297 lines
33 KiB
XML
297 lines
33 KiB
XML
<document id="006F6EF590A153E4C861972F0ECCB234" ID-DOI="10.5281/zenodo.281113" ID-GBIF-Dataset="6955a475-87ea-4eca-8017-5105bae32001" ID-ISSN="1175-5326" ID-Zenodo-Dep="281113" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460482285279" checkinUser="plazi" docAuthor="Fikáček, Martin" docDate="2012" docId="03F9AF63FFDCFFC5ADCA6FE124A73C32" docLanguage="en" docName="zt03316p068.pdf" docOrigin="Zootaxa 3316" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Georissus (Neogeorissus) smetanai Fikáček, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="66" masterDocId="FFC0D71BFFDCFFC6AD5D696224543F0A" masterDocTitle="A new species of leaf-litter inhabiting Georissus from Borneo (Coleoptera: Hydrophiloidea: Georissidae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="63" updateTime="1698307975117" updateUser="plazi">
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<mods:titleInfo id="5B42994ACD89504589524EFC4FE5B7D1">
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<mods:title id="3CA793C05B9563692810B08EBFC8380A">A new species of leaf-litter inhabiting Georissus from Borneo (Coleoptera: Hydrophiloidea: Georissidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="BE7A12491C9DA61C5B1010415F6E0D63" type="personal">
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<mods:namePart id="D57D417126BCE0F9EC1FF007A1E13D2D">Fikáček, Martin</mods:namePart>
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<mods:title id="7F45F8BB4F6B879A4A09ED3014860448">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="77CB3CAA7C9FFD6670FC819AFAB4CFD7">
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<mods:date id="D4CD9B5BBCF19312953A5ACA521E2304">2012</mods:date>
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<mods:detail id="F72ECB860E4C93A8A498DDCD5D0C7209" type="volume">
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<mods:number id="7936B30F0F26E6F079F4451027C6C791">3316</mods:number>
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<mods:start id="9CA42A1BBB3F106C1E8C119EFDA383EA">63</mods:start>
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<mods:classification id="B14606C738316A5548123AFD959B618F">journal article</mods:classification>
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<mods:identifier id="32E4EAF517CC0ED4210AD8B1A8586DDB" type="DOI">10.5281/zenodo.281113</mods:identifier>
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<mods:identifier id="963B34D471E036B01E8FF4B107A19DDF" type="GBIF-Dataset">6955a475-87ea-4eca-8017-5105bae32001</mods:identifier>
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<mods:identifier id="355039A2BAC8AB5E6D760D8A980CE879" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="80C5D0B394B799B8C0DA2946A9B77A1D" type="Zenodo-Dep">281113</mods:identifier>
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<treatment id="03F9AF63FFDCFFC5ADCA6FE124A73C32" ID-DOI="http://doi.org/10.5281/zenodo.6167752" ID-GBIF-Taxon="119517502" ID-Zenodo-Dep="6167752" LSID="urn:lsid:plazi:treatment:03F9AF63FFDCFFC5ADCA6FE124A73C32" httpUri="http://treatment.plazi.org/id/03F9AF63FFDCFFC5ADCA6FE124A73C32" lastPageId="3" lastPageNumber="66" pageId="0" pageNumber="63">
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<subSubSection id="C34A4DFEFFDCFFC6ADCA6FE124A039B4" pageId="0" pageNumber="63" type="nomenclature">
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<paragraph id="8BEF1E75FFDCFFC6ADCA6FE126F63997" blockId="0.[151,674,1667,1726]" box="[151,674,1667,1693]" pageId="0" pageNumber="63">
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<heading id="D0A7A919FFDCFFC6ADCA6FE126F63997" bold="true" box="[151,674,1667,1693]" fontSize="11" level="1" pageId="0" pageNumber="63" reason="1">
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<emphasis id="B924C267FFDCFFC6ADCA6FE126F63997" bold="true" box="[151,674,1667,1693]" pageId="0" pageNumber="63">
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||
<taxonomicName id="4C5065F6FFDCFFC6ADCA6FE126693997" ID-CoL="3FS35" box="[151,573,1667,1693]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov." subGenus="Neogeorissus">
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<emphasis id="B924C267FFDCFFC6ADCA6FE1255A3997" bold="true" box="[151,270,1667,1693]" italics="true" pageId="0" pageNumber="63">Georissus</emphasis>
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||
(
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<emphasis id="B924C267FFDCFFC6AC426FE125943997" bold="true" box="[287,448,1667,1693]" italics="true" pageId="0" pageNumber="63">Neogeorissus</emphasis>
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||
)
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<emphasis id="B924C267FFDCFFC6AC8C6FE126693997" bold="true" box="[465,573,1667,1693]" italics="true" pageId="0" pageNumber="63">smetanai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2177F1CFFDCFFC6AF186FE126F63997" box="[581,674,1667,1693]" pageId="0" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF1E75FFDCFFC6ADCA6FC424A039B4" blockId="0.[151,674,1667,1726]" box="[151,244,1702,1726]" pageId="0" pageNumber="63">
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<figureCitation id="136B02F0FFDCFFC6ADCA6FC424A039B4" box="[151,244,1702,1726]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1960,1983]" captionTargetBox="[151,1436,697,1939]" captionTargetId="figure@1.[151,1436,697,1939]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Georissus smetanai sp. nov. 1 – 2: habitus of the paratype (1: dorsal view, 2: lateral view); 3: aedeagus of the holotype; 4 – 5: sculpture of dorsal surface, paratype (4: head; 5: pronotum); 6: sculpture of the abdominal ventrites, paratype." httpUri="https://zenodo.org/record/281114/files/figure.png" pageId="0" pageNumber="63">Figs 1-6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A4DFEFFDCFFC7ADCA6F8C25973FDE" lastPageId="1" lastPageNumber="64" pageId="0" pageNumber="63" type="materials_examined">
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<paragraph id="8BEF1E75FFDCFFC6ADCA6F8C20AC3820" blockId="0.[151,1436,1773,2015]" pageId="0" pageNumber="63">
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<emphasis id="B924C267FFDCFFC6ADCA6F8C2562380C" bold="true" box="[151,310,1773,1798]" pageId="0" pageNumber="63">
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<typeStatus id="54EBA0D7FFDCFFC6ADCA6F8C2486380C" box="[151,210,1774,1798]" pageId="0" pageNumber="63">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3475EE5FFDCFFC6AC606F8C25F2380C" box="[317,422,1774,1798]" name="Malaysia" pageId="0" pageNumber="63">Malaysia</collectingCountry>
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, Sabah province, Mt. Kinabalu National Park, Layang Layang env.,
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<quantity id="4CA8B390FFDCFFC6A9F46F8C20AB380C" box="[1193,1279,1774,1798]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="0" pageNumber="63" unit="m" value="2600.0">2600 m</quantity>
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a.s.l. [the village is not mentioned on the locality labels, but the locality was specified on my request by I. Löbl].
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</paragraph>
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<paragraph id="8BEF1E75FFDCFFC7AD9A6E5425973FDE" blockId="0.[151,1436,1773,2015]" lastBlockId="1.[151,1436,151,644]" lastPageId="1" lastPageNumber="64" pageId="0" pageNumber="63">
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||
<emphasis id="B924C267FFDCFFC6AD9A6E54267C3845" bold="true" box="[199,552,1846,1871]" pageId="0" pageNumber="63">
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<typeStatus id="54EBA0D7FFDCFFC6AD9A6E5425563844" box="[199,258,1846,1870]" pageId="0" pageNumber="63">Type</typeStatus>
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material.
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<typeStatus id="54EBA0D7FFDCFFC6ACDD6E5426773845" box="[384,547,1846,1871]" pageId="0" pageNumber="63" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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male (
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<collectionCode id="ED4186B0FFDCFFC6AF246E5426823844" box="[633,726,1846,1870]" pageId="0" pageNumber="63">MNHG</collectionCode>
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): “
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<collectionCode id="ED4186B0FFDCFFC6AFA96E5427013844" box="[756,853,1846,1870]" pageId="0" pageNumber="63">SABAH</collectionCode>
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/ Mt. Kinabalu /
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<quantity id="4CA8B390FFDCFFC6A97C6E54202E3844" box="[1057,1146,1846,1870]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="0" pageNumber="63" unit="m" value="2600.0">2600 m</quantity>
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,
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<date id="FFEE38B5FFDCFFC6A9D56E5420B93844" box="[1160,1261,1846,1870]" pageId="0" pageNumber="63" value="1987-05-02">2.V.1987</date>
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/ Burckhardt – Löbl // 11a [on the opposite side of the label]”.
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<emphasis id="B924C267FFDCFFC6AF866E3827DD3879" bold="true" box="[731,905,1882,1907]" pageId="0" pageNumber="63">
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<typeStatus id="54EBA0D7FFDCFFC6AF866E3827D73879" box="[731,899,1882,1907]" pageId="0" pageNumber="63" type="paratype">PARATYPES</typeStatus>
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:
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</emphasis>
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121 spec. (
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<collectionCode id="ED4186B0FFDCFFC6A94B6E3820393878" box="[1046,1133,1882,1906]" pageId="0" pageNumber="63">MNHG</collectionCode>
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,
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<collectionCode id="ED4186B0FFDCFFC6A9236E3820843878" box="[1150,1232,1882,1906]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="63">NMPC</collectionCode>
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,
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<collectionCode id="ED4186B0FFDCFFC6A9BD6E3921633878" box="[1248,1335,1883,1906]" pageId="0" pageNumber="63">BMNH</collectionCode>
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,
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<collectionCode id="ED4186B0FFDCFFC6A81A6E3821CC3878" box="[1351,1432,1882,1906]" pageId="0" pageNumber="63">KSEM</collectionCode>
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,
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<collectionCode id="ED4186B0FFDCFFC6ADCA6E1D24A7389C" box="[151,243,1919,1942]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="63">NHMW</collectionCode>
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,
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<collectionCode id="ED4186B0FFDCFFC6ADA26E1C2502389C" box="[255,342,1918,1942]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="0" pageNumber="63">ZMUC</collectionCode>
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): same label data as the
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<typeStatus id="54EBA0D7FFDCFFC6AF316E1C2685389C" box="[620,721,1918,1942]" pageId="0" pageNumber="63" type="holotype">holotype</typeStatus>
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; 21 spec. (
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<collectionCode id="ED4186B0FFDCFFC6AE166E1C27F6389C" box="[843,930,1918,1942]" pageId="0" pageNumber="63">MNHG</collectionCode>
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,
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||
<collectionCode id="ED4186B0FFDCFFC6AEF26E1C2050389C" box="[943,1028,1918,1942]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="63">NMPC</collectionCode>
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):
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<collectionCode id="ED4186B0FFDCFFC6A9456E1C2023389C" box="[1048,1143,1918,1942]" pageId="0" pageNumber="63">SABAH</collectionCode>
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||
/ Mt. Kinabalu /
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||
<quantity id="4CA8B390FFDCFFC6A8626E1C21C3389C" box="[1343,1431,1918,1942]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="0" pageNumber="63" unit="m" value="2600.0">2600 m</quantity>
|
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,
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<date id="FFEE38B5FFDCFFC6ADCA6EC024AF38B0" box="[151,251,1954,1978]" pageId="0" pageNumber="63" value="1987-05-01">1.V.1987</date>
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/ Burckhardt – Löbl // 10a [on the opposite side of the label];
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<specimenCount id="9D56D5FCFFDCFFC6AE986EC0204038B0" box="[965,1044,1954,1978]" pageId="0" pageNumber="63" type="male">1 male</specimenCount>
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||
, 6 spec. (
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<collectionCode id="ED4186B0FFDCFFC6A9DD6EC0208338B0" box="[1152,1239,1954,1978]" pageId="0" pageNumber="63">MNHG</collectionCode>
|
||
,
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||
<collectionCode id="ED4186B0FFDCFFC6A9B96EC0216D38B0" box="[1252,1337,1954,1978]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="63">NMPC</collectionCode>
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): “BOR- NEO:
