314 lines
36 KiB
XML
314 lines
36 KiB
XML
<document ID-DOI="10.11646/zootaxa.5249.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687292" ID-ZooBank="52C66599-6FC8-4A2D-BEA4-8D1CAC33EEE6" approvalRequired="10" approvalRequired_for_textStreams="9" approvalRequired_for_treatments="1" checkinTime="1677655666126" checkinUser="plazi" docAuthor="Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S." docDate="2023" docId="03F95931630E426D288FFA71FEE0F83B" docLanguage="en" docName="zootaxa.5249.2.1.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philanthus variegatus subsp. ecoronatus Dufour 1854" docType="treatment" docVersion="2" lastPageNumber="182" masterDocId="FFC02149631242722818FF98FF85FFC8" masterDocTitle="The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species" masterLastPageNumber="189" masterPageNumber="151" pageNumber="179" updateTime="1677656267120" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Malash, Alyaa A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-6053-0693</mods:nameIdentifier>
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<mods:affiliation>https: // orcid. org / 0000 - 0001 - 6053 - 0693</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edmardash, Yusuf A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-3365-0822</mods:nameIdentifier>
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<mods:affiliation>https: // orcid. org / 0000 - 0002 - 3365 - 0822</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gadallah, Neveen S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4381-9599</mods:nameIdentifier>
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<mods:affiliation>https: // orcid. org / 0000 - 0002 - 4381 - 9599</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5249</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>151</mods:start>
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<mods:end>189</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.2.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5249.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7687292</mods:identifier>
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<mods:identifier type="ZooBank">52C66599-6FC8-4A2D-BEA4-8D1CAC33EEE6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03F95931630E426D288FFA71FEE0F83B" httpUri="http://treatment.plazi.org/id/03F95931630E426D288FFA71FEE0F83B" lastPageId="31" lastPageNumber="182" pageId="28" pageNumber="179">
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<subSubSection box="[151,728,1513,1540]" pageId="28" pageNumber="179" type="nomenclature">
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<paragraph blockId="28.[151,728,1513,1575]" box="[151,728,1513,1540]" pageId="28" pageNumber="179">
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<heading bold="true" box="[151,728,1513,1540]" fontSize="11" level="1" pageId="28" pageNumber="179" reason="1">
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<taxonomicName ID-CoL="5KCX5" authority="Dufour, 1854" authorityName="Dufour" authorityYear="1854" box="[151,728,1513,1540]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="ecoronatus">
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<emphasis bold="true" box="[151,728,1513,1540]" pageId="28" pageNumber="179">
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<emphasis bold="true" box="[151,559,1513,1540]" italics="true" pageId="28" pageNumber="179">Philanthus variegatus ecoronatus</emphasis>
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<bibRefCitation author="Dufour, L." box="[566,728,1513,1539]" pageId="28" pageNumber="179" pagination="375 - 382" refId="ref18099" refString="Dufour, L. (1854) Signalements de quelques especes nouvelles ou peu connues d'Hymenopteres algeriens. Annales de la Societe Entomologique de France, Serie 3, 1, 375 - 382." type="journal article" year="1854">Dufour, 1854</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,382,1549,1575]" pageId="28" pageNumber="179" type="description">
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<paragraph blockId="28.[151,728,1513,1575]" box="[151,382,1549,1575]" pageId="28" pageNumber="179">
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<figureCitation box="[151,290,1549,1575]" captionStart="FIGURE 20" captionStartId="31.[152,255,1363,1387]" captionTargetBox="[151,1436,181,1339]" captionTargetId="figure-18@31.[151,1436,181,1339]" captionTargetPageId="31" captionText="FIGURE 20. Philanthus variegatus ecoronatus Dufour, 1854: A, lateral habitus (♁); B, frontal view of head (♀); C, frontal view of head (♁)." figureDoi="http://doi.org/10.5281/zenodo.7687354" httpUri="https://zenodo.org/record/7687354/files/figure.png" pageId="28" pageNumber="179">Figs 20A–C</figureCitation>
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,
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<figureCitation box="[301,382,1549,1575]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">21A–C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,610,1618,1644]" pageId="28" pageNumber="179" type="reference_group">
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||
<paragraph blockId="28.[151,610,1618,1644]" box="[151,610,1618,1644]" pageId="28" pageNumber="179">
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<treatmentCitationGroup box="[151,610,1618,1644]" pageId="28" pageNumber="179">
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<treatmentCitation author="Dufour, L." box="[151,574,1618,1642]" page="380" pageId="28" pageNumber="179" year="1854">
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<taxonomicName ID-CoL="4G54G" authority="Dufour, 1854: 380" authorityName="Dufour" authorityPageNumber="380" authorityYear="1854" box="[151,574,1618,1642]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="ecoronatus">
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<emphasis box="[151,379,1618,1642]" italics="true" pageId="28" pageNumber="179">Philanthus ecoronatus</emphasis>
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<bibRefCitation author="Dufour, L." box="[385,574,1618,1642]" pageId="28" pageNumber="179" pagination="375 - 382" refId="ref18099" refString="Dufour, L. (1854) Signalements de quelques especes nouvelles ou peu connues d'Hymenopteres algeriens. Annales de la Societe Entomologique de France, Serie 3, 1, 375 - 382." type="journal article" year="1854">Dufour, 1854: 380</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, ♁.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="179" type="diagnosis">
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<paragraph blockId="28.[151,1437,1688,2040]" box="[151,814,1688,1715]" pageId="28" pageNumber="179">
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<emphasis bold="true" box="[151,274,1688,1714]" pageId="28" pageNumber="179">Diagnosis.</emphasis>
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Body length:
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<quantity box="[433,512,1688,1715]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="179" unit="mm" value="14.0">14 mm</quantity>
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(female); 11.6 mm (male).
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</paragraph>
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<paragraph blockId="28.[151,1437,1688,2040]" pageId="28" pageNumber="179">
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<taxonomicName authorityName="Dufour" authorityYear="1854" box="[199,573,1724,1750]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="ecoronatus">
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<emphasis box="[199,573,1724,1750]" italics="true" pageId="28" pageNumber="179">Philanthus variegatus ecoronatus</emphasis>
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</taxonomicName>
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differs from the nominotypical subspecies in the following: mesonotum shiny black (
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<figureCitation box="[225,324,1760,1786]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">Fig. 21B</figureCitation>
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) (shiny ferruginous in
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<taxonomicName authorityName="Dufour" authorityYear="1854" box="[574,842,1761,1786]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="variegatus">
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<emphasis box="[574,842,1761,1786]" italics="true" pageId="28" pageNumber="179">P. variegatus variegatus</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[856,957,1760,1786]" captionStart="FIGURE 19" captionStartId="30.[152,255,1944,1968]" captionTargetBox="[280,1308,181,1920]" captionTargetId="figure-23@30.[280,1308,181,1920]" captionTargetPageId="30" captionText="FIGURE 19. Philanthus variegatus Spinola, 1839: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster." figureDoi="http://doi.org/10.5281/zenodo.7687352" httpUri="https://zenodo.org/record/7687352/files/figure.png" pageId="28" pageNumber="179">Fig. 19D</figureCitation>
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)); scutellum and postscutellum ferruginous (
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<figureCitation box="[159,264,1796,1822]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">Fig. 21B</figureCitation>
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), propodeum black with two lateral yellow patches (
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<figureCitation box="[875,978,1796,1822]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">Fig. 21B</figureCitation>
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) (rest of thorax yellow in
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<taxonomicName authorityName="Dufour" authorityYear="1854" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="variegatus">
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<emphasis italics="true" pageId="28" pageNumber="179">P. variegatus variegatus</emphasis>
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</taxonomicName>
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, except black median sulcus (
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<figureCitation box="[612,747,1832,1858]" captionStart="FIGURE 19" captionStartId="30.[152,255,1944,1968]" captionTargetBox="[280,1308,181,1920]" captionTargetId="figure-23@30.[280,1308,181,1920]" captionTargetPageId="30" captionText="FIGURE 19. Philanthus variegatus Spinola, 1839: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster." figureDoi="http://doi.org/10.5281/zenodo.7687352" httpUri="https://zenodo.org/record/7687352/files/figure.png" pageId="28" pageNumber="179">Fig. 19C–E</figureCitation>
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)); mesoscutum sparsely punctate, punctures becoming dense posteriorly (
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<figureCitation box="[290,393,1868,1894]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">Fig. 21B</figureCitation>
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) (nearly smooth, impunctate in
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<taxonomicName authorityName="Dufour" authorityYear="1854" box="[764,1039,1869,1894]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="variegatus">
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<emphasis box="[764,1039,1869,1894]" italics="true" pageId="28" pageNumber="179">P. variegatus variegatus</emphasis>
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</taxonomicName>
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, densely punctate at junction with scutellum (
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<figureCitation box="[277,382,1904,1930]" captionStart="FIGURE 19" captionStartId="30.[152,255,1944,1968]" captionTargetBox="[280,1308,181,1920]" captionTargetId="figure-23@30.[280,1308,181,1920]" captionTargetPageId="30" captionText="FIGURE 19. Philanthus variegatus Spinola, 1839: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster." figureDoi="http://doi.org/10.5281/zenodo.7687352" httpUri="https://zenodo.org/record/7687352/files/figure.png" pageId="28" pageNumber="179">Fig. 19D</figureCitation>
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)); gastral T
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<subScript attach="left" box="[515,523,1922,1936]" fontSize="6" pageId="28" pageNumber="179">2</subScript>
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densely punctate (
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<figureCitation box="[735,838,1904,1930]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="28" pageNumber="179">Fig. 21C</figureCitation>
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) (sparsely punctate in
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<taxonomicName authorityName="Dufour" authorityYear="1854" box="[1099,1373,1905,1930]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="variegatus">
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<emphasis box="[1099,1373,1905,1930]" italics="true" pageId="28" pageNumber="179">P. variegatus variegatus</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="FIGURE 19" captionStartId="30.[152,255,1944,1968]" captionTargetBox="[280,1308,181,1920]" captionTargetId="figure-23@30.[280,1308,181,1920]" captionTargetPageId="30" captionText="FIGURE 19. Philanthus variegatus Spinola, 1839: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster." figureDoi="http://doi.org/10.5281/zenodo.7687352" httpUri="https://zenodo.org/record/7687352/files/figure.png" pageId="28" pageNumber="179">Fig. 19E</figureCitation>
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)).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="31" lastPageNumber="182" pageId="28" pageNumber="179" type="materials_examined">
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||
<paragraph blockId="28.[151,1437,1688,2040]" pageId="28" pageNumber="179">
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||
<emphasis bold="true" box="[199,434,1976,2002]" pageId="28" pageNumber="179">Material examined:</emphasis>
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||
<materialsCitation box="[440,1191,1976,2003]" collectingDate="1990-05-12" collectionCode="EFC" collectorName="Neveen Gadallah" location="Wadi El-Natroun" pageId="28" pageNumber="179" specimenCount="1" specimenCount-female="1">
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||
<specimenCount box="[440,475,1976,2002]" pageId="28" pageNumber="179" type="female">1♀</specimenCount>
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||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03F95931630E426D288FFA71FEE0F83B:8E8FBEFC630E426E29F0F820FD2CF81A" box="[488,681,1976,2002]" name="Wadi El-Natroun" pageId="28" pageNumber="179">Wadi El-Natroun</location>
|
||
,
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||
<date box="[691,804,1976,2002]" pageId="28" pageNumber="179" value="1990-05-12">
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||
<collectingDate box="[691,804,1976,2002]" pageId="28" pageNumber="179" value="1990-05-12">12.v.1990</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[815,1007,1976,2003]" pageId="28" pageNumber="179">Neveen Gadallah</collectorName>
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||
collector (
|
||
<collectionCode box="[1126,1183,1976,2002]" pageId="28" pageNumber="179">EFC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectionCode="EFC" location="Wadi El Natroun" pageId="28" pageNumber="179" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount box="[1201,1236,1976,2002]" pageId="28" pageNumber="179" type="female">1♀</specimenCount>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03F95931630E426D288FFA71FEE0F83B:8E8FBEFC630E426E2CF9F820FA19F81B" box="[1249,1436,1976,2003]" name="Wadi El Natroun" pageId="28" pageNumber="179">Wadi El Natroun</location>
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||
(Beo Ar Ashra), (18/94?) (
|
||
<collectionCode box="[446,503,2012,2038]" pageId="28" pageNumber="179">EFC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation box="[523,1127,2012,2040]" collectingDate="1954-09-28" collectionCode="ASUE" collectorName="Aly" location="Marsa" pageId="28" pageNumber="179" specimenCount="1" stateProvince="Matruh">
|
||
1♁,
|
||
<location LSID="urn:lsid:plazi:treatment:03F95931630E426D288FFA71FEE0F83B:8E8FBEFC630E426E2A24F845FD07F83F" box="[572,642,2013,2039]" name="Marsa" pageId="28" pageNumber="179" stateProvince="Matruh">Marsa</location>
|
||
<collectingRegion box="[649,734,2012,2038]" country="Egypt" name="Matruh" pageId="28" pageNumber="179">Matruh</collectingRegion>
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||
,
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||
<date box="[745,868,2012,2039]" pageId="28" pageNumber="179" value="1954-09-28">
|
||
<collectingDate box="[745,868,2012,2039]" pageId="28" pageNumber="179" value="1954-09-28">28.ix.1954</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[878,920,2012,2038]" pageId="28" pageNumber="179">Aly</collectorName>
|
||
collector (
|
||
<collectionCode box="[1039,1118,2012,2038]" pageId="28" pageNumber="179">ASUE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687350" ID-Zenodo-Dep="7687350" httpUri="https://zenodo.org/record/7687350/files/figure.png" pageId="29" pageNumber="180" startId="29.[152,255,1944,1968]" targetBox="[184,1404,181,1920]" targetPageId="29">
|
||
<paragraph blockId="29.[151,1436,1944,2005]" pageId="29" pageNumber="180">
|
||
<emphasis bold="true" box="[152,293,1944,1968]" pageId="29" pageNumber="180">FIGURE 18.</emphasis>
|
||
<taxonomicName authority="Lepeletier, 1845" authorityName="Lepeletier de Saint Fargeau" authorityYear="1845" box="[299,823,1944,1968]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="triangulum" subSpecies="abdelcader">
|
||
<emphasis box="[299,648,1944,1968]" italics="true" pageId="29" pageNumber="180">Philanthus triangulum abdelcader</emphasis>
|
||
<bibRefCitation author="Lepeletier de Saint Fargeau, A. L. M." box="[655,823,1944,1968]" pageId="29" pageNumber="180" pagination="25 - 36" refId="ref18736" refString="Lepeletier de Saint Fargeau, A. L. M. (1845) Histoire naturelle des Insectes. Hymenopteres. Vol. 3. Librairie Encyclopedique de Roret, Paris, 646 + 4 pp., pls. 25 - 36." type="book chapter" year="1845">Lepeletier, 1845</bibRefCitation>
|
||
</taxonomicName>
|
||
: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (♀); E, dorsal view of gaster.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687352" ID-Zenodo-Dep="7687352" httpUri="https://zenodo.org/record/7687352/files/figure.png" pageId="30" pageNumber="181" startId="30.[152,255,1944,1968]" targetBox="[280,1308,181,1920]" targetPageId="30">
|
||
<paragraph blockId="30.[151,1436,1944,2005]" pageId="30" pageNumber="181">
|
||
<emphasis bold="true" box="[152,293,1944,1968]" pageId="30" pageNumber="181">FIGURE 19.</emphasis>
|
||
<taxonomicName authority="Spinola, 1839" authorityName="Spinola" authorityYear="1839" box="[299,672,1944,1969]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis box="[299,522,1944,1968]" italics="true" pageId="30" pageNumber="181">Philanthus variegatus</emphasis>
|
||
<bibRefCitation author="Spinola, M." box="[528,672,1944,1969]" pageId="30" pageNumber="181" pagination="437 - 546" refId="ref19496" refString="Spinola, M. (1839) Compte-rendu des Hymenopteres recueillis par M. Fischer pendant son voyage en Egypte, et communiques par M. le Docteur Waltl a Maximilien Spinola. Annales de la Societe Entomologique de France, 7, 437 - 546." type="journal article" year="1839">Spinola, 1839</bibRefCitation>
|
||
</taxonomicName>
|
||
: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687354" ID-Zenodo-Dep="7687354" httpUri="https://zenodo.org/record/7687354/files/figure.png" pageId="31" pageNumber="182" startId="31.[152,255,1363,1387]" targetBox="[151,1436,181,1339]" targetPageId="31">
|
||
<paragraph blockId="31.[151,1437,1363,1425]" pageId="31" pageNumber="182">
|
||
<emphasis bold="true" box="[152,295,1363,1387]" pageId="31" pageNumber="182">FIGURE 20.</emphasis>
|
||
<taxonomicName authority="Dufour, 1854" authorityName="Dufour" authorityYear="1854" box="[302,794,1363,1387]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="ecoronatus">
|
||
<emphasis box="[302,647,1363,1387]" italics="true" pageId="31" pageNumber="182">Philanthus variegatus ecoronatus</emphasis>
|
||
<bibRefCitation author="Dufour, L." box="[654,794,1363,1387]" pageId="31" pageNumber="182" pagination="375 - 382" refId="ref18099" refString="Dufour, L. (1854) Signalements de quelques especes nouvelles ou peu connues d'Hymenopteres algeriens. Annales de la Societe Entomologique de France, Serie 3, 1, 375 - 382." type="journal article" year="1854">Dufour, 1854</bibRefCitation>
|
||
</taxonomicName>
|
||
: A, lateral habitus (♁); B, frontal view of head (♀); C, frontal view of head (♁).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="31.[151,1437,1469,2035]" box="[199,1268,1469,1496]" pageId="31" pageNumber="182">
|
||
<emphasis bold="true" box="[199,525,1469,1495]" pageId="31" pageNumber="182">Previous Egyptian records:</emphasis>
|
||
Wadi El-Natroun (
|
||
<bibRefCitation author="Gadallah, N. S." box="[737,907,1469,1495]" pageId="31" pageNumber="182" pagination="68 - 80" refId="ref18239" refString="Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80." type="journal article" year="1996">Gadallah 1996</bibRefCitation>
|
||
); Marsa
|
||
<collectingRegion box="[1004,1086,1469,1495]" country="Egypt" name="Matruh" pageId="31" pageNumber="182">Matruh</collectingRegion>
|
||
(present study).
|
||
</paragraph>
|
||
<paragraph blockId="31.[151,1437,1469,2035]" box="[199,1254,1505,1532]" pageId="31" pageNumber="182">
|
||
<emphasis bold="true" box="[199,503,1505,1531]" pageId="31" pageNumber="182">Extralimital distribution:</emphasis>
|
||
<collectingCountry box="[510,597,1505,1531]" name="Algeria" pageId="31" pageNumber="182">Algeria</collectingCountry>
|
||
,
|
||
<collectingCountry box="[607,708,1505,1531]" name="Armenia" pageId="31" pageNumber="182">Armenia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[719,781,1505,1531]" name="Chad" pageId="31" pageNumber="182">Chad</collectingCountry>
|
||
,
|
||
<collectingCountry box="[792,844,1505,1531]" name="Italy" pageId="31" pageNumber="182">Italy</collectingCountry>
|
||
,
|
||
<collectingCountry box="[855,923,1505,1531]" name="Libya" pageId="31" pageNumber="182">Libya</collectingCountry>
|
||
,
|
||
<collectingCountry box="[935,1039,1506,1532]" name="Morocco" pageId="31" pageNumber="182">Morocco</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1050,1154,1505,1531]" name="Palestine" pageId="31" pageNumber="182">Palestine</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1164,1250,1505,1531]" name="Tunisia" pageId="31" pageNumber="182">Tunisia</collectingCountry>
|
||
,
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="182" type="discussion">
|
||
<paragraph blockId="31.[151,1437,1469,2035]" pageId="31" pageNumber="182">
|
||
<emphasis bold="true" box="[199,327,1541,1567]" pageId="31" pageNumber="182">Comment:</emphasis>
|
||
This subspecies was recorded for the first time from
|
||
<collectingCountry box="[909,976,1542,1568]" name="Egypt" pageId="31" pageNumber="182">Egypt</collectingCountry>
|
||
by
|
||
<bibRefCitation author="Gadallah, N. S." box="[1014,1199,1541,1567]" pageId="31" pageNumber="182" pagination="68 - 80" refId="ref18239" refString="Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80." type="journal article" year="1996">Gadallah (1996)</bibRefCitation>
|
||
. The characters agree with de Beaumont’s key (1949: 179, couplet 6) in having mesopleuron and propodeum greatly marked with yellow (
|
||
<figureCitation box="[159,297,1613,1639]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21A, B</figureCitation>
|
||
); upper mesopleuron densely punctate (
|
||
<figureCitation box="[749,854,1613,1639]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21A</figureCitation>
|
||
); gaster mostly yellow, narrowly black medially on T
|
||
<subScript attach="left" box="[168,193,1668,1682]" fontSize="6" pageId="31" pageNumber="182">1–4</subScript>
|
||
, with terminal depressions ferruginous (
|
||
<figureCitation box="[659,764,1649,1675]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21C</figureCitation>
|
||
); T
|
||
<subScript attach="left" box="[804,812,1668,1682]" fontSize="6" pageId="31" pageNumber="182">2</subScript>
|
||
more or less densely, finely punctate (
|
||
<figureCitation box="[1261,1367,1649,1675]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21C</figureCitation>
|
||
); face entirely pale up to level of inner eye emargination in female (
|
||
<figureCitation box="[848,951,1685,1711]" captionStart="FIGURE 20" captionStartId="31.[152,255,1363,1387]" captionTargetBox="[151,1436,181,1339]" captionTargetId="figure-18@31.[151,1436,181,1339]" captionTargetPageId="31" captionText="FIGURE 20. Philanthus variegatus ecoronatus Dufour, 1854: A, lateral habitus (♁); B, frontal view of head (♀); C, frontal view of head (♁)." figureDoi="http://doi.org/10.5281/zenodo.7687354" httpUri="https://zenodo.org/record/7687354/files/figure.png" pageId="31" pageNumber="182">Fig. 20B</figureCitation>
|
||
), and dark reddish brown to ferruginous in male (
|
||
<figureCitation box="[223,328,1721,1747]" captionStart="FIGURE 20" captionStartId="31.[152,255,1363,1387]" captionTargetBox="[151,1436,181,1339]" captionTargetId="figure-18@31.[151,1436,181,1339]" captionTargetPageId="31" captionText="FIGURE 20. Philanthus variegatus ecoronatus Dufour, 1854: A, lateral habitus (♁); B, frontal view of head (♀); C, frontal view of head (♁)." figureDoi="http://doi.org/10.5281/zenodo.7687354" httpUri="https://zenodo.org/record/7687354/files/figure.png" pageId="31" pageNumber="182">Fig. 20C</figureCitation>
|
||
); scutellum and postscutellum ferruginous (
|
||
<figureCitation box="[826,930,1721,1747]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21B</figureCitation>
|
||
). It differs from de Beaumont description in having the pale colour of the face ivory white and extending very close to the middle ocellus (in de Beaumont’s specimens, 1949: 178, the pale colour of the face is yellow, and touching the middle ocellus, see fig. 1). The Tunisian specimen (
|
||
<bibRefCitation author="Ben Khedher, H. & Yildirim, E. & Braham, M." box="[272,582,1829,1855]" pageId="31" pageNumber="182" pagination="1 - 10" refId="ref17622" refString="Ben Khedher, H., Yildirim, E. & Braham, M. (2021) A checklist of Philanthini and Pseudoscoliini (Hymenoptera: Crabronidae) with new records from Tunisia. Transactions of the American Entomological Society, 147, 1 - 10. https: // doi. org / 10.3157 / 061.147.0101" type="journal article" year="2021">
|
||
Ben Khedher
|
||
<emphasis box="[431,488,1829,1855]" italics="true" pageId="31" pageNumber="182">et al.</emphasis>
|
||
2021: 8
|
||
</bibRefCitation>
|
||
, fig. 3J) is identical to de Beaumont’s. This may be a sexual difference (the Egyptian specimen is a female, while
|
||
<bibRefCitation author="de Beaumont, J." box="[577,807,1865,1891]" pageId="31" pageNumber="182" pagination="173 - 216" refId="ref17822" refString="de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216." type="journal article" year="1949">de Beaumont’s 1949</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Ben Khedher, H. & Yildirim, E. & Braham, M." box="[861,1139,1865,1892]" pageId="31" pageNumber="182" pagination="1 - 10" refId="ref17622" refString="Ben Khedher, H., Yildirim, E. & Braham, M. (2021) A checklist of Philanthini and Pseudoscoliini (Hymenoptera: Crabronidae) with new records from Tunisia. Transactions of the American Entomological Society, 147, 1 - 10. https: // doi. org / 10.3157 / 061.147.0101" type="journal article" year="2021">
|
||
Ben Khedher
|
||
<emphasis box="[1017,1073,1865,1891]" italics="true" pageId="31" pageNumber="182">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
, specimens are males).
|
||
</paragraph>
|
||
<paragraph blockId="31.[151,1437,1469,2035]" pageId="31" pageNumber="182">
|
||
The specimen of Wadi El Natroun,
|
||
<date box="[611,720,1901,1927]" pageId="31" pageNumber="182" value="1990-05-12">12.v.1990</date>
|
||
(EFC) was misidentified by Giles Roche as
|
||
<taxonomicName authority="Lucas. However" authorityName="Lucas. However" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="dufouri">
|
||
<emphasis box="[1240,1352,1901,1927]" italics="true" pageId="31" pageNumber="182">P. dufouri</emphasis>
|
||
Lucas. However
|
||
</taxonomicName>
|
||
, it differs from
|
||
<taxonomicName box="[441,553,1937,1963]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="dufouri">
|
||
<emphasis box="[441,553,1937,1963]" italics="true" pageId="31" pageNumber="182">P. dufouri</emphasis>
|
||
</taxonomicName>
|
||
in having the scutellum ferruginous (
|
||
<figureCitation box="[989,1092,1937,1963]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21B</figureCitation>
|
||
) (clearly black in
|
||
<taxonomicName box="[1307,1419,1937,1963]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="dufouri">
|
||
<emphasis box="[1307,1419,1937,1963]" italics="true" pageId="31" pageNumber="182">P. dufouri</emphasis>
|
||
</taxonomicName>
|
||
); gastral terga predominantly yellow, narrowly black medially (
|
||
<figureCitation box="[845,946,1973,1999]" captionStart="FIGURE 21" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-23@32.[165,1422,181,1919]" captionTargetPageId="32" captionText="FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster." figureDoi="http://doi.org/10.5281/zenodo.7687356" httpUri="https://zenodo.org/record/7687356/files/figure.png" pageId="31" pageNumber="182">Fig. 21C</figureCitation>
|
||
) (black in
|
||
<taxonomicName box="[1067,1177,1973,1999]" class="Insecta" family="Crabronidae" genus="Philanthus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="dufouri">
|
||
<emphasis box="[1067,1177,1973,1999]" italics="true" pageId="31" pageNumber="182">P. dufouri</emphasis>
|
||
</taxonomicName>
|
||
, with yellow markings clearly delimited).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |