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<document ID-DOI="http://doi.org/10.5281/zenodo.5689401" ID-GBIF-Dataset="6ea2d9a4-cff3-42df-ad0c-cfbc6f9034ed" ID-GBIF-Taxon="119377351" ID-ISSN="1175-5326" ID-Zenodo-Dep="5689401" checkinTime="1459966256126" checkinUser="plazi" docAuthor="Pola, Marta, Rudman, William B. &amp; Gosliner, Terrence M." docDate="2009" docId="03F98793FF8272292BCFF95DA428FDE5" docLanguage="en" docName="zt01975p057.pdf" docOrigin="Zootaxa 1975" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2814bb39" docStyleName="zootaxa.2007.monograph" docTitle="Bornella sarape Bertsch 1980" docType="treatment" docVersion="8" lastPageId="25" lastPageNumber="26" masterDocId="FFC0FFEBFF9472302B58FFD8A00AFFA7" masterDocTitle="Systematics and preliminary phylogeny of Bornellidae (Mollusca: Nudibranchia: Dendronotina) based on morphological characters with description of four new species" masterLastPageNumber="57" masterPageNumber="1" pageId="22" pageNumber="23" updateTime="1636786038398" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics and preliminary phylogeny of Bornellidae (Mollusca: Nudibranchia: Dendronotina) based on morphological characters with description of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pola, Marta</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rudman, William B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gosliner, Terrence M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="volume">
<mods:number>1975</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>57</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.185130</mods:identifier>
<mods:identifier type="GBIF-Dataset">6ea2d9a4-cff3-42df-ad0c-cfbc6f9034ed</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">185130</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5689401" ID-GBIF-Taxon="119377351" ID-Zenodo-Dep="5689401" LSID="urn:lsid:plazi:treatment:03F98793FF8272292BCFF95DA428FDE5" httpUri="http://treatment.plazi.org/id/03F98793FF8272292BCFF95DA428FDE5" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection pageId="22" pageNumber="23" type="nomenclature">
<paragraph blockId="22.[151,520,1668,1729]" box="[151,520,1668,1695]" pageId="22" pageNumber="23">
<heading bold="true" box="[151,520,1668,1695]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName authority="Bertsch, 1980" authorityName="Bertsch" authorityYear="1980" box="[151,520,1668,1695]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="sarape">
<emphasis bold="true" box="[151,520,1668,1695]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[151,343,1669,1695]" italics="true" pageId="22" pageNumber="23">Bornella sarape</emphasis>
<bibRefCitation author="Bertsch" box="[351,520,1668,1694]" pageId="22" pageNumber="54" refString="Bertsch, H. (1980) A new species of Bornella from tropical west-America (Mollusca, Opisthobranchia). Spixiana, 3, 33 - 42." type="journal article" year="1980">
Bertsch,
<number box="[462,520,1668,1694]" pageId="22" pageNumber="23" value="1980.0">1980</number>
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="22.[151,520,1668,1729]" box="[151,393,1703,1729]" pageId="22" pageNumber="23">
(
<figureCitation box="[159,267,1703,1729]" captionStart="FIGURE 1" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[198,1387,194,1713]" captionTargetId="figure@7.[197,1390,194,1715]" captionTargetPageId="7" captionText="FIGURE 1. Living animals. A B, Bornella stellifer (Adams and Reeve in Adams, 1848); A, Malaysia, Malay Peninsula, South China Sea, Photo: T. M. Gosliner. B, Papua New Guinea, north coast, Madang, Photo: R. C. Willan. C D, Bornella calcarata Mörch, 1863; C, Brazil, Photo: C. M. Cunha. D, Guarapi, Espiritu Santo, Brazil, Photo: J. P. Cauduro Filho. E, Bornella sarape Bertsch, 1980, Pacific coast of Mexico, Photo: A. Hermosillo. F, Bornella anguilla Johnson, 1984, Japan, Ryukyu Island, Okinawa, Photo: R. F. Bolland. G, Bornella irvingi Edmunds and Preece, 1996, Pitcairn Islands, Photo: J. Jamieson. H, Bornella dotoides sp. nov. Papua New Guinea, north Madang, “ The Quarry ”, Photo: T. M. Gosliner." httpUri="https://zenodo.org/record/185131/files/figure.png" pageId="22" pageNumber="23">
Figures
<number box="[256,267,1703,1729]" pageId="22" pageNumber="23" value="1.0">1</number>
</figureCitation>
E,
<number box="[303,319,1703,1729]" pageId="22" pageNumber="23" value="4.0">4</number>
D,
<number box="[353,383,1703,1729]" pageId="22" pageNumber="23" value="12.0">12</number>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,684,1775,1798]" pageId="22" pageNumber="23" type="reference_group">
<paragraph blockId="22.[151,684,1775,1798]" box="[151,684,1775,1798]" pageId="22" pageNumber="23">
<treatmentCitationGroup box="[151,684,1775,1798]" pageId="22" pageNumber="23">
<taxonomicName authority="Bertsch, 1980: 33" authorityName="Bertsch" authorityPageNumber="33" authorityYear="1980" box="[151,518,1775,1798]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="sarape">
<emphasis box="[151,323,1775,1798]" italics="true" pageId="22" pageNumber="23">Bornella sarape</emphasis>
<treatmentCitation author="Bertsch" box="[330,518,1775,1798]" page="33" pageId="22" pageNumber="23" year="1980">
<bibRefCitation author="Bertsch" box="[330,482,1775,1798]" pageId="22" pageNumber="54" refString="Bertsch, H. (1980) A new species of Bornella from tropical west-America (Mollusca, Opisthobranchia). Spixiana, 3, 33 - 42." type="journal article" year="1980">
Bertsch,
<number box="[425,482,1775,1798]" pageId="22" pageNumber="23" value="1980.0">1980</number>
</bibRefCitation>
:
<number box="[492,518,1775,1798]" pageId="22" pageNumber="23" value="37.5" valueMax="42.0" valueMin="33.0">33</number>
</treatmentCitation>
</taxonomicName>
42, figs.
<number box="[625,681,1775,1798]" pageId="22" pageNumber="23" value="6.0" valueMax="11.0" valueMin="1.0">111</number>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="materials_examined">
<paragraph blockId="22.[151,1436,1847,2033]" lastBlockId="23.[151,1437,152,2018]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis bold="true" box="[151,402,1847,1873]" pageId="22" pageNumber="23">Material examined:</emphasis>
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[414,499,1847,1873]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23">LACM</collectionCode>
<number box="[511,581,1847,1873]" pageId="22" pageNumber="23" value="53.5" valueMax="71.0" valueMin="36.0">36-71</number>
,
<collectingCountry box="[599,691,1847,1873]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
, Gulf of California,
<collectingCountry box="[948,990,1847,1873]" name="Iceland" pageId="22" pageNumber="23">Isla</collectingCountry>
Partida, off White Rock, 28.55.5'N, 113.05.6'W, 0
<date box="[313,478,1887,1913]" pageId="22" pageNumber="23" value="1936-03-08">
<number box="[313,326,1887,1913]" pageId="22" pageNumber="23" value="8.0">8</number>
March
<number box="[417,478,1887,1913]" pageId="22" pageNumber="23" value="1936.0">1936</number>
</date>
, one adult specimen
<quantity box="[723,805,1887,1913]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="23" unit="mm" value="25.0">25 mm</quantity>
preserved, dissected,
<quantity box="[1062,1121,1887,1913]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="22" pageNumber="23" unit="m" value="82.0">82 m</quantity>
depth, coll: R/V Velero III (
<collectionCode box="[159,220,1927,1953]" httpUri="http://grbio.org/cool/95c7-rrrz" name="Allan Hancock Foundation, University of Southern California" pageId="22" pageNumber="23">AHF</collectionCode>
<number box="[229,277,1927,1953]" pageId="22" pageNumber="23" value="557.0">557</number>
-
<number box="[286,317,1927,1953]" pageId="22" pageNumber="23" value="36.0">36</number>
).
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[339,424,1927,1953]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23">LACM</collectionCode>
<number box="[434,505,1927,1953]" pageId="22" pageNumber="23" value="46.5" valueMax="53.0" valueMin="40.0">40-53</number>
,
<collectingCountry box="[519,612,1927,1953]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
, Gulf of California, North of
<collectingCountry box="[973,1015,1927,1953]" name="Iceland" pageId="22" pageNumber="23">Isla</collectingCountry>
Partida, 28.56.2'N, 113.02.6'W, 0
<date pageId="22" pageNumber="23" value="1940-02-05">
<number box="[1422,1436,1927,1953]" pageId="22" pageNumber="23" value="5.0">5</number>
February
<number box="[265,326,1967,1993]" pageId="22" pageNumber="23" value="1940.0">1940</number>
</date>
, one adult specimen
<quantity box="[576,659,1967,1993]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="23" unit="mm" value="25.0">25 mm</quantity>
preserved, dissected,
<quantity box="[920,1039,1967,1993]" metricMagnitude="2" metricUnit="m" metricValue="1.115" metricValueMax="1.39" metricValueMin="0.84" pageId="22" pageNumber="23" unit="m" value="111.5" valueMax="139.0" valueMin="84.0">84139 m</quantity>
depth, coll: R/V Velero III (
<collectionCode box="[1376,1436,1967,1993]" httpUri="http://grbio.org/cool/95c7-rrrz" name="Allan Hancock Foundation, University of Southern California" pageId="22" pageNumber="23">AHF</collectionCode>
<number box="[151,212,2007,2033]" pageId="22" pageNumber="23" value="1081.0">1081</number>
-
<number box="[220,253,2007,2033]" pageId="22" pageNumber="23" value="40.0">40</number>
).
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[273,358,2007,2033]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23">LACM</collectionCode>
<number box="[365,441,2007,2033]" pageId="22" pageNumber="23" value="14801.0">14801</number>
,
<collectingCountry box="[453,545,2007,2033]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
, Jalisco, Melaque,
<date box="[768,927,2007,2033]" pageId="22" pageNumber="23" value="1976-01">
January
<number box="[866,927,2007,2033]" pageId="22" pageNumber="23" value="1976.0">1976</number>
</date>
, one immature specimen
<quantity box="[1226,1308,2007,2033]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="mm" value="11.0">11 mm</quantity>
preserved, dissected, intertidal, coll: Forest &amp; Roy Poorman.
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[764,849,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<number box="[860,936,152,178]" pageId="23" pageNumber="24" value="24996.0">24996</number>
,
<collectingCountry box="[951,1044,152,178]" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingCountry>
, Sonora, San Carlos Bay, trailer park,
<date box="[220,384,192,218]" pageId="23" pageNumber="24" value="1976-10">
October
<number box="[323,384,192,218]" pageId="23" pageNumber="24" value="1976.0">1976</number>
</date>
, five adult specimens
<number box="[652,730,192,218]" pageId="23" pageNumber="24" value="15.15">15, 15</number>
,
<number box="[744,819,192,218]" pageId="23" pageNumber="24" value="16.17">16, 17</number>
&amp;
<quantity box="[861,945,192,218]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="24" unit="mm" value="17.0">17 mm</quantity>
preserved, two dissected, intertidal, coll: Forest &amp; Roy Poorman.
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[441,526,232,258]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<number box="[534,610,232,258]" pageId="23" pageNumber="24" value="25007.0">25007</number>
,
<collectingCountry box="[622,714,232,258]" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingCountry>
, Sonora, Guaymas, San Carlos Bay, Honeymoon
<collectingCountry box="[1302,1376,232,258]" name="Iceland" pageId="23" pageNumber="24">Island</collectingCountry>
, diving on
<taxonomicName box="[232,316,272,297]" class="Phaeophyceae" family="Dictyotaceae" genus="Padina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dictyotales" pageId="23" pageNumber="24" phylum="Phaeophyta" rank="genus">
<emphasis box="[232,316,272,297]" italics="true" pageId="23" pageNumber="24">Padina</emphasis>
</taxonomicName>
,
<number box="[332,393,272,298]" pageId="23" pageNumber="24" value="1976.0">1976</number>
, one adult specimen
<quantity box="[638,721,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="mm" value="20.0">20 mm</quantity>
preserved, dissected, coll: R. Poorman &amp; Shy.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="distribution">
<paragraph blockId="23.[151,1437,152,2018]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[198,361,311,337]" pageId="23" pageNumber="24">Distribution:</emphasis>
From
<collectingCountry box="[439,531,312,338]" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingCountry>
, Gulf of California to Bahia Banderas, and
<collectingCountry box="[1037,1130,312,338]" name="Panama" pageId="23" pageNumber="24">Panama</collectingCountry>
(
<bibRefCitation author="Bertsch" box="[1145,1305,312,338]" pageId="23" pageNumber="54" refString="Bertsch, H. (1980) A new species of Bornella from tropical west-America (Mollusca, Opisthobranchia). Spixiana, 3, 33 - 42." type="journal article" year="1980">
Bertsch
<number box="[1243,1305,312,338]" pageId="23" pageNumber="24" value="1980.0">1980</number>
</bibRefCitation>
;
<bibRefCitation author="Bertsch" pageId="23" pageNumber="54" refString="Bertsch, H. &amp; Kerstitch. A. (1984) Distribution and radular morphology of various nudibranchs (Gastropoda: Opisthobranchia) from the Gulf of California, Mexico. The Vel iger, 26, 264 - 273." type="journal article" year="1984">
Bertsch &amp; Kerstitch
<number box="[266,328,352,378]" pageId="23" pageNumber="24" value="1984.0">1984</number>
</bibRefCitation>
;
<bibRefCitation author="Hermosillo" box="[339,609,352,378]" pageId="23" pageNumber="55" refString="Hermosillo, A., Behrens, D. W. &amp; Rios Jara, E. (2006) Opisthobranquios de Mexico. Guia de babosas marinas del Pacifico, Golfo de California y las islas oceanicas. CONABIO, Guadalajara, Jalisco, Mexico, 144 pp." type="book" year="2006">
Hermosillo
<emphasis box="[479,537,352,377]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
<number box="[545,609,352,378]" pageId="23" pageNumber="24" value="2006.0">2006</number>
</bibRefCitation>
).
</paragraph>
<paragraph blockId="23.[151,1437,152,2018]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[197,476,391,417]" pageId="23" pageNumber="24">External morphology:</emphasis>
The general body shape is elongate and limaciform with the posterior end of the foot being long and tapering (
<figureCitation box="[510,578,432,458]" captionStart="FIGURE 1" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[198,1387,194,1713]" captionTargetId="figure@7.[197,1390,194,1715]" captionTargetPageId="7" captionText="FIGURE 1. Living animals. A B, Bornella stellifer (Adams and Reeve in Adams, 1848); A, Malaysia, Malay Peninsula, South China Sea, Photo: T. M. Gosliner. B, Papua New Guinea, north coast, Madang, Photo: R. C. Willan. C D, Bornella calcarata Mörch, 1863; C, Brazil, Photo: C. M. Cunha. D, Guarapi, Espiritu Santo, Brazil, Photo: J. P. Cauduro Filho. E, Bornella sarape Bertsch, 1980, Pacific coast of Mexico, Photo: A. Hermosillo. F, Bornella anguilla Johnson, 1984, Japan, Ryukyu Island, Okinawa, Photo: R. F. Bolland. G, Bornella irvingi Edmunds and Preece, 1996, Pitcairn Islands, Photo: J. Jamieson. H, Bornella dotoides sp. nov. Papua New Guinea, north Madang, “ The Quarry ”, Photo: T. M. Gosliner." httpUri="https://zenodo.org/record/185131/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[567,578,432,458]" pageId="23" pageNumber="24" value="1.0">1</number>
</figureCitation>
E). The living adults reach to
<quantity box="[945,1029,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="24" unit="mm" value="40.0">40 mm</quantity>
in length (
<bibRefCitation author="Hermosillo" box="[1162,1369,432,458]" pageId="23" pageNumber="55" refString="Hermosillo, A. (2002) Bornella sarape from Mexico. [Message in] Sea Slug Forum. Australian Museum, Sydney. Available from http: // www. seaslugforum. net / find. cfm? id = 7179 (15 June 2002), (Last accessed: 26 October 2008)." type="book" year="2002">
Hermosillo
<number box="[1305,1369,432,458]" pageId="23" pageNumber="24" value="2002.0">2002</number>
</bibRefCitation>
). The general color pattern is pale yellow-brown, with dark brown to black reticulate pattern, which sometimes coalesces into dark patches. The dorsolateral processes and rhinophore sheaths have the same coloration (
<figureCitation captionStart="FIGURE 1" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[198,1387,194,1713]" captionTargetId="figure@7.[197,1390,194,1715]" captionTargetPageId="7" captionText="FIGURE 1. Living animals. A B, Bornella stellifer (Adams and Reeve in Adams, 1848); A, Malaysia, Malay Peninsula, South China Sea, Photo: T. M. Gosliner. B, Papua New Guinea, north coast, Madang, Photo: R. C. Willan. C D, Bornella calcarata Mörch, 1863; C, Brazil, Photo: C. M. Cunha. D, Guarapi, Espiritu Santo, Brazil, Photo: J. P. Cauduro Filho. E, Bornella sarape Bertsch, 1980, Pacific coast of Mexico, Photo: A. Hermosillo. F, Bornella anguilla Johnson, 1984, Japan, Ryukyu Island, Okinawa, Photo: R. F. Bolland. G, Bornella irvingi Edmunds and Preece, 1996, Pitcairn Islands, Photo: J. Jamieson. H, Bornella dotoides sp. nov. Papua New Guinea, north Madang, “ The Quarry ”, Photo: T. M. Gosliner." httpUri="https://zenodo.org/record/185131/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[151,162,552,578]" pageId="23" pageNumber="24" value="1.0">1</number>
</figureCitation>
E). A medio-dorsal ridge extends lengthwise down the animals body. The ridge is broken into two sections. The short anterior section runs from just in front of the first pair of dorsolateral processes, forward to just behind the rhinophore sheaths where it forks, with a branch going to the base of each rhinophore. The long posterior section of the ridge runs from just behind the first pair of dorsolateral processes back to the tip of the foot. Anteriorly this ridge also forks, this time with a branch going out to each of the first pair of dorsolateral processes. On either side of the mouth is an oral lobe, bearing nine-ten papillae arranged in two rows, four smaller papillae at the base of five-six larger ones. There is a pair of large, conical perfoliate rhinophores, each bearing about
<number box="[319,348,832,858]" pageId="23" pageNumber="24" value="19.0">19</number>
lamellae; completely retractile into elevated rhinophoral sheaths. Each rhinophore is nestled in a large, flattened rhinophoral sheath, into which it is able to completely retract. The rhinophore sheaths have a tall, slightly laterally compressed stalk from which the rhinophore protrudes. One anterior and two anterolateral finger-like papillae, as well as a taller, quite elaborate posterior structure which looks quite similar in shape to one of the dorsolateral processes, surround each rhinophore, at the upper edge of its stalk. Posterior to the rhinophores are three pairs of dorsolateral five branched processes, followed by three-four small, single processes arranged along the dorsal midline. The translucent gills on each dorsolateral process are inserted at the junction of the main stalk and the branches. There are three well-developed, nonretractile, tripinnate gills per dorsolateral process. The reproductive opening is located on the right side, midway between the rhinophore sheath and the first dorsolateral process. The anus is smaller and it is located on the right side of the dorsum between the first and second pair of dorsolateral processes, closer to the second. The sole of the foot is narrow and pale yellow in color.
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph blockId="23.[151,1437,152,2018]" lastBlockId="24.[151,1436,152,218]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis bold="true" box="[197,432,1311,1337]" pageId="23" pageNumber="24">Alimentary Canal:</emphasis>
The buccal bulb is relatively large. The labial cuticle is thick, and consists of elongate and numerous overlapping scales, arranged in fairly regular rows (
<figureCitation box="[952,1064,1352,1378]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[1010,1040,1352,1378]" pageId="23" pageNumber="24" value="12.0">12</number>
A
</figureCitation>
). The jaws are roughly oval in shape, without a distinct masticatory process (
<figureCitation box="[695,803,1392,1418]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[751,783,1392,1418]" pageId="23" pageNumber="24" value="12.0">12</number>
B
</figureCitation>
). The radular formulae of some of the specimens dissected are:
<date box="[281,332,1432,1458]" pageId="23" pageNumber="24" value="1937-10">
<number box="[281,310,1432,1458]" pageId="23" pageNumber="24" value="37.0">37</number>
x
</date>
<date box="[338,426,1432,1458]" pageId="23" pageNumber="24" value="21-01-21">
21.1.
<number box="[397,426,1432,1458]" pageId="23" pageNumber="24" value="21.0">21</number>
</date>
(
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[441,528,1432,1458]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<quantity box="[534,703,1432,1458]" metricMagnitude="1" metricUnit="m" metricValue="2.50072" pageId="23" pageNumber="24" unit="mm" value="25007.2">25007, 20 mm</quantity>
preserved),
<number box="[847,920,1432,1458]" pageId="23" pageNumber="24" value="36.0" valueMax="37.0" valueMin="35.0">
35
<date box="[891,920,1432,1458]" pageId="23" pageNumber="24" value="1937-10">37</date>
</number>
x
<date box="[948,1035,1432,1458]" pageId="23" pageNumber="24" value="13-01-13">
13.1.
<number box="[1006,1035,1432,1458]" pageId="23" pageNumber="24" value="13.0">13</number>
</date>
(
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[1050,1137,1432,1458]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<quantity box="[1143,1308,1432,1458]" metricMagnitude="-2" metricUnit="m" metricValue="5.3625" metricValueMax="7.125" metricValueMin="3.6" pageId="23" pageNumber="24" unit="mm" value="53.625" valueMax="71.25" valueMin="36.0">36-71, 25 mm</quantity>
preserved;
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[151,236,1472,1498]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<quantity box="[244,418,1472,1498]" metricMagnitude="1" metricUnit="m" metricValue="2.4996169999999998" pageId="23" pageNumber="24" unit="mm" value="24996.17">24996, 17 mm</quantity>
preserved),
<number box="[566,639,1472,1498]" pageId="23" pageNumber="24" value="31.5" valueMax="33.0" valueMin="30.0">
30
<date box="[611,639,1472,1498]" pageId="23" pageNumber="24" value="1933-10-10">33</date>
</number>
x
<number box="[672,699,1472,1498]" pageId="23" pageNumber="24" value="10.0">10</number>
<date box="[715,805,1472,1498]" pageId="23" pageNumber="24" value="10-11-01">11.1.10</date>
<number box="[821,846,1472,1498]" pageId="23" pageNumber="24" value="11.0">11</number>
(
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[862,950,1472,1498]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<quantity box="[950,1122,1472,1498]" metricMagnitude="1" metricUnit="m" metricValue="2.4996169999999998" pageId="23" pageNumber="24" unit="mm" value="24996.17">24996, 17 mm</quantity>
preserved;
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[1262,1347,1472,1498]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<quantity metricMagnitude="1" metricUnit="m" metricValue="1.480111" pageId="23" pageNumber="24" unit="mm" value="14801.11">14801, 11 mm</quantity>
preserved). The rachidian teeth are higher than wider (
<figureCitation box="[885,1033,1512,1538]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Figs.
<number box="[952,983,1512,1538]" pageId="23" pageNumber="24" value="12.0">12</number>
D,E
</figureCitation>
). They lack well-developed denticles on each side of the large cusp but have a number of irregular faint denticles at the posterior base of the cusp (
<figureCitation box="[220,336,1592,1618]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Figs.
<number box="[286,317,1592,1618]" pageId="23" pageNumber="24" value="12.0">12</number>
F
</figureCitation>
). Most of the specimens examined have from
<number box="[872,966,1592,1618]" pageId="23" pageNumber="24" value="11.5" valueMax="13.0" valueMin="10.0">10 to 13</number>
laterals (
<figureCitation box="[1074,1181,1592,1618]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[1128,1158,1592,1618]" pageId="23" pageNumber="24" value="12.0">12</number>
D
</figureCitation>
), but in one specimen (
<collectionCode LSID="urn:lsid:biocol.org:col:34802" box="[159,246,1632,1658]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
<number box="[256,331,1632,1658]" pageId="23" pageNumber="24" value="25007.0">25007</number>
) the number rises to
<number box="[594,623,1632,1658]" pageId="23" pageNumber="24" value="21.0">21</number>
(
<figureCitation box="[643,753,1632,1658]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[701,732,1632,1658]" pageId="23" pageNumber="24" value="12.0">12</number>
E
</figureCitation>
). The laterals are all blade-like, increasing outwards but always the two or three outer laterals a little bit shorter (
<figureCitation box="[833,982,1672,1698]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Figs.
<number box="[901,932,1672,1698]" pageId="23" pageNumber="24" value="12.0">12</number>
D,E
</figureCitation>
). A long unpaired oral gland is found ventrally (
<figureCitation box="[270,378,1712,1738]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="23" pageNumber="24">
Fig.
<number box="[325,356,1712,1738]" pageId="23" pageNumber="24" value="12.0">12</number>
C
</figureCitation>
). It opens on the mid-ventral side of the mouth and runs back to the region of the reproductive system. A pair of yellowish, elongate and highly branched salivary glands is attached to the posterior side of the oesophagus. The oesophagus is short and wide. The two anterior digestive glands open on the upper surface of the stomach. Both glands are large and elongate but the gland on the right is larger than the left one. They each have a single branch that runs to their respective anterior dorsolateral process. The posterior lobe of the digestive gland opens on the lower left surface of the stomach. All pairs of dorsolateral processes, behind the first pair, receive branches from the posterior portion of the digestive gland. The posterior chamber of the stomach is tubular and armed, in its anterior part, with about
<number box="[886,985,1992,2018]" pageId="23" pageNumber="24" value="22.5" valueMax="25.0" valueMin="20.0">20 to 25</number>
longitudinal rows of chitinous brown spines. These spines are elongate, straight and rounded at the tip (
<figureCitation box="[916,1025,152,178]" captionStart="FIGURE 12. A G" captionStartId="24.[151,255,1438,1462]" captionTargetBox="[251,1335,277,1411]" captionTargetId="figure@24.[251,1336,277,1412]" captionTargetPageId="24" captionText="FIGURE 12. A G, Scanning electron micrographs of Bornella sarape Bertsch, 1980. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100 Μm). B, Left jaw (LACM 24996, scale bar: 100 Μm). C, Oral gland (LACM 25007, scale bar: 100 Μm). D, Half radula (LACM 24996, scale bar: 10 Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20 Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10 Μm). G, Stomach spines (LACM 24966, scale bar: 100 Μm)." httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="24" pageNumber="25">
Fig.
<number box="[970,1001,152,178]" pageId="24" pageNumber="25" value="12.0">12</number>
G
</figureCitation>
). From the stomach the short intestine descends to the ventral side and then bends dorsally to the anus.
</paragraph>
<caption httpUri="https://zenodo.org/record/185142/files/figure.png" pageId="24" pageNumber="25" targetBox="[251,1335,277,1411]" targetPageId="24">
<paragraph blockId="24.[151,1436,1438,1603]" pageId="24" pageNumber="25">
<emphasis bold="true" box="[151,290,1438,1462]" pageId="24" pageNumber="25">FIGURE 12</emphasis>
. AG, Scanning electron micrographs of
<taxonomicName authority="Bertsch, 1980" authorityName="Bertsch" authorityYear="1980" box="[748,1085,1438,1462]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="sarape">
<emphasis box="[748,922,1439,1462]" italics="true" pageId="24" pageNumber="25">Bornella sarape</emphasis>
Bertsch, 1980
</taxonomicName>
. A, Rodlets of the labial cuticle (LACM 24996, scale bar: 100Μm). B, Left jaw (LACM 24996, scale bar: 100Μm). C, Oral gland (LACM 25007, scale bar: 100Μm). D, Half radula (LACM 24996, scale bar: 10Μm). E, Right half-rows of radular teeth (LACM 25007, scale bar: 20Μm). F, Lateral view of the rachidian tooth (LACM 24966, scale bar: 10Μm). G, Stomach spines (LACM 24966, scale bar: 100Μm).
</paragraph>
</caption>
<paragraph blockId="24.[151,1436,1646,1992]" pageId="24" pageNumber="25">
<emphasis bold="true" box="[197,474,1646,1672]" pageId="24" pageNumber="25">Reproductive system:</emphasis>
(
<figureCitation box="[497,599,1646,1672]" captionStart="FIGURE 4. A E" captionStartId="11.[151,255,1904,1928]" captionTargetBox="[187,1406,226,1838]" captionTargetId="figure@11.[125,1461,174,1887]" captionTargetPageId="11" captionText="FIGURE 4. A E, Drawings of reproductive systems of Bornella spp. A, Bornella stellifer (CASIZ 075076). B, B. calcarata (CASIZ 118665). C, B. hermanni (AM C. 113793). D, B. sarape (LACM 24996). E, B. anguilla (CASIZ 173389). Scale bar = 1 mm. Abbreviations: am, ampulla; fglm, female gland mass; hd, hermaphrodite duct; p, penis; pb, penial bulb; pr, prostate; rs, distal allosperm receptacle; va, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/185134/files/figure.png" pageId="24" pageNumber="25">
Fig.
<number box="[559,576,1646,1672]" pageId="24" pageNumber="25" value="4.0">4</number>
D
</figureCitation>
): The ovotestis consists of between five and eight closely packed, rounded or somewhat pyriform follicles lying over the posterior digestive gland and each follicle is joined to the hermaphrodite duct by a fine tube. The hermaphrodite duct runs around the stomach on the left side and expands into the large C-shaped ampulla on the dorsal side of the female gland mass. From the ampullar region the narrow hermaphrodite duct divides, giving rise to the separate oviduct and vas deferens. The visible oviduct is very short before it disappears into the female gland mass. The vas deferens is relatively long and coiled, and appears to have a layer of prostate gland tissue along its entire length. The large elongate penial bulb lacks any recognizable penial papilla, and no traces of penial armature were found in any of the specimens dissected. A small distal pyriform allosperm receptacle opens at the genital aperture through a long duct.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="discussion">
<paragraph blockId="25.[151,1436,151,578]" pageId="25" pageNumber="26">
<emphasis bold="true" box="[197,321,151,177]" pageId="25" pageNumber="26">Remarks:</emphasis>
<taxonomicName box="[330,521,152,177]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="sarape">
<emphasis box="[330,521,152,177]" italics="true" pageId="25" pageNumber="26">Bornella sarape</emphasis>
</taxonomicName>
is known from several specimens from the Pacific coast of
<collectingCountry box="[1234,1323,152,178]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
and
<collectingCountry name="Panama" pageId="25" pageNumber="26">Panama</collectingCountry>
(
<bibRefCitation author="Bertsch" box="[215,377,192,218]" pageId="25" pageNumber="54" refString="Bertsch, H. (1980) A new species of Bornella from tropical west-America (Mollusca, Opisthobranchia). Spixiana, 3, 33 - 42." type="journal article" year="1980">
Bertsch
<number box="[315,377,192,218]" pageId="25" pageNumber="26" value="1980.0">1980</number>
</bibRefCitation>
;
<bibRefCitation author="Bertsch" box="[389,695,192,218]" pageId="25" pageNumber="54" refString="Bertsch, H. &amp; Kerstitch. A. (1984) Distribution and radular morphology of various nudibranchs (Gastropoda: Opisthobranchia) from the Gulf of California, Mexico. The Vel iger, 26, 264 - 273." type="journal article" year="1984">
Bertsch &amp; Kerstitch
<number box="[633,695,192,218]" pageId="25" pageNumber="26" value="1984.0">1984</number>
</bibRefCitation>
;
<bibRefCitation author="Hermosillo" box="[707,977,192,218]" pageId="25" pageNumber="55" refString="Hermosillo, A., Behrens, D. W. &amp; Rios Jara, E. (2006) Opisthobranquios de Mexico. Guia de babosas marinas del Pacifico, Golfo de California y las islas oceanicas. CONABIO, Guadalajara, Jalisco, Mexico, 144 pp." type="book" year="2006">
Hermosillo
<emphasis box="[847,906,192,217]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
<number box="[913,977,192,218]" pageId="25" pageNumber="26" value="2006.0">2006</number>
</bibRefCitation>
). The external anatomy, coloration and radula described by the previous authors agree with those of our specimens, except for the presence of a constant number of three pairs of branching dorsolateral processes, and the presence of tripinnate gills, which have been previously recorded as uni- or bi-pinnate. The reproductive system is described here for the first time. The colour pattern of this species clearly distinguishes it from other species of
<taxonomicName box="[1132,1234,352,377]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis box="[1132,1234,352,377]" italics="true" pageId="25" pageNumber="26">Bornella</emphasis>
</taxonomicName>
. The pale yellow body with its brown reticulate pattern distinguishes it from the species described from the Pacific and the Caribbean, which have orange or reddish reticulations. Internally the main distinguishing features are that the penis lacks a distinct papilla and is unarmed and the rachidian tooth has many irregular and faint denticles at the base of the median cusp. Also the rounded tips of the stomach spines are more similar to those of
<taxonomicName class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="anguilla">
<emphasis italics="true" pageId="25" pageNumber="26">B. anguilla</emphasis>
</taxonomicName>
than to the other species, but in
<taxonomicName box="[628,759,552,577]" class="Gastropoda" family="Bornellidae" genus="Bornella" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="anguilla">
<emphasis box="[628,759,552,577]" italics="true" pageId="25" pageNumber="26">B. anguilla</emphasis>
</taxonomicName>
they are more numerous.
</paragraph>
</subSubSection>
</treatment>
</document>