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<collectionCode id="ED4186B0FFDCFFC6ADBD6EA4256B38D4" box="[224,319,1990,2014]" pageId="0" pageNumber="63">SABAH</collectionCode>
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Mt. / Kinabalu Nat. Pk. / Layang Layang /
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<quantity id="4CA8B390FFDCFFC6AE766EA4272838D4" box="[811,892,1990,2014]" metricMagnitude="3" metricUnit="m" metricValue="2.16" pageId="0" pageNumber="63" unit="m" value="2160.0">2160m</quantity>
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,
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<date id="FFEE38B5FFDCFFC6AED56EA427E238D4" box="[904,950,1990,2014]" pageId="0" pageNumber="63">2.V.</date>
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[19]87 / A. Smetana”;
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<specimenCount id="9D56D5FCFFDCFFC6A9E86EA4215738D4" box="[1205,1283,1990,2014]" pageId="0" pageNumber="63" type="male">1 male</specimenCount>
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,
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<specimenCount id="9D56D5FCFFDCFFC6A84D6EA4212038D4" box="[1296,1396,1990,2014]" pageId="0" pageNumber="63" type="female">1 female</specimenCount>
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, 45 spec. (
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<collectionCode id="ED4186B0FFDDFFC7AD8269F525623FA5" box="[223,310,151,175]" pageId="1" pageNumber="64">MNHG</collectionCode>
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,
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<collectionCode id="ED4186B0FFDDFFC7AC1C69F525C23FA5" box="[321,406,151,175]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="64">NMPC</collectionCode>
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): “BORNEO:
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<collectionCode id="ED4186B0FFDDFFC7AF6469F526CC3FA5" box="[569,664,151,175]" pageId="1" pageNumber="64">SABAH</collectionCode>
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Mt. / Kinabalu N. P. Sum- / mit
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<collectingCountry id="F3475EE5FFDDFFC7A95369FA207D3FA5" box="[1038,1065,152,175]" name="Chile" pageId="1" pageNumber="64">Tr</collectingCountry>
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. Pondok Lo- / wii
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<quantity id="4CA8B390FFDDFFC7A9A369F521D83FA5" box="[1278,1420,151,175]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.4" metricValueMin="2.3" pageId="1" pageNumber="64" unit="m" value="2350.0" valueMax="2400.0" valueMin="2300.0">2300-2400m</quantity>
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||
/
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<date id="FFEE38B5FFDDFFC7ADCA69DE248A3FDE" box="[151,222,188,212]" pageId="1" pageNumber="64">28.IV.</date>
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[19]87 A. Smetana”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A4DFEFFDDFFC7AD9A69BD268C3D8E" pageId="1" pageNumber="64" type="diagnosis">
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<paragraph id="8BEF1E75FFDDFFC7AD9A69BD268C3D8E" blockId="1.[151,1436,151,644]" pageId="1" pageNumber="64">
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<emphasis id="B924C267FFDDFFC7AD9A69BD25153FF2" bold="true" box="[199,321,223,248]" pageId="1" pageNumber="64">Diagnosis.</emphasis>
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By the arrangement of the protuberances and depressions on the pronotum,
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<taxonomicName id="4C5065F6FFDDFFC7A9E96982216E3FF2" box="[1204,1338,224,248]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
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<emphasis id="B924C267FFDDFFC7A9E96982216E3FF2" box="[1204,1338,224,248]" italics="true" pageId="1" pageNumber="64">G. smetanai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2177F1CFFDDFFC7A819698221C83FFD" box="[1348,1436,224,247]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
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belongs to the
|
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<taxonomicName id="4C5065F6FFDDFFC7AC666867259E3E16" box="[315,458,260,284]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="laesicollis">
|
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<emphasis id="B924C267FFDDFFC7AC666867259E3E16" box="[315,458,260,284]" italics="true" pageId="1" pageNumber="64">G. laesicollis</emphasis>
|
||
</taxonomicName>
|
||
species group defined by
|
||
<bibRefCitation id="EFC16384FFDDFFC7AFB2686627F43E16" author="Deleve" box="[751,928,260,284]" pageId="1" pageNumber="64" refString="Deleve, J. (1967 a) Notes sur les Georissus d? Afrique et descriptions d? especes nouvelles (Coleoptera Georissidae). Bulletin de l? Institut Royal des Sciences Naturelles de Belgique, 43, 26, 3 - 23." type="journal article" year="1967" yearSuffix="a">Delève (1967a)</bibRefCitation>
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: the pronotum bears a large rhomboid central depression, a pair of oval depression posterolaterally of the central depression, and a protuberance on each side laterally of the oval depressions. It may be easily distinguished from all remaining Oriental taxa of the species group by the combination of following characters: pronotum without large lateral projections; elytra with elevated suture and intervals 2, 4 and 6; intervals 1, 3 and 5 flat, largely eliminated by coarse and densely arranged punctation; and ridge on interval 2 with tubercles situated basally and at midlength, hence appearing angular in lateral view. When the aedeagus of
|
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<taxonomicName id="4C5065F6FFDDFFC7AC1368BF25863EFE" box="[334,466,477,500]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
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<emphasis id="B924C267FFDDFFC7AC1368BF25863EFE" box="[334,466,477,500]" italics="true" pageId="1" pageNumber="64">G. smetanai</emphasis>
|
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</taxonomicName>
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<taxonomicNameLabel id="A2177F1CFFDDFFC7AC8768BF26663EFE" box="[474,562,477,500]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is compared with other species of the
|
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<taxonomicName id="4C5065F6FFDDFFC7AEB868BF20233EFE" box="[997,1143,476,500]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="laesicollis">
|
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<emphasis id="B924C267FFDDFFC7AEB868BF20233EFE" box="[997,1143,476,500]" italics="true" pageId="1" pageNumber="64">G. laesicollis</emphasis>
|
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</taxonomicName>
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group (at least those with known male, as few species are described according to females only), it resembles
|
||
<taxonomicName id="4C5065F6FFDDFFC7A9656B62211B3D1D" authority="Deleve, 1969" authorityName="Deleve" authorityYear="1969" box="[1080,1359,511,536]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="calculus">
|
||
<emphasis id="B924C267FFDDFFC7A9656B6220E73D12" box="[1080,1203,512,536]" italics="true" pageId="1" pageNumber="64">G. calculus</emphasis>
|
||
Delève, 1969
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5065F6FFDDFFC7A8D96B6225D93D36" authority="Deleve, 1972" authorityName="Deleve" authorityYear="1972" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="B924C267FFDDFFC7A8D96B6224BF3D36" italics="true" pageId="1" pageNumber="64">G. inflatus</emphasis>
|
||
Delève, 1972
|
||
</taxonomicName>
|
||
by the continually arcuate lateral margins of parameres and phallobase ca. as long as the parameres; both latter species may be easily distinguished by all intervals being of the same height (in contrast to elevated alternate internals in
|
||
<taxonomicName id="4C5065F6FFDDFFC7ACBA6B0F263E3D8E" box="[487,618,621,644]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
|
||
<emphasis id="B924C267FFDDFFC7ACBA6B0F263E3D8E" box="[487,618,621,644]" italics="true" pageId="1" pageNumber="64">G. smetanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2177F1CFFDDFFC7AF2C6B0F26983D8E" box="[625,716,620,644]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2F4EFDFFDDFFC7ADCA6ECA21DD38D7" httpUri="https://zenodo.org/record/281114/files/figure.png" pageId="1" pageNumber="64" targetBox="[151,1436,697,1939]" targetPageId="1">
|
||
<paragraph id="8BEF1E75FFDDFFC7ADCA6ECA21DD38D7" blockId="1.[151,1436,1960,2013]" pageId="1" pageNumber="64">
|
||
<emphasis id="B924C267FFDDFFC7ADCA6ECA256F38B4" bold="true" box="[151,315,1960,1983]" pageId="1" pageNumber="64">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C5065F6FFDDFFC7AC1F6EC8265F38B5" box="[322,523,1962,1983]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
|
||
<emphasis id="B924C267FFDDFFC7AC1F6EC8265F38B5" box="[322,523,1962,1983]" italics="true" pageId="1" pageNumber="64">Georissus smetanai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B924C267FFDDFFC7AF4F6ECB263738B4" bold="true" box="[530,611,1961,1982]" pageId="1" pageNumber="64">
|
||
<taxonomicNameLabel id="A2177F1CFFDDFFC7AF4F6ECB263738B4" box="[530,611,1961,1982]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1–2: habitus of the paratype (1: dorsal view, 2: lateral view); 3: aedeagus of the holotype; 4–5: sculpture of dorsal surface, paratype (4: head; 5: pronotum); 6: sculpture of the abdominal ventrites, paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2F4EFDFFDEFFC4ADCA6A9B263B3B44" httpUri="https://zenodo.org/record/281115/files/figure.png" pageId="2" pageNumber="65" targetBox="[151,1436,193,999]" targetPageId="2">
|
||
<paragraph id="8BEF1E75FFDEFFC4ADCA6A9B263B3B44" blockId="2.[151,1436,1017,1102]" pageId="2" pageNumber="65">
|
||
<emphasis id="B924C267FFDEFFC4ADCA6A9B256F3B05" bold="true" box="[151,315,1017,1040]" pageId="2" pageNumber="65">FIGURES 7–8.</emphasis>
|
||
Other leaf-litter inhabiting
|
||
<taxonomicName id="4C5065F6FFDEFFC4AF056A9926E93B1A" box="[600,701,1019,1040]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924C267FFDEFFC4AF056A9926E93B1A" box="[600,701,1019,1040]" italics="true" pageId="2" pageNumber="65">Georissus</emphasis>
|
||
</taxonomicName>
|
||
species. 7:
|
||
<taxonomicName id="4C5065F6FFDEFFC4AE6A6A9920DD3B1A" authority="Deleve, 1967" authorityName="Deleve" authorityYear="1967" box="[823,1161,1018,1040]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="lateralis">
|
||
<emphasis id="B924C267FFDEFFC4AE6A6A9927AD3B1A" box="[823,1017,1018,1040]" italics="true" pageId="2" pageNumber="65">Georissus lateralis</emphasis>
|
||
Delève, 1967
|
||
</taxonomicName>
|
||
(paratype, Republic of the Congo, Kindamba, Méya, Bangou forest, 9.xi.1963, coll. IRSNB). 8:
|
||
<taxonomicName id="4C5065F6FFDEFFC4AE3E6D7B20F83B24" authority="Deleve, 1972" authorityName="Deleve" authorityYear="1972" box="[867,1196,1048,1070]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="B924C267FFDEFFC4AE3E6D7B20483B24" box="[867,1052,1048,1070]" italics="true" pageId="2" pageNumber="65">Georissus inflatus</emphasis>
|
||
Delève, 1972
|
||
</taxonomicName>
|
||
(paratype, northern Sri Lanka, Puliyankulam, 6.ii.1970, coll. IRSNB).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34A4DFEFFDEFFC5AD9A6D1627823E8D" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="description">
|
||
<paragraph id="8BEF1E75FFDEFFC4AD9A6D1627853BF2" blockId="2.[151,1437,1140,2028]" pageId="2" pageNumber="65">
|
||
<emphasis id="B924C267FFDEFFC4AD9A6D16250D3B87" bold="true" box="[199,345,1140,1165]" pageId="2" pageNumber="65">Description.</emphasis>
|
||
Body widely oval, rather highly convex. Body length
|
||
<quantity id="4CA8B390FFDEFFC4AE9C6D1620313B86" box="[961,1125,1140,1164]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="2" pageNumber="65" unit="mm" value="1.45" valueMax="1.7" valueMin="1.2">1.20–1.70 mm</quantity>
|
||
(
|
||
<typeStatus id="54EBA0D7FFDEFFC4A9296D16208C3B86" box="[1140,1240,1140,1164]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA8B390FFDEFFC4A9826D16211C3B86" box="[1247,1352,1140,1164]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="65" unit="mm" value="1.45">1.45 mm</quantity>
|
||
), width of head
|
||
<quantity id="4CA8B390FFDEFFC4ADA56DFA25CA3BBA" box="[248,414,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="65" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40–0.60 mm</quantity>
|
||
(
|
||
<typeStatus id="54EBA0D7FFDEFFC4ACED6DFA26403BBA" box="[432,532,1176,1200]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA8B390FFDEFFC4AF436DFA26DD3BBA" box="[542,649,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="65" unit="mm" value="0.55">0.55 mm</quantity>
|
||
), maximum width of pronotum
|
||
<quantity id="4CA8B390FFDEFFC4AEA26DFA20F13BBA" box="[1023,1189,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="2" pageNumber="65" unit="mm" value="0.725" valueMax="0.8" valueMin="0.65">0.65–0.80 mm</quantity>
|
||
(
|
||
<typeStatus id="54EBA0D7FFDEFFC4A9EA6DFA214F3BBA" box="[1207,1307,1176,1200]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA8B390FFDEFFC4A8796DFA21C43BBA" box="[1316,1424,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="65" unit="mm" value="0.7">0.70 mm</quantity>
|
||
), maximum width of elytra 0.85–1.00 mm (
|
||
<typeStatus id="54EBA0D7FFDEFFC4AF336DDE26863BDE" box="[622,722,1212,1236]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA8B390FFDEFFC4AF846DDE27163BDE" box="[729,834,1212,1236]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="65" unit="mm" value="0.95">0.95 mm</quantity>
|
||
). Coloration of elytra and pronotum reddish brown to dark brown, anterior portion of pronotum slightly paler, head dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BEF1E75FFDEFFC4AD9A6C6620093A82" blockId="2.[151,1437,1140,2028]" pageId="2" pageNumber="65">Head. Clypeus with narrow depressed anterior portion divided from posterior portion by low ridges, bearing weak and sparse granulation. Posterior portion subdivided by three longitudinal ridges into “cells”. Frons with a deep rhomboid central depression divided from lateral portions by high ridges; lateral portions further subdivided by two longitudinal ridges on each side. Eyes small, narrowly elongate in lateral view.</paragraph>
|
||
<paragraph id="8BEF1E75FFDEFFC4AD9A6CF62709398E" blockId="2.[151,1437,1140,2028]" pageId="2" pageNumber="65">
|
||
Prothorax. Pronotum 1.2× as wide as long, with maximum width at anterior third; lateral portions only very slightly projecting laterad. Anterior portion with two closely situated submedian ridges delimiting median groove, each side laterally of the ridges with two low and wide bulges. Central portion of pronotum with large and deep rhomboid depression delimited by high ridges, lateral angles of the central depression projecting to high protuberances; posterolaterally of the central depression with one oval depression on each side, laterally of these depressions with a small protuberance bearing few setiferous granules. Posterolateral pits large and conspicuous. Ventral morphology of the prothorax corresponding with
|
||
<taxonomicName id="4C5065F6FFDEFFC4AF9F6F0F2702398F" box="[706,854,1645,1669]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="crenulatus">
|
||
<emphasis id="B924C267FFDEFFC4AF9F6F0F2702398F" box="[706,854,1645,1669]" italics="true" pageId="2" pageNumber="65">G. crenulatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF1E75FFDEFFC4AD9A6FF221D438AE" blockId="2.[151,1437,1140,2028]" pageId="2" pageNumber="65">Mesothorax. Elytra combined as wide as long, 1.7× as long as pronotum; base of elytra ca. as wide as maximum width of pronotum, maximum width of elytra in anterior quarter, median portion of elytra slightly bulging posteriad. Elytral suture and intervals 2, 4 and 6, 7, and 8 elevated into sharp narrow strongly denticulate ridges, ridge on interval 2 projecting into slightly higher bulge basally and at midlength, hence slightly angular in lateral view; intervals 1, 3, and 5 not elevated, largely obsolete among extremely coarse and very closely aggregated punctation; punctation retaining its serial arrangement between suture and first elytral ridge, serial arrangement largely eliminated between more lateral ridges. Lateral portion projecting into small wing-like denticulate projections in anterior quarter. Median pentagonal protuberance of mesoventrite with two large circular pits next to each other.</paragraph>
|
||
<paragraph id="8BEF1E75FFDEFFC4AD9A6ED225D238E6" blockId="2.[151,1437,1140,2028]" pageId="2" pageNumber="65">Metathorax. Metathoracic wings absent in all specimens examined for this character (n=6). Metaventrite ca. as long as mesoventrite.</paragraph>
|
||
<paragraph id="8BEF1E75FFDFFFC5AD9A69F527CE3FDE" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="66">Abdomen with lateral margins of ventrites 1 and 2 strongly narrowing posteriad. Ventrites 1 and 2 bearing scattered setiferous tubercles; ventrites 1 and 2 without any large bulges.</paragraph>
|
||
<paragraph id="8BEF1E75FFDFFFC5AD9A69BD27823E8D" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="66">
|
||
Aedeagus
|
||
<quantity id="4CA8B390FFDFFFC5AC6369BD25B63FFD" box="[318,482,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="3" pageNumber="66" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
|
||
long (
|
||
<typeStatus id="54EBA0D7FFDFFFC5AF7669BD26DB3FFD" box="[555,655,223,247]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA8B390FFDFFFC5AFC569BD26A73FFD" box="[664,755,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="66" unit="mm" value="0.4">0.4 mm</quantity>
|
||
). Parameres ca. as long as phallobase, when combined their base slightly narrower than phallobase; lateral margins of parameres nearly parallel-sided in basal 0.8, arcuately bent inward apically, inner margins straight, well sclerotized. Median lobe narrowly triangular, gonopore situated subapically, struts ca. 0.4× as long as apical portion of median lobe. Phallobase slightly widening posteriad, widely open posteriorly, basal foramen occupying posterior 0.4 of the phallobase.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A4DFEFFDFFFC5AD9A68F620403EC5" pageId="3" pageNumber="66" type="etymology">
|
||
<paragraph id="8BEF1E75FFDFFFC5AD9A68F620403EC5" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="66">
|
||
<emphasis id="B924C267FFDFFFC5AD9A68F6251A3EA6" bold="true" box="[199,334,404,428]" pageId="3" pageNumber="66">Etymology.</emphasis>
|
||
This species is dedicated to one of its collectors, Aleš Smetana, on the occasion of his 80th birthday, and as my thanks for his kind support of my studies of the Hydrophiloidea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A4DFEFFDFFFC5AD9A68B921223DE5" pageId="3" pageNumber="66" type="biology_ecology">
|
||
<paragraph id="8BEF1E75FFDFFFC5AD9A68B921223DE5" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="66">
|
||
<emphasis id="B924C267FFDFFFC5AD9A68B925733EFE" bold="true" box="[199,295,475,500]" pageId="3" pageNumber="66">Biology.</emphasis>
|
||
All known specimens of
|
||
<taxonomicName id="4C5065F6FFDFFFC5AF1F68BF26913EFE" box="[578,709,477,500]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
|
||
<emphasis id="B924C267FFDFFFC5AF1F68BF26913EFE" box="[578,709,477,500]" italics="true" pageId="3" pageNumber="66">G. smetanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2177F1CFFDFFFC5AF9668BF27753EFE" box="[715,801,477,500]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were sifted from cloud forest leaf litter under the following circumstances (I. Löbl & A. Smetana, pers. comm., ordered according to collecting date): (i)
|
||
<date id="FFEE38B5FFDFFFC5A9AC689D213E3D1D" box="[1265,1386,511,535]" pageId="3" pageNumber="66" value="1987-04-28">28.iv.1987</date>
|
||
, lgt. Smetana: rather wet moss forest on a steep slope, sifting of fallen leaves with detritus mainly in small forest floor depressions, sometimes including the root system of the vegetation; (ii)
|
||
<date id="FFEE38B5FFDFFFC5AE846B25206F3D55" box="[985,1083,583,607]" pageId="3" pageNumber="66" value="1987-05-01">1.v.1987</date>
|
||
, lgt. Burckhard & Löbl: moist gully below Layang Layang, sifted cloud forest floor litter; (iii)
|
||
<date id="FFEE38B5FFDFFFC5AED96B0E27B23D8E" box="[900,998,620,644]" pageId="3" pageNumber="66" value="1987-05-02">2.v.1987</date>
|
||
, lgt. Burckhard & Löbl: proximity of Layang Layang, sifted moss and moist leaf litter in cloud forest; (iv)
|
||
<date id="FFEE38B5FFDFFFC5AEC36BED20543DAD" box="[926,1024,655,679]" pageId="3" pageNumber="66" value="1987-05-02">2.v.1987</date>
|
||
, lgt. Smetana: sifting various debris, leaf litter and fallen leaves in more or less wet spots in the forest. See below for the discussion on the leaf litter association of this species and its comparison with other
|
||
<taxonomicName id="4C5065F6FFDFFFC5AE496BBA27D03DE5" box="[788,900,728,751]" class="Insecta" family="Georissidae" genus="Georissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924C267FFDFFFC5AE496BBA27D03DE5" box="[788,900,728,751]" italics="true" pageId="3" pageNumber="66">Georissus</emphasis>
|
||
</taxonomicName>
|
||
species collected under such circumstances.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A4DFEFFDFFFC5AD9A6B9924A73C32" pageId="3" pageNumber="66" type="distribution">
|
||
<paragraph id="8BEF1E75FFDFFFC5AD9A6B9924A73C32" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="66">
|
||
<emphasis id="B924C267FFDFFFC5AD9A6B9925343C1E" bold="true" box="[199,352,763,788]" pageId="3" pageNumber="66">Distribution.</emphasis>
|
||
Only known from two areas in Mt. Kinabalu National Park, at elevations between
|
||
<date id="FFEE38B5FFDFFFC5A86C6B9E249B3C3D" pageId="3" pageNumber="66" value="2300" valueMax="2600">
|
||
2300 and
|
||
<quantity id="4CA8B390FFDFFFC5ADCA6A7D249B3C3D" box="[151,207,799,823]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="66" unit="m" value="2600.0">2600</quantity>
|
||
</date>
|
||
m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |