujitani 2011</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C574D446353085AFD3AF982DE1AF9E1" box="[661,867,1617,1641]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56353085AFD3AF982DE1AF9E1" box="[661,867,1617,1641]" italics="true" pageId="4" pageNumber="192">Baetis frequentus</emphasis>
</taxonomicName>
was attributed to
<taxonomicName id="4C574D446353085AFBF7F983D9A1F9E1" box="[1112,1240,1616,1641]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFBF7F983D9A1F9E1" box="[1112,1240,1616,1641]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
in the website “
<taxonomicName id="4C574D446353085AFF0FF9A7DC37F905" box="[160,334,1652,1677]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
of the World” (http://www.insecta.bio.spbu.ru/z/Eph-spp/%29
<taxonomicName id="4C574D446353085AFBBFF9A7D9F4F905" box="[1040,1165,1652,1677]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus">frequentus</taxonomicName>
%28
<taxonomicName id="4C574D446353085AFB11F9A7D87FF905" box="[1214,1286,1652,1677]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">Baetis</taxonomicName>
%29.htm, last visited
<date id="FFE910076353085AFF42F94BDCB8F939" box="[237,449,1688,1713]" pageId="4" pageNumber="192" value="2015-02-26">26 February 2015</date>
), but the combination was never formally made. Considering that
<taxonomicName id="4C574D446353085AFB7AF94AD8E5F939" box="[1237,1436,1689,1713]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56353085AFB7AF94AD8E5F939" box="[1237,1436,1689,1713]" italics="true" pageId="4" pageNumber="192">Baetis frequentus</emphasis>
</taxonomicName>
perfectly fits the diagnosis and characters of
<taxonomicName id="4C574D446353085AFD22F96FDE74F95D" box="[653,781,1724,1749]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD56353085AFD22F96FDE74F95D" box="[653,781,1724,1749]" italics="true" pageId="4" pageNumber="192">Tenuibaetis</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="EFC64B366353085AFC1EF96ED906F95D" box="[945,1151,1724,1749]" pageId="4" pageNumber="192" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article">
Kang
<emphasis id="B923EAD56353085AFC5AF96ED95EF95D" box="[1013,1063,1724,1749]" italics="true" pageId="4" pageNumber="192">et al</emphasis>
. (1994)
</bibRefCitation>
and
<bibRefCitation id="EFC64B366353085AFB19F96FD8E1F95D" author="Fujitani" box="[1206,1432,1724,1749]" pageId="4" pageNumber="192" refString="Fujitani, T., Hirowatari, T. &amp; Tanida, K. (2003) Nymphs of Nigrobaetis, Alainites, Labiobaetis, Tenuibaetis and Baetis from Japan (Ephemerotpera: Baetidae): diagnosis and keys for genera and species. In: Gaino, E. (Ed.), Research update on Ephemeroptera &amp; Plecoptera. University of Perugia, Perugia, pp. 127 - 133." type="book chapter" year="2003">
Fujitani
<emphasis id="B923EAD56353085AFABBF96ED834F95D" box="[1300,1357,1724,1749]" italics="true" pageId="4" pageNumber="192">et al.</emphasis>
(2003
</bibRefCitation>
,
<bibRefCitation id="EFC64B366353085AFF38F932DDA9F971" author="Fujitani" box="[151,208,1761,1785]" pageId="4" pageNumber="192" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. http: // dx. doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">2011</bibRefCitation>
) and regarding the close affinity between
<taxonomicName id="4C574D446353085AFD1BF932DE32F971" box="[692,843,1761,1785]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56353085AFD1BF932DE32F971" box="[692,843,1761,1785]" italics="true" pageId="4" pageNumber="192">B. frequentus</emphasis>
</taxonomicName>
and
<emphasis id="B923EAD56353085AFC2AF932D917F971" box="[901,1134,1760,1785]" italics="true" pageId="4" pageNumber="192">
<taxonomicName id="4C574D446353085AFC2AF932D912F971" box="[901,1131,1760,1785]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="pseudofrequentus">T. pseudofrequentus</taxonomicName>
,
</emphasis>
we formally proposed the new combination
<taxonomicName id="4C574D446353085AFECAF8D7D977F895" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[357,1038,1796,1821]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="frequentus">
<emphasis id="B923EAD56353085AFECAF8D7DF1DF895" box="[357,612,1796,1821]" italics="true" pageId="4" pageNumber="192">Tenuibaetis frequentus</emphasis>
(
<bibRefCitation id="EFC64B366353085AFDD9F8D7D97FF895" author="Muller-Liebenau" box="[630,1030,1796,1821]" pageId="4" pageNumber="192" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental region (Insecta, Ephemeroptera). The Florida Entomologist, 68, 537 - 561. http: // dx. doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
)
</taxonomicName>
. The species was already reported from
<collectingCountry id="F34076576353085AFF77F8FBDC6BF8C9" box="[216,274,1832,1857]" name="India" pageId="4" pageNumber="192">India</collectingCountry>
(
<bibRefCitation id="EFC64B366353085AFE8AF8FBDC8EF8C9" author="Balaji" box="[293,503,1832,1857]" pageId="4" pageNumber="192" refString="Balaji, A., Vatheeswaran, M. &amp; Venkataraman, K. (1990) Laboratory observations on the life cycle patterns of two Baetis spp. Geobios, 17 (1), 15 - 17. [India]" type="journal article" year="1990">
Balaji
<emphasis id="B923EAD56353085AFEDBF8FADCC8F8C9" box="[372,433,1832,1857]" italics="true" pageId="4" pageNumber="192">et al.</emphasis>
1990
</bibRefCitation>
;
<bibRefCitation id="EFC64B366353085AFDA8F8FBDE9CF8C9" author="Sivaramakrishnan" box="[519,997,1832,1857]" pageId="4" pageNumber="192" refString="Sivaramakrishnan, K. G. &amp; Venkataraman, K. (1990) Abundance, altitudinal distribution and swarming of Ephemeroptera in Palani hills, South India. In: Campbell, I. C. (Ed.), Mayflies and Stoneflies: Life Histories and Biology. Series Entomologica. Vol. 44. Kluwer Academic Publishers, Dordrecht, pp. 209 - 213." type="book" year="1990">Sivaramakrishnan &amp; Venkataraman 1990</bibRefCitation>
). The genus is known therefore from
<collectingCountry id="F34076576353085AFF38F89EDDA3F8EC" box="[151,218,1869,1892]" name="Japan" pageId="4" pageNumber="192">Japan</collectingCountry>
,
<collectingCountry id="F34076576353085AFF4AF89FDC43F8ED" box="[229,314,1868,1893]" name="Taiwan" pageId="4" pageNumber="192">Taiwan</collectingCountry>
,
<collectingCountry id="F34076576353085AFEEAF89EDCB2F8ED" box="[325,459,1869,1893]" name="Hong Kong" pageId="4" pageNumber="192">Hong Kong</collectingCountry>
,
<collectingCountry id="F34076576353085AFE79F89FDF3DF8ED" box="[470,580,1868,1893]" name="Sri Lanka" pageId="4" pageNumber="192">Sri Lanka</collectingCountry>
and
<collectingCountry id="F34076576353085AFDD5F89FDFCEF8ED" box="[634,695,1868,1893]" name="India" pageId="4" pageNumber="192">India</collectingCountry>
.
</paragraph>
<paragraph id="8BE836C76353085AFF68F8A3D9C1F859" blockId="4.[151,1436,1148,2001]" pageId="4" pageNumber="192">
<emphasis id="B923EAD56353085AFF68F8A3DC5DF801" bold="true" box="[199,292,1904,1929]" pageId="4" pageNumber="192">Ecology</emphasis>
. The larvae were collected in a perennial stream of the Vaigai River basin with
<quantity id="4CAF9B226353085AFB02F8A3D873F800" box="[1197,1290,1904,1929]" metricMagnitude="0" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="4" pageNumber="192" unit="m" value="2.75" valueMax="3.5" valueMin="2.0">23.5 m</quantity>
wide and
<quantity id="4CAF9B226353085AFA2FF8A3DDC0F824" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="192" unit="cm" value="60.0">60 cm</quantity>
depth and medium water current (
<quantity id="4CAF9B226353085AFDF2F847DFDFF825" box="[605,678,1940,1965]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="192" unit="m" value="0.6">0.6 m</quantity>
/sec.) on the eastern part of southern Western Ghats. The water temperature ranged between 22˚C25˚C (seasonal variations) and the pH between 6.5 and 7.4.
</paragraph>
</subSubSection>
</treatment>
</document>