treatments-xml/data/03/F9/87/03F987C08A7B017C3AEE8DA0FBA94A95.xml
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<document id="4CE1C47B4F13E5E3F00B679294958DAC" ID-CLB-Dataset="28135" ID-DOI="10.11646/zootaxa.4563.3.3" ID-GBIF-Dataset="6c280343-607d-4ff5-b2ba-1ed24db90c36" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601342" ID-ZooBank="86E75433-D91B-48E5-9807-9A0A460EFCB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172235077" checkinUser="plazi" docAuthor="Salgado, Alexandre &amp; Ruiz, Gustavo R. S." docDate="2019" docId="03F987C08A7B017C3AEE8DA0FBA94A95" docLanguage="en" docName="zootaxa.4563.3.3.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus araripe Salgado &amp; Ruiz 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="465" masterDocId="FFC0FFB88A7201723A798B0DFFC84C76" masterDocTitle="Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini)" masterLastPageNumber="481" masterPageNumber="451" pageNumber="460" updateTime="1698703444826" updateUser="ExternalLinkService">
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<mods:title id="E85E71E07BEC5C3CF1EF79C3907D0859">Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini)</mods:title>
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<mods:namePart id="085B98A7E4CAAA8590B48255D4F11EB4">Salgado, Alexandre</mods:namePart>
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<mods:namePart id="D5A64219A15186E40A40AAC5ECA529CB">Ruiz, Gustavo R. S.</mods:namePart>
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<treatment id="03F987C08A7B017C3AEE8DA0FBA94A95" ID-DOI="http://doi.org/10.5281/zenodo.5931669" ID-GBIF-Taxon="154474848" ID-Zenodo-Dep="5931669" LSID="urn:lsid:plazi:treatment:03F987C08A7B017C3AEE8DA0FBA94A95" httpUri="http://treatment.plazi.org/id/03F987C08A7B017C3AEE8DA0FBA94A95" lastPageId="14" lastPageNumber="465" pageId="9" pageNumber="460">
<subSubSection id="C34A655D8A7B017B3AEE8DA0FE394AB1" box="[151,497,1709,1735]" pageId="9" pageNumber="460" type="nomenclature">
<paragraph id="8BEF36D68A7B017B3AEE8DA0FE394AB1" blockId="9.[151,497,1709,1768]" box="[151,497,1709,1735]" pageId="9" pageNumber="460">
<heading id="D0A781BA8A7B017B3AEE8DA0FE394AB1" bold="true" box="[151,497,1709,1735]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
<emphasis id="B924EAC48A7B017B3AEE8DA0FE394AB1" bold="true" box="[151,497,1709,1735]" pageId="9" pageNumber="460">
<taxonomicName id="4C504D558A7B017B3AEE8DA0FE444AB1" ID-CoL="66FZW" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[151,396,1709,1735]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7B017B3AEE8DA0FE444AB1" bold="true" box="[151,396,1709,1735]" italics="true" pageId="9" pageNumber="460">Amphidraus araripe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BF8A7B017B3BEA8DA0FE394AB1" box="[403,497,1709,1735]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A7B017B3AEE8DC2FEDB4A9E" box="[151,275,1743,1768]" pageId="9" pageNumber="460" type="description">
<paragraph id="8BEF36D68A7B017B3AEE8DC2FEDB4A9E" blockId="9.[151,497,1709,1768]" box="[151,275,1743,1768]" pageId="9" pageNumber="460">
<figureCitation id="136B2A538A7B017B3AEE8DC2FEDB4A9E" box="[151,275,1743,1768]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStartId-0="11.[151,250,1888,1910]" captionStartId-1="12.[151,250,1946,1968]" captionStartId-2="13.[151,250,1851,1873]" captionStartId-3="14.[151,250,1032,1054]" captionTargetBox-0="[151,1436,193,1867]" captionTargetBox-1="[187,1409,836,1922]" captionTargetBox-2="[198,1418,208,1790]" captionTargetBox-3="[163,1419,197,1010]" captionTargetId-0="figure@11.[151,1436,193,1867]" captionTargetId-1="figure@12.[173,1417,805,1931]" captionTargetId-2="figure@13.[151,1436,193,1829]" captionTargetId-3="figure@14.[163,1419,197,1010]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." captionText-1="FIGURE 12. Amphidraus araripe sp. nov. AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral, D dorsal). Abbreviations: cd, copulatory duct; fd, fertilization duct; gl, gland; S spermatheca. White arrow in C shows copulatory opening." captionText-2="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." captionText-3="FIGURE 14. Map of northern/northeastern Brazil showing records of Amphidraus araripe, A. boomerang, A. cornipalpis and A. sacrificatus. White lines represent the borders of Brazilian states. Map modified from Google Earth ®." httpUri-0="https://zenodo.org/record/2601366/files/figure.png" httpUri-1="https://zenodo.org/record/2601368/files/figure.png" httpUri-2="https://zenodo.org/record/2601370/files/figure.png" httpUri-3="https://zenodo.org/record/2601372/files/figure.png" pageId="9" pageNumber="460">Figs 1114</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A7B017B3AEE8C1AFEB74BEA" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="8BEF36D68A7B017B3AEE8C1AFEB74BEA" blockId="9.[151,1436,1815,1949]" pageId="9" pageNumber="460">
<emphasis id="B924EAC48A7B017B3AEE8C1AFE174B46" bold="true" box="[151,479,1815,1840]" pageId="9" pageNumber="460">
Type material.
<materialsCitation id="3B383C8B8A7B017B3B358C1AFE174B46" ID-GBIF-Occurrence="2013952918" box="[332,479,1815,1840]" collectingDate="2011-02-06" collectionCode="MPEG" collectorName="L. S. Carvalho" country="Brazil" latitude="-7.3110003" location="Floresta Nacional do Araripe-Apodi" longLatPrecision="1" longitude="-39.447002" municipality="Crato" pageId="9" pageNumber="460" specimenCode="MPEG 35011" specimenCount="1" stateProvince="Ceara" typeStatus="holotype">
<typeStatus id="54EB88748A7B017B3B358C1AFE754B46" box="[332,445,1815,1840]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
: Ƌ
</materialsCitation>
</emphasis>
from
<location id="8E8F600D8A7B017B385D8C1AFC754B46" LSID="urn:lsid:plazi:treatment:03F987C08A7B017C3AEE8DA0FBA94A95:8E8F600D8A7B017B385D8C1AFC754B46" box="[548,957,1815,1840]" country="Brazil" latitude="-7.3110003" longLatPrecision="1" longitude="-39.447002" municipality="Crato" name="Floresta Nacional do Araripe-Apodi" pageId="9" pageNumber="460" stateProvince="Ceara">Floresta Nacional do Araripe-Apodi</location>
,
<geoCoordinate id="EE6450118A7B017B39B18C1AFB984B46" box="[968,1104,1815,1840]" direction="south" orientation="latitude" pageId="9" pageNumber="460" precision="1" value="-7.3110003">7°18'39.6&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6450118A7B017B3E258C1AFB344B46" box="[1116,1276,1815,1840]" direction="west" orientation="longitude" pageId="9" pageNumber="460" precision="1" value="-39.447002">39°26'49.2&quot;W</geoCoordinate>
,
<collectingMunicipality id="6B8BACAC8A7B017B3F7E8C15FA804B46" box="[1287,1352,1816,1840]" pageId="9" pageNumber="460">Crato</collectingMunicipality>
,
<collectingRegion id="4994F8348A7B017B3F2D8C15FA504B46" box="[1364,1432,1816,1840]" country="Brazil" name="Ceara" pageId="9" pageNumber="460">Ceará</collectingRegion>
,
<collectingCountry id="F34776468A7B017B3AEE8C31FF164B23" box="[151,222,1852,1877]" name="Brazil" pageId="9" pageNumber="460">Brazil</collectingCountry>
,
<date id="FFEE10168A7B017B3A948C31FEAB4B23" box="[237,355,1852,1877]" pageId="9" pageNumber="460" value="2011-02-06">
<collectingDate id="EFAAE9FE8A7B017B3A948C31FEAB4B23" box="[237,355,1852,1877]" pageId="9" pageNumber="460" value="2011-02-06">06.II.2011</collectingDate>
</date>
,
<collectorName id="26A553008A7B017B3B0A8C31FDDB4B23" box="[371,531,1852,1877]" pageId="9" pageNumber="460">L.S. Carvalho</collectorName>
<emphasis id="B924EAC48A7B017B38678C30FD924B22" box="[542,602,1852,1876]" italics="true" pageId="9" pageNumber="460">et al.</emphasis>
leg. (
<specimenCode id="DBF69EAD8A7B017B38D98C31FC8C4B23" box="[672,836,1852,1877]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="9" pageNumber="460">MPEG 35011</specimenCode>
);
<materialsCitation id="3B383C8B8A7B017B39258C31FEB34BEA" ID-GBIF-Occurrence="2013952929" collectingDate="2013-05-14" collectionCode="UFMG" collectorName="da Samambaia &amp; L. S. Carvalho" country="Brazil" county="Ubajara" latitude="-3.8387222" location="Parque Nacional de Ubajara" longLatPrecision="1" longitude="-40.899612" municipality="Trilha da Samambaia" pageId="9" pageNumber="460" specimenCode="UFMG 14887" specimenCount="1" specimenCount-female="1" stateProvince="Ceara" typeStatus="paratype">
<emphasis id="B924EAC48A7B017B39258C31FC154B23" bold="true" box="[860,989,1852,1877]" pageId="9" pageNumber="460">
<typeStatus id="54EB88748A7B017B39258C31FC104B23" box="[860,984,1852,1877]" pageId="9" pageNumber="460" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1
<emphasis id="B924EAC48A7B017B398F8C31FBC34B23" bold="true" box="[1014,1035,1852,1877]" pageId="9" pageNumber="460">Ƌ</emphasis>
and
<specimenCount id="9D56FD5F8A7B017B3E308C31FBA44B22" box="[1097,1132,1852,1876]" pageId="9" pageNumber="460" type="female">1♀</specimenCount>
from
<location id="8E8F600D8A7B017B3EC18C30FF3B4B0E" LSID="urn:lsid:plazi:treatment:03F987C08A7B017C3AEE8DA0FBA94A95:8E8F600D8A7B017B3EC18C30FF3B4B0E" country="Brazil" county="Ubajara" latitude="-3.8387222" longLatPrecision="1" longitude="-40.899612" municipality="Trilha da Samambaia" name="Parque Nacional de Ubajara" pageId="9" pageNumber="460" stateProvince="Ceara">Parque Nacional de Ubajara</location>
,
<collectingMunicipality id="6B8BACAC8A7B017B3A878C52FE274B0E" box="[254,495,1887,1912]" pageId="9" pageNumber="460">
Trilha
<collectorName id="26A553008A7B017B3B318C52FE274B0E" box="[328,495,1887,1912]" pageId="9" pageNumber="460">da Samambaia</collectorName>
</collectingMunicipality>
,
<geoCoordinate id="EE6450118A7B017B3B838C52FD4A4B0E" box="[506,642,1887,1912]" direction="south" orientation="latitude" pageId="9" pageNumber="460" precision="1" value="-3.8387222">3°50'19.4&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6450118A7B017B38F78C52FCE54B0E" box="[654,813,1887,1912]" direction="west" orientation="longitude" pageId="9" pageNumber="460" precision="1" value="-40.899612">40°53'58.6&quot;W</geoCoordinate>
,
<collectingCounty id="628E4E5A8A7B017B39418C52FC5B4B0E" box="[824,915,1887,1912]" pageId="9" pageNumber="460">Ubajara</collectingCounty>
,
<collectingRegion id="4994F8348A7B017B39E68C6DFC2B4B0E" box="[927,995,1888,1912]" country="Brazil" name="Ceara" pageId="9" pageNumber="460">Ceará</collectingRegion>
,
<collectingCountry id="F34776468A7B017B39978C52FBFD4B0E" box="[1006,1077,1887,1912]" name="Brazil" pageId="9" pageNumber="460">Brazil</collectingCountry>
,
<collectingDate id="EFAAE9FE8A7B017B3E388C6DFB7E4B0E" box="[1089,1206,1888,1912]" pageId="9" pageNumber="460" value="2013-05-14">
<date id="FFEE10168A7B017B3E388C6DFB7E4B0E" box="[1089,1206,1888,1912]" pageId="9" pageNumber="460" value="2013-05-14">14.V.2013</date>
</collectingDate>
,
<collectorName id="26A553008A7B017B3EB88C6DFA964B0E" box="[1217,1374,1887,1912]" pageId="9" pageNumber="460">L.S. Carvalho</collectorName>
<emphasis id="B924EAC48A7B017B3F1D8C6CFA544B0E" box="[1380,1436,1888,1912]" italics="true" pageId="9" pageNumber="460">et al.</emphasis>
leg. (
<specimenCode id="DBF69EAD8A7B017B3AB78C89FEBA4BEA" box="[206,370,1924,1948]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="9" pageNumber="460">UFMG 14887</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2F665E8A7801783AEE8C5BFB834BBE" httpUri="https://zenodo.org/record/2601364/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,250,1878,1900]" targetBox="[166,1378,217,1855]" targetPageId="10">
<paragraph id="8BEF36D68A7801783AEE8C5BFB834BBE" blockId="10.[151,1436,1878,1992]" pageId="10" pageNumber="461">
<emphasis id="B924EAC48A7801783AEE8C5BFED74B1A" bold="true" box="[151,287,1878,1900]" pageId="10" pageNumber="461">FIGURE 10.</emphasis>
<taxonomicName id="4C504D558A7801783B5F8C5AFDD74B1A" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[294,543,1879,1900]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="sacrificatus" status="sp. nov.">
<emphasis id="B924EAC48A7801783B5F8C5AFDD74B1A" box="[294,543,1879,1900]" italics="true" pageId="10" pageNumber="461">Amphidraus sacrificatus</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A780178385C8C5AFDBE4B1A" bold="true" box="[549,630,1879,1900]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="A21757BF8A780178385C8C5AFDBE4B1A" box="[549,630,1879,1900]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, left male palp (A ventral; B embolic disc, ventral; C bulb removed, ventral; D retrolateral; E dorsal). Abbreviations: cc, cymbial conductor; Cy, cymbium; e, embolus shaft; ed, embolic disc; ef, embolic filament; dPED, distal processes on embolic disc; pPED, proximal process on embolic disc; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc.
</paragraph>
</caption>
<caption id="DF2F665E8A7901793AEE8C6DFA494BA4" httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1888,1910]" targetBox="[151,1436,193,1867]" targetPageId="11">
<paragraph id="8BEF36D68A7901793AEE8C6DFA494BA4" blockId="11.[151,1436,1888,2002]" pageId="11" pageNumber="462">
<emphasis id="B924EAC48A7901793AEE8C6DFEE94B03" bold="true" box="[151,289,1888,1910]" pageId="11" pageNumber="462">FIGURE 11.</emphasis>
<taxonomicName id="4C504D558A7901793B538C6DFE334B00" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[298,507,1888,1910]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7901793B538C6DFE334B00" box="[298,507,1888,1910]" italics="true" pageId="11" pageNumber="462">Amphidraus araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A790179387C8C6DFD914B03" bold="true" box="[517,601,1888,1909]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="A21757BF8A790179387C8C6DFD914B03" box="[517,601,1888,1909]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA.
</paragraph>
</caption>
<subSubSection id="C34A655D8A7E017E3ABE8B9AFB3B4CC6" box="[199,1267,151,176]" pageId="12" pageNumber="463" type="etymology">
<paragraph id="8BEF36D68A7E017E3ABE8B9AFB3B4CC6" blockId="12.[151,1437,151,752]" box="[199,1267,151,176]" pageId="12" pageNumber="463">
<emphasis id="B924EAC48A7E017E3ABE8B9AFE864CC6" bold="true" box="[199,334,151,176]" pageId="12" pageNumber="463">Etymology.</emphasis>
The specific epithet is a noun in apposition taken from the locality of the
<typeStatus id="54EB88748A7E017E3EF08B9AFB274CC6" box="[1161,1263,151,176]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A7E017E3ABE8BB1FE334E86" pageId="12" pageNumber="463" type="diagnosis">
<paragraph id="8BEF36D68A7E017E3ABE8BB1FE334E86" blockId="12.[151,1437,151,752]" pageId="12" pageNumber="463">
<emphasis id="B924EAC48A7E017E3ABE8BB1FE894CA3" bold="true" box="[199,321,188,213]" pageId="12" pageNumber="463">Diagnosis.</emphasis>
Among all species of
<taxonomicName id="4C504D558A7E017E38318BB1FD194CA2" authorityName="Simon" authorityYear="1900" box="[584,721,188,212]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAC48A7E017E38318BB1FD194CA2" box="[584,721,188,212]" italics="true" pageId="12" pageNumber="463">Amphidraus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D558A7E017E389B8BB1FC9F4CA2" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[738,855,188,212]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E389B8BB1FC9F4CA2" box="[738,855,188,212]" italics="true" pageId="12" pageNumber="463">A. araripe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BF8A7E017E39188BB1FC744CA2" box="[865,956,188,213]" pageId="12" pageNumber="463" rank="species">
<emphasis id="B924EAC48A7E017E39188BB1FC7E4CA3" bold="true" box="[865,950,188,213]" pageId="12" pageNumber="463">sp. nov</emphasis>
.
</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C504D558A7E017E3EE18BB1FED94C8E" authority="Salgado &amp; Ruiz, 2017" authorityName="Salgado &amp; Ruiz" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pulvinus">
<emphasis id="B924EAC48A7E017E3EE18BB1FAD14CA2" box="[1176,1305,188,212]" italics="true" pageId="12" pageNumber="463">A. pulvinus</emphasis>
<bibRefCitation id="EFC14B278A7E017E3F5A8BB1FED94C8E" author="Salgado, A. &amp; Ruiz, G. R." pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz, 2017</bibRefCitation>
</taxonomicName>
by having the tegulum with small retrolateral spermophore loop (RSPL) (
<figureCitation id="136B2A538A7E017E3E338BD2FB7B4C8E" box="[1098,1203,223,248]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Figs 11D</figureCitation>
,
<figureCitation id="136B2A538A7E017E3EC78BEDFB384C8E" box="[1214,1264,224,248]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">13B</figureCitation>
; see
<bibRefCitation id="EFC14B278A7E017E3F5F8BD2FEC74D6A" author="Salgado, A. &amp; Ruiz, G. R." pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 8C) and the same tibial apophysis pattern: the retroventral tibial apophysis (RvTA) and intercalary tibial apophysis (ITA) are short and blade-shaped (the ITA with an internal basal projection; black arrow in
<figureCitation id="136B2A538A7E017E3F098A2AFF0E4D12" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Fig. 11E</figureCitation>
; see also
<bibRefCitation id="EFC14B278A7E017E3B4B8A41FDE14D12" author="Salgado, A. &amp; Ruiz, G. R." box="[306,553,332,357]" pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 8D) and the RTA is linked to an inflated membrane (
<figureCitation id="136B2A538A7E017E3ECA8A41FACB4D12" box="[1203,1283,332,357]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Figs 11</figureCitation>
EG, 13BC; see Salgado &amp; Ruiz: figs 8B, 8D). These species also have an embolic disc with a distal process projected to prolateral side and a ventral process projected to retrolateral side (
<figureCitation id="136B2A538A7E017E39F38A99FC3C4DDA" box="[906,1012,404,429]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Figs 11C</figureCitation>
,
<figureCitation id="136B2A538A7E017E3E7B8A99FBFD4DDA" box="[1026,1077,404,428]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">13A</figureCitation>
; see
<bibRefCitation id="EFC14B278A7E017E3E168A99FAA24DDA" author="Salgado, A. &amp; Ruiz, G. R." box="[1135,1386,404,429]" pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 8C). However,
<taxonomicName id="4C504D558A7E017E3B3F8AB5FE734DB9" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[326,443,440,463]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3B3F8AB5FE734DB9" box="[326,443,440,463]" italics="true" pageId="12" pageNumber="463">A. araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E3BBD8ABAFDE84DA6" bold="true" box="[452,544,439,464]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E3BBD8ABAFDE84DA6" box="[452,544,439,464]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by having the RvTA and ITA more separated from each other (
<figureCitation id="136B2A538A7E017E3A9A8AD1FE834D82" box="[227,331,476,501]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Figs 11F</figureCitation>
,
<figureCitation id="136B2A538A7E017E3B228AD1FE444D82" box="[347,396,476,500]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">13B</figureCitation>
), whereas these are almost fused in
<taxonomicName id="4C504D558A7E017E39448AD1FC084D82" authorityName="Salgado &amp; Ruiz" authorityYear="2017" box="[829,960,476,500]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pulvinus">
<emphasis id="B924EAC48A7E017E39448AD1FC084D82" box="[829,960,476,500]" italics="true" pageId="12" pageNumber="463">A. pulvinus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC14B278A7E017E3E7D8AD1FACE4D82" author="Salgado, A. &amp; Ruiz, G. R." box="[1028,1286,476,501]" pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 8D). In addition,
<taxonomicName id="4C504D558A7E017E3B78890DFEBD4E61" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[257,373,512,535]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3B78890DFEBD4E61" box="[257,373,512,535]" italics="true" pageId="12" pageNumber="463">A. araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E3B048AF2FE1F4E6E" bold="true" box="[381,471,511,536]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E3B048AF2FE1F4E6E" box="[381,471,511,536]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a vPED placed near the distal processes (
<figureCitation id="136B2A538A7E017E39A48AF2FB8F4E6E" box="[989,1095,511,536]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="12" pageNumber="463">Figs 11C</figureCitation>
,
<figureCitation id="136B2A538A7E017E3E2A890DFB4E4E6E" box="[1107,1158,512,536]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">13A</figureCitation>
), whereas in
<taxonomicName id="4C504D558A7E017E3F64890DFA544E6E" authorityName="Salgado &amp; Ruiz" authorityYear="2017" box="[1309,1436,512,536]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pulvinus">
<emphasis id="B924EAC48A7E017E3F64890DFA544E6E" box="[1309,1436,512,536]" italics="true" pageId="12" pageNumber="463">A. pulvinus</emphasis>
</taxonomicName>
the vPED is placed on the prolateral border of embolic disc (see
<bibRefCitation id="EFC14B278A7E017E390A8929FBA44E4A" author="Salgado, A. &amp; Ruiz, G. R." box="[883,1132,548,573]" pageId="12" pageNumber="463" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 8C). The female of
<taxonomicName id="4C504D558A7E017E3FFD8929FF3E4E16" authorityName="Salgado &amp; Ruiz" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pulvinus">
<emphasis id="B924EAC48A7E017E3FFD8929FF3E4E16" italics="true" pageId="12" pageNumber="463">A. pulvinus</emphasis>
</taxonomicName>
is unknown, but among the
<taxonomicName id="4C504D558A7E017E38388945FD024E16" authorityName="Simon" authorityYear="1900" box="[577,714,584,608]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAC48A7E017E38388945FD024E16" box="[577,714,584,608]" italics="true" pageId="12" pageNumber="463">Amphidraus</emphasis>
</taxonomicName>
species currently known, the epigyne of
<taxonomicName id="4C504D558A7E017E3EDF8945FAD34E29" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1190,1307,584,607]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3EDF8945FAD34E29" box="[1190,1307,584,607]" italics="true" pageId="12" pageNumber="463">A. araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E3F5D894AFA484E16" bold="true" box="[1316,1408,583,608]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E3F5D894AFA484E16" box="[1316,1408,583,608]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to that of
<taxonomicName id="4C504D558A7E017E3BD88961FDEF4EF2" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[417,551,620,644]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="manaura" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3BD88961FDEF4EF2" box="[417,551,620,644]" italics="true" pageId="12" pageNumber="463">A. manaura</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E38488961FD454EF3" bold="true" box="[561,653,620,645]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E38488961FD454EF3" box="[561,653,620,645]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, both with glands near copulatory openings and with epigynal plate with border slightly bilobed (
<figureCitation id="136B2A538A7E017E3B918982FD9D4EDE" box="[488,597,655,680]" captionStart="FIGURE 12" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[187,1409,836,1922]" captionTargetId="figure@12.[173,1417,805,1931]" captionTargetPageId="12" captionText="FIGURE 12. Amphidraus araripe sp. nov. AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral, D dorsal). Abbreviations: cd, copulatory duct; fd, fertilization duct; gl, gland; S spermatheca. White arrow in C shows copulatory opening." httpUri="https://zenodo.org/record/2601368/files/figure.png" pageId="12" pageNumber="463">Figs 12C</figureCitation>
,
<figureCitation id="136B2A538A7E017E381A899DFD5E4EDE" box="[611,662,656,680]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">13D</figureCitation>
,
<figureCitation id="136B2A538A7E017E38DD8982FD1F4EDE" box="[676,727,655,680]" captionStart="FIGURE 16" captionStartId="16.[151,250,1571,1593]" captionTargetBox="[167,1436,193,1539]" captionTargetId="figure@16.[151,1436,193,1549]" captionTargetPageId="16" captionText="FIGURE 16. Amphidraus manauara sp. nov. AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral; D ventral, cleared). Abbreviations: co, copulatory opening; fd, fertilization duct; po, coupling pocket; S, spermatheca." httpUri="https://zenodo.org/record/2601376/files/figure.png" pageId="12" pageNumber="463">16D</figureCitation>
,
<figureCitation id="136B2A538A7E017E389C899DFCD14EDE" box="[741,793,656,680]" captionStart="FIGURE 17" captionStartId="18.[151,250,1883,1905]" captionTargetBox="[157,1401,276,1828]" captionTargetId="figure@18.[151,1436,248,1861]" captionTargetPageId="18" captionText="FIGURE 17. Amphidraus manauara sp. nov. AC left male palp (A ventral; B dorsal; C retrolateral); D epigyne/vulva, cleared, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening; Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; po, coupling pocket; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; T, tegulum; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601378/files/figure.png" pageId="12" pageNumber="463">17D</figureCitation>
). However, the female of
<taxonomicName id="4C504D558A7E017E3E31899DFB764ED1" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1096,1214,656,679]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3E31899DFB764ED1" box="[1096,1214,656,679]" italics="true" pageId="12" pageNumber="463">A. araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E3EB18982FAEC4EDE" bold="true" box="[1224,1316,655,680]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E3EB18982FAEC4EDE" box="[1224,1316,655,680]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs by having a longer copulatory duct (with one loop ventrally projected) and by having the less expanded border of epigynal plate (
<figureCitation id="136B2A538A7E017E3B3A89DAFE674E86" box="[323,431,727,752]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="12" pageNumber="463">Figs 13D</figureCitation>
,
<figureCitation id="136B2A538A7E017E3BC389D5FE264E86" box="[442,494,728,752]" captionStart="FIGURE 17" captionStartId="18.[151,250,1883,1905]" captionTargetBox="[157,1401,276,1828]" captionTargetId="figure@18.[151,1436,248,1861]" captionTargetPageId="18" captionText="FIGURE 17. Amphidraus manauara sp. nov. AC left male palp (A ventral; B dorsal; C retrolateral); D epigyne/vulva, cleared, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening; Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; po, coupling pocket; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; T, tegulum; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601378/files/figure.png" pageId="12" pageNumber="463">17D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2F665E8A7E017E3AEE8C97FF384B9B" httpUri="https://zenodo.org/record/2601368/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,1946,1968]" targetBox="[187,1409,836,1922]" targetPageId="12">
<paragraph id="8BEF36D68A7E017E3AEE8C97FF384B9B" blockId="12.[151,1437,1946,2029]" pageId="12" pageNumber="463">
<emphasis id="B924EAC48A7E017E3AEE8C97FEEB4BD9" bold="true" box="[151,291,1946,1968]" pageId="12" pageNumber="463">FIGURE 12.</emphasis>
<taxonomicName id="4C504D558A7E017E3B578C97FDC84BC6" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[302,512,1946,1968]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7E017E3B578C97FDC84BC6" box="[302,512,1946,1968]" italics="true" pageId="12" pageNumber="463">Amphidraus araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7E017E38728C97FD974BD9" bold="true" box="[523,607,1946,1967]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A21757BF8A7E017E38728C97FD974BD9" box="[523,607,1946,1967]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral, D dorsal). Abbreviations: cd, copulatory duct; fd, fertilization duct; gl, gland; S spermatheca. White arrow in C shows copulatory opening.
</paragraph>
</caption>
<caption id="DF2F665E8A7F017F3AEE8C36FD914BBD" httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="13" pageNumber="464" startId="13.[151,250,1851,1873]" targetBox="[198,1418,208,1790]" targetPageId="13">
<paragraph id="8BEF36D68A7F017F3AEE8C36FD914BBD" blockId="13.[151,1436,1851,1995]" pageId="13" pageNumber="464">
<emphasis id="B924EAC48A7F017F3AEE8C36FEE84B26" bold="true" box="[151,288,1851,1873]" pageId="13" pageNumber="464">FIGURE 13.</emphasis>
<taxonomicName id="4C504D558A7F017F3B5E8C36FE3D4B27" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[295,501,1851,1873]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="araripe" status="sp. nov.">
<emphasis id="B924EAC48A7F017F3B5E8C36FE3D4B27" box="[295,501,1851,1873]" italics="true" pageId="13" pageNumber="464">Amphidraus araripe</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A7F017F3B858C36FD854B26" bold="true" box="[508,589,1851,1872]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="A21757BF8A7F017F3B858C36FD854B26" box="[508,589,1851,1872]" pageId="13" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc.
</paragraph>
</caption>
<caption id="DF2F665E8A7C017C3AEE8F05FB7C4836" httpUri="https://zenodo.org/record/2601372/files/figure.png" pageId="14" pageNumber="465" startId="14.[151,250,1032,1054]" targetBox="[163,1419,197,1010]" targetPageId="14">
<paragraph id="8BEF36D68A7C017C3AEE8F05FB7C4836" blockId="14.[151,1435,1032,1088]" pageId="14" pageNumber="465">
<emphasis id="B924EAC48A7C017C3AEE8F05FEE8486B" bold="true" box="[151,288,1032,1054]" pageId="14" pageNumber="465">FIGURE 14.</emphasis>
Map of northern/northeastern Brazil showing records of
<taxonomicName id="4C504D558A7C017C391E8F04FBFE4868" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[871,1078,1033,1054]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="araripe">
<emphasis id="B924EAC48A7C017C391E8F04FBFE4868" box="[871,1078,1033,1054]" italics="true" pageId="14" pageNumber="465">Amphidraus araripe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D558A7C017C3E3A8F04FB1B4868" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1091,1235,1033,1054]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="boomerang">
<emphasis id="B924EAC48A7C017C3E3A8F04FB1B4868" box="[1091,1235,1033,1054]" italics="true" pageId="14" pageNumber="465">A. boomerang</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D558A7C017C3E998F04FAB84868" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1248,1392,1033,1054]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cornipalpis">
<emphasis id="B924EAC48A7C017C3E998F04FAB84868" box="[1248,1392,1033,1054]" italics="true" pageId="14" pageNumber="465">A. cornipalpis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D558A7C017C3AEE8F27FEE14849" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[151,297,1066,1087]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="sacrificatus">
<emphasis id="B924EAC48A7C017C3AEE8F27FEE14849" box="[151,297,1066,1087]" italics="true" pageId="14" pageNumber="465">A. sacrificatus</emphasis>
</taxonomicName>
. White lines represent the borders of Brazilian states. Map modified from Google Earth
<superScript id="7C259B9E8A7C017C3EDD8F2AFB674845" attach="left" box="[1188,1199,1063,1075]" fontSize="5" pageId="14" pageNumber="465">®</superScript>
.
</paragraph>
</caption>
<subSubSection id="C34A655D8A7C017C3ABE8F6AFE164AC9" pageId="14" pageNumber="465" type="description">
<paragraph id="8BEF36D68A7C017C3ABE8F6AFE8A49D6" blockId="14.[151,1437,1127,1764]" pageId="14" pageNumber="465">
<emphasis id="B924EAC48A7C017C3ABE8F6AFD0048F6" bold="true" box="[199,712,1127,1152]" pageId="14" pageNumber="465">
Description. Male
<typeStatus id="54EB88748A7C017C3BDB8F6AFDC148F6" box="[418,521,1127,1152]" pageId="14" pageNumber="465" type="holotype">holotype</typeStatus>
(MPEG 35011).
</emphasis>
Total length: 2.73. Carapace 1.32 long, 0.88 wide and 0.65 high. Ocular quadrangle 0.54 long. Anterior eye row 0.93 wide, posterior 0.81 wide. Legs 4312. Length of leg: I 2.09 (0.63 + 0.77 + 0.69); II 2.04 (0.62 + 0.70 + 0.72); III 2.50 (0.75 + 0.82 + 0.93); IV 2.78 (0.79 + 0.94 + 1.05). Palp (
<figureCitation id="136B2A538A7C017C3AE68FDEFF27489D" box="[159,239,1235,1260]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="14" pageNumber="465">Figs 11</figureCitation>
CG, 13AC): femur and patella without modifications; cymbium oval (
<figureCitation id="136B2A538A7C017C3E618FDEFBB6489D" box="[1048,1150,1235,1260]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="14" pageNumber="465">Fig. 13C</figureCitation>
); tegulum with thick lobe curved to prolateral side (
<figureCitation id="136B2A538A7C017C3BCB8FFAFDD24979" box="[434,538,1271,1296]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="14" pageNumber="465">Figs 11C</figureCitation>
,
<figureCitation id="136B2A538A7C017C385C8FFAFD914979" box="[549,601,1271,1295]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="14" pageNumber="465">13A</figureCitation>
); embolus shaft (e) strongly arched to prolateral side, with tip retrolaterally projected (
<figureCitation id="136B2A538A7C017C3B768E16FEBF4945" box="[271,375,1307,1332]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="14" pageNumber="465">Fig. 13A</figureCitation>
); embolic filament (ef) short, with nearly half the length of embolus shaft (
<figureCitation id="136B2A538A7C017C3EBA8E16FAE44945" box="[1219,1324,1307,1332]" captionStart="FIGURE 13" captionStartId="13.[151,250,1851,1873]" captionTargetBox="[198,1418,208,1790]" captionTargetId="figure@13.[151,1436,193,1829]" captionTargetPageId="13" captionText="FIGURE 13. Amphidraus araripe sp. nov., left male palp (A ventral, B retrolateral, C dorsal, after bulb removal). D epigyne/ vulva, ventral. Abbreviations: cc, cymbial conductor; cd, copulatory duct; co, copulatory opening. Cy, cymbium; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPEd, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601370/files/figure.png" pageId="14" pageNumber="465">Fig. 13A</figureCitation>
). Color in alcohol (
<figureCitation id="136B2A538A7C017C3A828E32FE984921" box="[251,336,1343,1368]" captionStart="FIGURE 11" captionStartId="11.[151,250,1888,1910]" captionTargetBox="[151,1436,193,1867]" captionTargetId="figure@11.[151,1436,193,1867]" captionTargetPageId="11" captionText="FIGURE 11. Amphidraus araripe sp. nov. AB male (A dorsal, B ventral); CG left male palp (C bulb, ventral; D bulb, retrolateral; E tibia, ventral, F tibia, retrolateral; G tibia, dorsal). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; ITA, intercalary tibial apophysis; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. Black arrow in E shows the internal spike of ITA." httpUri="https://zenodo.org/record/2601366/files/figure.png" pageId="14" pageNumber="465">Figs 11</figureCitation>
AB): cephalic area black; thoracic area brown with pale longitudinal stripe; abdomen dorsally variegated with brown, and with a pale longitudinal stripe bearing white setae; ventrally pale with posterior portion dark; legs pale.
</paragraph>
<paragraph id="8BEF36D68A7C017C3ABE8EA6FE164AC9" blockId="14.[151,1437,1127,1764]" pageId="14" pageNumber="465">
<emphasis id="B924EAC48A7C017C3ABE8EA6FD3B49B2" bold="true" box="[199,755,1451,1476]" pageId="14" pageNumber="465">
Description. Female
<typeStatus id="54EB88748A7C017C3BB88EA6FDE549B2" box="[449,557,1451,1476]" pageId="14" pageNumber="465" type="paratype">paratype</typeStatus>
(UFMG 14887).
</emphasis>
Total length: 2.95. Carapace 1.40 long, 0.97 wide and 0.67 high. Ocular quadrangle 0.68 long. Anterior eye row 1.02 wide, posterior 0.90 wide. Legs 4312. Length of leg: I 2.18 (0.71 + 0.82 + 0.65); II 2.06 (0.66 + 0.76 + 0.64); III 2.50 (0.79 + 0.86 + 0.85); IV 2.88 (0.81+ 0.99 + 1.08). Epigyne/vulva (
<figureCitation id="136B2A538A7C017C3B378D1AFE6E4A59" box="[334,422,1559,1584]" captionStart="FIGURE 12" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[187,1409,836,1922]" captionTargetId="figure@12.[173,1417,805,1931]" captionTargetPageId="12" captionText="FIGURE 12. Amphidraus araripe sp. nov. AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral, D dorsal). Abbreviations: cd, copulatory duct; fd, fertilization duct; gl, gland; S spermatheca. White arrow in C shows copulatory opening." httpUri="https://zenodo.org/record/2601368/files/figure.png" pageId="14" pageNumber="465">Figs 12</figureCitation>
CD, 13D: copulatory openings separated by about two diameters of copulatory duct; spermathecae kidney-shaped; fertilization ducts anteriorly placed and laterally projected. Color in alcohol (
<figureCitation id="136B2A538A7C017C3F138D36FF7B4A01" captionStart="FIGURE 12" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[187,1409,836,1922]" captionTargetId="figure@12.[173,1417,805,1931]" captionTargetPageId="12" captionText="FIGURE 12. Amphidraus araripe sp. nov. AB female (A dorsal, B ventral); CD epigyne/vulva (C ventral, D dorsal). Abbreviations: cd, copulatory duct; fd, fertilization duct; gl, gland; S spermatheca. White arrow in C shows copulatory opening." httpUri="https://zenodo.org/record/2601368/files/figure.png" pageId="14" pageNumber="465">Figs 12</figureCitation>
AB): cephalic area black; thoracic area dark brown with a longitudinal light brown stripe; abdomen dorsally black with longitudinal interrupted pale stripe; ventrally pale; femora pale with black spots; other leg segments light brown with black spots.
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A7C017C3ABE8DC6FBA94A95" box="[199,1121,1739,1764]" pageId="14" pageNumber="465" type="distribution">
<paragraph id="8BEF36D68A7C017C3ABE8DC6FBA94A95" blockId="14.[151,1437,1127,1764]" box="[199,1121,1739,1764]" pageId="14" pageNumber="465">
<emphasis id="B924EAC48A7C017C3ABE8DC6FEA84A92" bold="true" box="[199,352,1739,1764]" pageId="14" pageNumber="465">Distribution.</emphasis>
Known only from the
<typeStatus id="54EB88748A7C017C38198DC1FD584A92" box="[608,656,1740,1764]" pageId="14" pageNumber="465">type</typeStatus>
locality (state of
<collectingRegion id="4994F8348A7C017C392D8DC6FC504A95" box="[852,920,1739,1763]" country="Brazil" name="Ceara" pageId="14" pageNumber="465">Ceará</collectingRegion>
,
<collectingCountry id="F34776468A7C017C39DA8DC6FC214A92" box="[931,1001,1739,1764]" name="Brazil" pageId="14" pageNumber="465">Brazil</collectingCountry>
) (
<figureCitation id="136B2A538A7C017C3E798DC6FB9C4A95" box="[1024,1108,1739,1764]" captionStart="FIGURE 14" captionStartId="14.[151,250,1032,1054]" captionTargetBox="[163,1419,197,1010]" captionTargetId="figure@14.[163,1419,197,1010]" captionTargetPageId="14" captionText="FIGURE 14. Map of northern/northeastern Brazil showing records of Amphidraus araripe, A. boomerang, A. cornipalpis and A. sacrificatus. White lines represent the borders of Brazilian states. Map modified from Google Earth ®." httpUri="https://zenodo.org/record/2601372/files/figure.png" pageId="14" pageNumber="465">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>